The AusTraits Plant Dictionary (APD) Ontology

Release 2.1.0

Latest version:
https://w3id.org/APD/index.html
This version:
https://w3id.org/APD/release/2.1.0/index.html
Previous version:
https://w3id.org/APD/release/2.0.0/index.html
Authors:
Elizabeth Wenk, Daniel Falster, Rachael Gallagher, Hervé Sauquet, Rowan Brownlee, Carl Boettiger
Contributors:
Tony Auld, Russell Barrett, Timothy Brodribb, Brendan Choat, David Coleman, Félix de Tombeur, Lily Dun, David Ellsworth, Carl Gosper, Lydia Guja, Gregory J. Jordan, Tom Le Breton, Andy Leigh, Patricia Lu-Irving, Belinda Medlyn, Rachael Nolan, Mark Ooi, Karen D. Sommerville, Ruby Stephens, Peter Vesk, Matthew White, Ian J. Wright, Sophie Yang
Human readable tables:

Traits categorical_values

Serializations:
JSON-LD N-Triples N-Quads TTL
License:
http://purl.org/NET/rdflicense/cc-by4.0
Vocabulary service:
JSON-LD
Source:

10.5281/zenodo.8040789

JSON-LD
Contributing:
contributions welcome
Cite as:
Wenk EH, Sauquet H, Gallagher RV, Brownlee R, Boettiger C, Coleman D, Yang S, Auld T, Barrett RL, Brodribb T, Choat B, Dun L, Ellsworth D, Gosper C, Guja L, Jordan GJ, Breton T, Leigh A, Irving PMedlyn B, Nolan R, Ooi M, Sommerville KD, Vesk P, White M, Wright IJ & Falster DS (2024) The AusTraits Plant Dictionary. Scientific Data 11, 537. DOI: 10.1038/s41597-024-03368-z

Abstract

This document describes the schema available from the APD namespace. The APD offers detailed descriptions for more than 500 plant trait concepts. It includes trait focused on plant morphology, plant nutrient concentrations, plant physiology, plant life history, and plant fire response. The definitions will be useful to researchers from diverse disciplines, including plant functional ecology, plant taxonomy, and conservation biology. All trait concepts are supported by comprehensive metadata including trait descriptions, allowable trait values, allowable ranges, preferred units, keywords, references, and links to matches in a selection of trait databases. The traits describe here also fully support the AusTraits plant trait database, <doi.org/10.5281/zenodo.3568417>.

Table of Contents

  1. Introduction
  2. Trait groups: Hierarchical trait groupings, included to enhance trait findability.
  3. Trait concepts: Traits defined within the vocabulary.
  4. Values for categorical traits: Allowed categorical trait values to support the defined trait concepts.
  5. Glossary: Terms used as keywords that were not available in other published vocabularies.
  6. Acknowledgements

1. Introduction

The APD is a compilation of the traits concepts used by AusTraits, a database of Australian plant traits. APD includes definitions for nearly 500 traits pertaining to plant functional ecology and plant morphology. The metadata for each trait concept has been reviewed by multiple people. Metadata includes a trait definition, accepted units and ranges (for numeric traits), allowable values (for categorical traits), keywords and trait groupings to enhance trait findability, references, and links to identical/similar traits in other trait databases. APD will be published in a machine-readable format (rdf representation), allowing the traits to be readily re-used by other databases. This is a human-friendly output, with embedded links to the references, other trait databases and keywords/concepts borrowed from other ontologies.

The APD is a community resource and we welcome any suggestions for edits, including additional key references for each trait or other trait databases for which we should add links.

2. Trait groups

The APD Trait groups express broad trait groupings, with each trait concept mapped into one or more trait groups as narrower concepts.

plant trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000000
description A measurable or observable characteristic of a cellular component [GO:0005575], biological process [GO:0008150] or molecular function [GO:0003674] that is part of, or has participant a plant anatomical entity [PO:0025131] and/or a plant structure development stage [PO:0009012]. This includes physiological, molecular and biochemical traits as well.
has narrower biochemical trait
plant morphology trait
biological process trait, physiological process trait
life history trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000000

biochemical trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000002
description A plant trait [TO:0000387] which is a quality of a chemical entity [CHEBI:24431], a plant metabolite [CHEBI:76924], or a molecular function [GO:0003674], which occurs in a cellular component [GO:0005575], plant anatomical entity [PO:0025131] or during a plant structure development stage [PO:0009012].
has narrower mineral and ion content trait
metabolite content trait
stable isotope ratio determination
has broader plant trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000002

mineral and ion content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000004
description A biochemical trait [TO:0000277] that is the amount of a particular mineral or ion constituent occurring in a plant anatomical entity [PO:0025131].
has narrower leaf mineral and ion content trait
stem mineral and ion content trait
bark mineral and ion content trait
root mineral and ion content trait
reproductive shoot system mineral and ion content trait
cell or tissue mineral and ion content trait
has broader biochemical trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000004

leaf mineral and ion content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000006
description A mineral and ion content trait which is measured on a leaf [PO:0025034].
has narrower live leaf mineral and ion content trait
senesced leaf mineral and ion content trait
has broader mineral and ion content trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000006

live leaf mineral and ion content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000008
description A mineral and ion content trait which is measured on a live [PATO:0001421] leaf [PO:0025034].
has narrower Leaf aluminium (Al) content per unit leaf dry mass
Leaf boron (B) content per unit leaf dry mass
Leaf carbon (C) content per unit leaf dry mass
Leaf calcium (Ca) content per unit leaf dry mass
Leaf chlorine (Cl) content per unit leaf dry mass
Leaf chromium (Cr) content per unit leaf dry mass
Leaf cobalt (Co) content per unit leaf dry mass
Leaf copper (Cu) content per unit leaf dry mass
Leaf iron (Fe) content per unit leaf dry mass
Leaf potassium (K) content per unit leaf area
Leaf potassium (K) content per unit leaf dry mass
Leaf magnesium (Mg) content per unit leaf dry mass
Leaf manganese (Mn) content per unit leaf dry mass
Leaf molybdenum (Mo) content per unit leaf dry mass
Leaf nitrogen (N) content per unit leaf area
Leaf nitrogen (N) content per unit leaf dry mass
Leaf sodium (Na) content per unit leaf dry mass
Leaf nickel (Ni) content per unit leaf dry mass
Leaf phosphorus (P) content per unit leaf area
Leaf phosphorus (P) content per unit leaf dry mass
Leaf sulphur (S) content per unit leaf dry mass
Leaf selenium (Se) content per unit leaf dry mass
Leaf silicon (Si) content per unit leaf dry mass
Leaf zinc (Zn) content per unit leaf dry mass
Leaf carbon to nitrogen ratio (C/N
Leaf nitrogen to phosphorus ratio (N/P) per unit leaf dry mass
Leaf cell wall nitrogen (N) per unit cell wall dry mass
Leaf cell wall nitrogen (N) per unit leaf N content
Lipid fraction of leaf P content per dry mass
Nucleic acid fraction of leaf P content per dry mass
Residual P fraction of leaf P content per dry mass
Inorganic P fraction of leaf P content per dry mass
Metabolite fraction of leaf P content per dry mass
Leaf ash content per unit leaf dry mass
Leaf photosynthetic nitrogen use efficiency during Amax measurement (PNUE
Leaf photosynthetic nitrogen use efficiency during Asat measurement (PNUE
Leaf photosynthetic phosphorus use efficiency during Amax measurement PPUE
Leaf photosynthetic phosphorus use efficiency during Asat measurement (PPUE
has broader leaf mineral and ion content trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000008

senesced leaf mineral and ion content trait

stem mineral and ion content trait

wood mineral and ion content trait

senesced wood mineral and ion content trait

bark mineral and ion content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000608
description A mineral and ion content trait which is measured on bark [PO:0004518].
has broader mineral and ion content trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000608

root mineral and ion content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0000808
description A mineral and ion content trait which is measured on a root [PO:0009005].
has narrower Root carbon (C) content per unit root dry mass
Root nitrogen (N) content per unit root dry mass
Root phosphorus (P) content per unit root dry mass
has broader mineral and ion content trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0000808

reproductive shoot system mineral and ion content trait

cell or tissue mineral and ion content trait

metabolite content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0002004
description The trait associated with an assay to determine the affected metabolites profile.
has narrower carbohydrate content trait
lipid content trait
phenolic compound content trait
pigment content trait
protein content trait
has broader biochemical trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0002004

carbohydrate content trait

lipid content trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0002006
description The total lipid [CHEBI:18059] content measured in a plant or a plant part.
has narrower Seed oil content per unit seed dry mass
has broader metabolite content trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0002006

phenolic compound content trait

pigment content trait

protein content trait

stable isotope ratio determination

plant morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0010001
description A plant trait [TO:0000387] which is a morphological quality of a plant anatomical entity [PO:0025131] or a constituent cellular component [GO:0005575] contained therein. Includes traits of plant structures [PO:0009011] such as plant cell [PO:0009002], plant organ [PO:0009008], portion of plant tissue [PO:0009007], vascular system [PO:0000034], whole plant [PO:0000003] and seed [PO:0009010], as well as those of plant anatomical spaces [PO:0025117] and portions of plant substance [PO:0025161].
has narrower whole plant morphology trait
plant structure morphology trait
vascular tissue morphology trait
plant structure strength trait
has broader plant trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0010001

whole plant morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0010004
description A plant structure morphology trait [TO:0000839] which is a quality of a whole plant [PO:0000003].
has narrower plant embryo morphology trait
Plant canopy width
Plant canopy breadth
Plant vegetative height
Height of a climbing plant
Plant reproductive height
Plant growth form
Life form
has broader plant morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0010004

plant embryo morphology trait

plant structure morphology trait

leaf morphology trait

leaf size trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0011204
description Refer to size [PATO:0000117]: A morphology quality inhering in a bearer by virtue of the bearers physical magnitude.
has narrower Leaf area
Leaflet area
Leaf length
Leaf width
Leaf thickness
Petiole length
Petiole width
Leaflet length
Leaflet width
Leaf mass per area
Leaf lamina mass per area
Leaf area ratio (LAR
Leaf fresh mass per leaf area
has broader leaf morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0011204

leaf mass trait

leaf shape trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0011304
description A leaf morphology trait [TO:0000748] which is the variation in shapes and forms of a leaf [PO:0025034]. Refer to shape [PATO:0000052]: A morphological quality inhering in a bearer by virtue of the bearers ratios of distances between its features [points, edges, surfaces and also holes etc].
has narrower Plant spinescence
Leaf type
Leaf shape
Leaf base shape
Leaf margin
Leaf margin posture
Leaf lobation
Leaf compoundness
Leaf divisions
Leaf lamina posture (leaf 3-dimensionality
Leaf lamina posture (leaf 3-dimensional shape
has broader leaf morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0011304

leaf position trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0011402
description A leaf morphology trait [TO:0000748] which refers to the position [PATO:0000140] of leaves [PO:0025034] relative to the stem [PO:0009047].
has narrower Leaf phyllotaxis
Leaf arrangement
Leaf axil angle
Leaf inclination angle
Leaf pendulousness
has broader leaf morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0011402

portion of plant tissue morphology trait

plant cell morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0011503
description A plant structure morphology trait [TO:0000839] which is a quality of a plant cell [PO:0009002].
has narrower Average leaf epidermal cell density
Lower leaf side epidermal cell density
Upper leaf side epidermal cell density
Palisade cell length
Palisade cell width
Cell cross-sectional area
has broader plant structure morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0011503

leaf stomatal complex morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0011602
description A vascular leaf morphology trait [TO:0000419] which is a quality of the stomatal complexes [PO:0002000] present on the leaf surface.
has narrower Stomatal density on the lower leaf surface
Stomatal density on the upper leaf surface
Stomatal density averaged across both leaf surfaces
Stomatal distribution
Stomatal hairiness
Guard cell length
has broader leaf morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0011602

leaf optical properties trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0011702
description Leaf morphology trait [TO:0000748] associated with the leafs optical [NCIT:C16940] properties, including the absorption [SWEET:soprocw:Absorption], transmission [SWEET:soprocw:Transmission], and reflection [SWEET:soprocw:Reflection] of light [OMIT:0009112].
has narrower Leaf visible light transmission
Leaf visible light absorption
Leaf visible light reflection
Leaf infra-red light reflection
has broader leaf morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0011702

stem morphology trait

stem mass trait

bark morphology trait

bark mass trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0012006
description A bark morphology trait which is the mass of bark.
has broader bark morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0012006

root system morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0012102
description A collective plant organ structure morphology trait [TO:0000842] which is a quality of a root system [PO:0025025].
has narrower root morphology trait
root mass trait
root branchiness trait
has broader plant structure morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0012102

root morphology trait

root mass trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0012106
description A root morphology trait [TO:0000043] which is the weight of a root [PO:0009005].
has broader root system morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0012106

root branchiness trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0012108
description A root system morphology trait [TO:0000985] which is the branchiness [PATO:0002009] of the root system [PO:0025025].
has broader root system morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0012108

reproductive shoot system morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0012202
description A shoot system morphology trait [TO:0000077] that is a quality of a reproductive shoot system [PO:0025082].
has narrower floral organ morphology trait
fruit morphology trait
seed morphology trait
Seed accessory cost fraction
Seed accessory cost mass
Life form
has broader plant structure morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0012202

floral organ morphology trait

perianth morphology trait

androecium morphology trait

gynoecium morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0012409
description A collective phyllome structure morphology trait [TO:0000845] that is a quality of the gynoecium [PO:0009062].
has narrower Gynoecium phyllotaxis
Placentation
Fusion of ovaries
Ovary position
Number of ovules per functional carpel
Number of structural carpels
Style differentiation
Fusion of styles
has broader floral organ morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0012409

fruit morphology trait

seed morphology trait

vascular tissue morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0013001
description A portion of plant tissue morphology trait [TO:0000843] which is a quality of a vascular tissue and its parts.
has narrower leaf vein morphology trait
xylem vessel morphology trait
Wood axial parenchyma fraction
Wood conduit fraction
Wood fibre fraction
Wood ray parenchyma fraction
Wood tracheid fraction
has broader plant morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0013001

leaf vein morphology trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0013006
description A vascular leaf morphology trait [TO:0000419] which is a quality of a leaf lamina vein [PO:0020138].
has narrower Leaf secondary vein angle
Major leaf vein density
Length of all minor and major leaf lamina veins per unit area
Leaf vein frequency
has broader vascular tissue morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0013006

xylem vessel morphology trait

plant structure strength trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0014001
description A plant structure morphology trait [TO:0000736] pertaining to the strength [PATO:0001230] of a plant structure [PO:0009011] and often indicating the amount of work [PATO:0001026] required to break or deform the plant structure.
has narrower Leaf work to punch
Leaf specific work to punch
Leaf work to shear
Leaf specific work to shear (fracture toughness
Leaf work to tear
Leaf specific work to tear
Bark modulus of elasticity
Stem modulus of elasticity
Xylem modulus of elasticity
Modulus of rupture
has broader plant morphology trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0014001

biological process trait, physiological process trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0020001
description Refers to process quality [PATO:0001236]: A quality which inheres in an process. A plant trait [TO:0000387] which is a process quality [PATO:0001236] of a biological process [GO:0008150] which occurs in a plant anatomical entity [PO:0025131], a cellular component [GO:0005575], or during a plant structure development stage [PO:0009012].
has narrower photosynthetic trait
water transport trait, hydraulic trait
nutrient recycling trait
has broader plant trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0020001

photosynthetic trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0020202
description A plant physiological process trait [TO:0000283] associated with photosynthesis [GO:0015979].
has narrower gas exchange trait
photosystem performance trait
Plant photosynthetic pathway
Plant photosynthetic organ
has broader biological process trait, physiological process trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0020202

gas exchange trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0020204
description A plant photosynthesis trait pertaining to the movement [NCIT:C37994] of carbon dioxide [CHEBI:16526], water [CHEBI:15377], and oxygen [CHEBI:15379] into and out of a plant cellular metabolic processes [GO:0044237] where these molecules are the reactant [SIO:000879] or product [SIO:010363].
has narrower photosynthetic rate trait
respiration rate trait
transpiration rate trait
carbon dioxide concentration trait
Leaf stomatal conductance to water vapour per unit leaf area under ambient conditions (gsw
Leaf stomatal conductance to water vapour per unit leaf area during Amax measurement (gsw
Leaf stomatal conductance to water vapour per unit leaf area during Asat measurement (gsw
Leaf stomatal water vapour resistance under ambient conditions
Leaf mesophyll conductance to carbon dioxide per unit leaf area (gm
Leaf mesophyll conductance to carbon dioxide per unit leaf mass (gm
Leaf cuticular conductance
Leaf conductance to surface water (ksurf
Leaf foliar water uptake
Leaf photosynthetic nitrogen use efficiency during Amax measurement (PNUE
Leaf photosynthetic nitrogen use efficiency during Asat measurement (PNUE
Leaf photosynthetic phosphorus use efficiency during Amax measurement PPUE
Leaf photosynthetic phosphorus use efficiency during Asat measurement (PPUE
Integrated water use efficiency
Intrinsic water use efficiency (WUEi
Instantaneous water use efficiency (WUE
has broader photosynthetic trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0020204

photosynthetic rate trait

respiration rate trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0020208
description A plant gas exchange trait which is the net rate of carbon dioxide [CHEBI:16526] released from a whole plant [PO:0000003] or plant organ [PO:0009008] due to cellular respiration [GO:0045333] and photorespiration [GO:0009853].
has narrower Leaf respiration rate per unit leaf area, in the dark (Rdark
Leaf respiration rate per unit leaf dry mass, in the dark (Rdark
Leaf respiration rate per unit leaf area, in the light (Rday
Stem respiration rate per unit stem area, in the dark
has broader gas exchange trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0020208

transpiration rate trait

carbon dioxide concentration trait

photosystem performance trait

water transport trait, hydraulic trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0021002
description A plant physiological process trait [TO:0000283] pertaining to the movement [NCIT:C37994] of water [CHEBI:15377] through a plants vascular system [PO:0000034], including water stress traits [TO:0000237].
has narrower Leaf secondary vein angle
Major leaf vein density
Length of all minor and major leaf lamina veins per unit area
Leaf vein frequency
Stem xylem vessel density
Leaf xylem vessel density
Stem xylem vessel diameter
Stem xylem vessel hydraulic mean diameter
Leaf xylem vessel diameter
Stem xylem vessel lumen fraction
Stem xylem vessel multiple fraction
Stem non-lumen fraction
xylem vessel wall fraction
Xylem vulnerability index
Leaf xylem vessel wall thickness
Stem xylem vessel wall thickness
Leaf xylem vessel length
Stem xylem vessel length
Wood axial parenchyma fraction
Wood conduit fraction
Wood fibre fraction
Wood ray parenchyma fraction
Wood tracheid fraction
Water band index
Stem hydraulic conductivity (Kh
Sapwood specific hydraulic conductivity (Ks
Theoretical sapwood specific hydraulic conductivity (Ks
Stem specific hydraulic conductivity (Ks
Leaf specific hydraulic conductance (kleaf
Leaf specific hydraulic conductivity (Kl
Root hydraulic conductivity (Kh
Root specific hydraulic conductivity
Pre-dawn water potential
Midday water potential
Stem sapwood capacitance (C
Leaf capacitance (Cleaf
Root sapwood capacitance (C
Leaf xylem pressure, 50% lost conductance (P50) (leaf hydraulic vulnerability
Stem xylem pressure, 12% lost conductivity (P12
Stem xylem pressure, 50% lost conductivity (P50
Stem xylem pressure, 88% lost conductivity (P88
Hydraulic safety margin, 50
Hydraulic safety margin, 88
Leaf turgor loss point
Osmotic potential
Osmotic potential at full tugor
Bulk modulus of elasticity (e
Leaf relative water content predawn
Leaf relative water content at turgor loss point
Root xylem pressure, 50% lost conductivity (P50
Root xylem pressure, 12% lost conductivity (P12
Root xylem pressure, 88% lost conductivity (P88
Leaf xylem pressure, 12% lost conductance (P12
Leaf xylem pressure, 88% lost conductance (P88
has broader biological process trait, physiological process trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0021002

nutrient recycling trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0022002
description A plant physiological process trait [TO:0000283] associated with the resorption [NCIT:C93204] and recycling [NCIT:C126395] of macronutrients [CHEBI:33937] and micronutrients [CHEBI:27027] as plant structures [PO:0009011] senesce [PATO:0001487].
has narrower Leaf nitrogen resorption efficiency
Leaf phosphorus resorption efficiency
has broader biological process trait, physiological process trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0022002

life history trait

whole plant phenotype trait

plant phenological trait

interspecific interactions trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0030009
description A plant life history trait that captures an interspecific interaction between organisms [GO:0044419].
has narrower Dispersal appendage
Plant nitrogen fixation capacity
Plant root structures
Plant parasitism status
Pollination syndrome
Pollination system
Diaspore dispersal syndrome
Diaspore dispersal agents
has broader life history trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0030009

genetic structure trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0030059
description A plant life history trait that captures information about genetic structures.
has narrower Plant genome size
Chromosome ploidy
has broader life history trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0030059

reproductive structure life history trait

fire response trait

chemical stress sensitivity trait

Property Value
URI https://w3id.org/APD/traits/trait_group_0030702
description Response by the plant in terms of resistivity or senility to chemical stress.
has narrower Plant soil salinity tolerance
Plant salt tolerance strategy
Plant calcium sensitivity
has broader life history trait
is in scheme https://w3id.org/APD/traits
identifier trait_group_0030702

environmental tolerance trait

3. Trait concepts

The APD Trait concepts express the traits defined by the APD.

Leaf aluminium (Al) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000012
alternative label leaf_Al_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of aluminium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0.01
maxAllowedValue 12000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword aluminium atom
scope note none
has exact match
has close match
has related match
examples exact match: Leaf aluminium content per leaf dry mass [TOP449] (https://top-thesaurus.org/index)
exact match: Leaf aluminium (Al) content per leaf dry mass [TRY:249] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000012

Leaf boron (B) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000014
alternative label leaf_B_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of boron in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0.01
maxAllowedValue 1000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword boron atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf boron content per leaf dry mass [TOP450] (https://top-thesaurus.org/index)
exact match: Leaf boron (B) content per leaf dry mass [TRY:250] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000014

Leaf carbon (C) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000016
alternative label leaf_C_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of carbon in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 50
maxAllowedValue 750
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon atom
scope note none
has exact match leaf carbon content
Leaf carbon content
has close match
has related match
examples exact match: Leaf carbon content per leaf dry mass [TOP452] (https://top-thesaurus.org/index)
exact match: Leaf carbon (C) content per leaf dry mass [TRY:13] (https://www.try-db.org/de/de.php)
exact match: leaf carbon content per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000016

Leaf calcium (Ca) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000018
alternative label leaf_Ca_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of calcium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 1000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf calcium content per leaf dry mass [TOP451] (https://top-thesaurus.org/index)
exact match: Leaf calcium (Ca) content per leaf dry mass [TRY:252] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000018

Leaf chlorine (Cl) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000020
alternative label leaf_Cl_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorine [CHEBI:23116] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of chlorine in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 10
maxAllowedValue 10000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword chlorine atom
scope note none
has exact match
has close match
has related match
examples exact match: Leaf chlorine content per leaf dry mass [TOP453] (https://top-thesaurus.org/index)
exact match: Leaf chlorine (Cl) content per leaf dry mass [TRY:539] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000020

Leaf chromium (Cr) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000022
alternative label leaf_Cr_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chromium [CHEBI:28073] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of chromium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit ppm
parts per million
minAllowedValue 0.01
maxAllowedValue 150
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword chromium atom
scope note none
has exact match
has close match
has related match
examples exact match: Leaf chromium (Cr) content per leaf dry mass [TRY:254] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000022

Leaf cobalt (Co) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000024
alternative label leaf_Co_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of cobalt [CHEBI:27638] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of cobalt in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit ppm
parts per million
minAllowedValue 0.01
maxAllowedValue 150
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword cobalt atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf cobalt content per leaf dry mass [TOP454] (https://top-thesaurus.org/index)
exact match: Leaf cobalt (Co) content per leaf dry mass [TRY:253] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000024

Leaf copper (Cu) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000026
alternative label leaf_Cu_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of copper in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0.01
maxAllowedValue 150
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword copper atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf copper content per leaf dry mass [TOP455] (https://top-thesaurus.org/index)
exact match: Leaf copper (Cu) content per leaf dry mass [TRY:255] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000026

Leaf iron (Fe) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000028
alternative label leaf_Fe_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of iron in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 1
maxAllowedValue 10000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword iron atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf iron content per leaf dry mass [TOP456] (https://top-thesaurus.org/index)
exact match: Leaf iron (Fe) content per leaf dry mass [TRY:256] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000028

Leaf potassium (K) content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0000029
alternative label leaf_K_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of potassium in a leaf to leaf area.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 10
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword potassium atom
macronutrient
scope note none
has exact match leaf potassium content
has close match
has related match
examples exact match: Leaf potassium content per leaf area [TOP483] (https://top-thesaurus.org/index)
exact match: Leaf potassium (K) content per leaf area [TRY:52] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000029

Leaf potassium (K) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000030
alternative label leaf_K_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of potassium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
macronutrient
scope note none
has exact match leaf potassium content
has close match
has related match
examples exact match: Leaf potassium content per leaf dry mass [TOP464] (https://top-thesaurus.org/index)
exact match: Leaf potassium (K) content per leaf dry mass [TRY:44] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000030

Leaf magnesium (Mg) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000032
alternative label leaf_Mg_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of magnesium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf magnesium content per leaf dry mass [TOP457] (https://top-thesaurus.org/index)
exact match: Leaf magnesium (Mg) content per leaf dry mass [TRY:257] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000032

Leaf manganese (Mn) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000034
alternative label leaf_Mn_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of manganese in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 1
maxAllowedValue 20000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword manganese atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf manganese content per leaf dry mass [TOP458] (https://top-thesaurus.org/index)
exact match: Leaf manganese (Mn) content per leaf dry mass [TRY:258] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000034

Leaf molybdenum (Mo) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000036
alternative label leaf_Mo_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of molybdenum [CHEBI:28685] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of molybdenum in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0
maxAllowedValue 100
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword molybdenum atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf molybdenum content per leaf dry mass [TOP459] (https://top-thesaurus.org/index)
exact match: Leaf molybdenum (Mo) content per leaf dry mass [TRY:259] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000036

Leaf nitrogen (N) content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0000037
alternative label leaf_N_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of nitrogen in a leaf to leaf area.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.08
maxAllowedValue 50
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword nitrogen atom
macronutrient
scope note none
has exact match Leaf nitrogen content
leaf nitrogen content
leaf nitrogen concentration
has close match
has related match
examples exact match: Leaf nitrogen content per leaf area [TOP481] (https://top-thesaurus.org/index)
exact match: Leaf nitrogen (N) content per leaf area [TRY:50] (https://www.try-db.org/de/de.php)
exact match: leaf nitrogen content per leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000037

Leaf nitrogen (N) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000038
alternative label leaf_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of nitrogen in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.4
maxAllowedValue 100
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
scope note none
has exact match Leaf nitrogen content
leaf nitrogen content
leaf nitrogen concentration
has close match
has related match
examples exact match: Leaf nitrogen content per leaf dry mass [TOP462] (https://top-thesaurus.org/index)
exact match: Leaf nitrogen (N) content per leaf dry mass [TRY:14] (https://www.try-db.org/de/de.php)
exact match: leaf nitrogen content per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Mass-based leaf nitrogen content [BROT:7] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000038

Leaf sodium (Na) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000040
alternative label leaf_Na_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of sodium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 1
maxAllowedValue 20000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword sodium atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf sodium content per leaf dry mass [TOP460] (https://top-thesaurus.org/index)
exact match: Leaf sodium (Na) content per leaf dry mass [TRY:260] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000040

Leaf nickel (Ni) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000042
alternative label leaf_Ni_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nickel [CHEBI:28112] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of nickel in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0.1
maxAllowedValue 100
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nickel atom
scope note none
has exact match
has close match
has related match
examples exact match: Leaf nickel content per leaf dry mass [TOP461] (https://top-thesaurus.org/index)
exact match: Leaf nickel (Ni) content per leaf dry mass [TRY:261] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000042

Leaf phosphorus (P) content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0000043
alternative label leaf_P_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of phosphorus in a leaf to leaf area.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.005
maxAllowedValue 1.2
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword phosphorus atom
macronutrient
scope note none
has exact match leaf phosphorus content
has close match
has related match
examples exact match: Leaf phosphorus content per leaf area [TOP482] (https://top-thesaurus.org/index)
exact match: Leaf phosphorus (P) content per leaf area [TRY:51] (https://www.try-db.org/de/de.php)
exact match: leaf phosphorus content per leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000043

Leaf phosphorus (P) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000044
alternative label leaf_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of phosphorus in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.03
maxAllowedValue 12
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
macronutrient
scope note none
has exact match leaf phosphorus content
has close match
has related match
examples exact match: Leaf phosphorus content per leaf dry mass [TOP463] (https://top-thesaurus.org/index)
exact match: Leaf phosphorus (P) content per leaf dry mass [TRY:15] (https://www.try-db.org/de/de.php)
exact match: leaf phosphorus content per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000044

Leaf sulphur (S) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000046
alternative label leaf_S_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of sulphur in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 50
maxAllowedValue 50000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword sulfur atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf sulfur content per leaf dry mass [TOP466] (https://top-thesaurus.org/index)
exact match: Leaf sulfur (S) content per leaf dry mass [TRY:262] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000046

Leaf selenium (Se) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000048
alternative label leaf_Se_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of selenium [CHEBI:27568] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of selenium in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit ppm
parts per million
minAllowedValue 0.01
maxAllowedValue 100
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword selenium atom
scope note none
has exact match
has close match
has related match
examples exact match: Leaf selenium (Se) content per leaf dry mass [TRY:263] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000048

Leaf silicon (Si) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000050
alternative label leaf_Si_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of silicon [CHEBI:27573] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of silicon in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades. Leaf silicon (Si) must not be confused with leaf silica (SiO2). In plants, Si is mostly found as hydrated amorphous silica (SiO2 , nH2O).
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 200
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword silicon atom
scope note none
has exact match
has close match
has related match
examples exact match: Leaf silicon content per leaf dry mass [TOP465] (https://top-thesaurus.org/index)
exact match: Leaf silicon (Si) content per leaf dry mass [TRY:264] (https://www.try-db.org/de/de.php)
references Cooke & Leishman 2011
de Tombeur 2023
date created 14/07/2021
date modified 05/02/2024
date reviewed 13/02/2023
reviewed by Félix de Tombeur
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000050

Leaf zinc (Zn) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000052
alternative label leaf_Zn_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of zinc in a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0.1
maxAllowedValue 1000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword zinc atom
micronutrient
scope note none
has exact match
has close match
has related match
examples exact match: Leaf zinc content per leaf dry mass [TOP467] (https://top-thesaurus.org/index)
exact match: Leaf zinc (Zn) content per leaf dry mass [TRY:268] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000052

Leaf carbon to nitrogen ratio (C/N)

Property Value
URI https://w3id.org/APD/traits/trait_0000090
alternative label leaf_CN_ratio
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a leaf [PO:0025034] to the mass of nitrogen [CHEBI:25555] in the leaf.
The ratio of the mass of carbon in a leaf to the mass of nitrogen in the leaf.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 10
maxAllowedValue 1000
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
dimensionless ratio
ratio; proportion
keyword carbon atom
macronutrient
nitrogen atom
scope note none
has exact match Ratio leaf nitrogen to leaf carbon
C/N
has close match
has related match
examples exact match: Leaf carbon/nitrogen (C/N) ratio [TRY:146] (https://www.try-db.org/de/de.php)
exact match: leaf carbon content per leaf nitrogen content [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000090

Leaf nitrogen to phosphorus ratio (N/P) per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000091
alternative label leaf_NP_ratio
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to the mass of phosphorus [CHEBI:28659] in the leaf.
The ratio of the mass of nitrogen in a leaf to the mass of phosphorus in the leaf.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 1
maxAllowedValue 100
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
dimensionless ratio
ratio; proportion
keyword nitrogen atom
macronutrient
phosphorus atom
scope note
has exact match
has close match
has related match
examples exact match: Leaf nitrogen/phosphorus (N/P) ratio [TRY:56] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note N_to_P_ratio
is in scheme https://w3id.org/APD/traits
identifier trait_0000091

Senesced leaf aluminium (Al) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000112
alternative label leaf_senesced_Al_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of aluminium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword aluminium atom
senescent
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter aluminium content per litter dry mass [TOP583] (https://top-thesaurus.org/index)
exact match: Litter aluminium (Al ) content per litter dry mass [TRY:987] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Al_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000112

Senesced leaf boron (B) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000114
alternative label leaf_senesced_B_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of boron in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword boron atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter boron content per litter dry mass [TOP584] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_B_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000114

Senesced leaf carbon (C) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000116
alternative label leaf_senesced_C_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of carbon in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.1
maxAllowedValue 1
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon atom
senescent
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter carbon content per litter dry mass [TOP586] (https://top-thesaurus.org/index)
exact match: Litter carbon (C) content per litter dry mass [TRY:72] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_C_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000116

Senesced leaf calcium (Ca) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000118
alternative label leaf_senesced_Ca_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of calcium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 1000
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter calcium content per litter dry mass [TOP585] (https://top-thesaurus.org/index)
exact match: Litter (leaf) calcium (Ca) content per leaf litter dry mass [TRY:943] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Ca_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000118

Senesced leaf copper (Cu) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000126
alternative label leaf_senesced_Cu_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of copper in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0.01
maxAllowedValue 100
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword copper atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter copper content per litter dry mass [TOP589] (https://top-thesaurus.org/index)
exact match: Litter (leaf) copper (Cu) content per leaf litter dry mass [TRY:947] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Cu_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000126

Senesced leaf iron (Fe) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000128
alternative label leaf_senesced_Fe_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of iron in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 1
maxAllowedValue 10000
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword iron atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter iron content per litter dry mass [TOP590] (https://top-thesaurus.org/index)
exact match: Litter (leaf) iron (Fe) content per leaf litter dry mass [TRY:950] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Fe_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000128

Senesced leaf potassium (K) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000130
alternative label leaf_senesced_K_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of potassium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
senescent
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter potassium content per litter dry mass [TOP598] (https://top-thesaurus.org/index)
exact match: Litter (leaf) potassium (K) content per leaf litter dry mass [TRY:958] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_K_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000130

Senesced leaf magnesium (Mg) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000132
alternative label leaf_senesced_Mg_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of magnesium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 100
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter magnesium content per litter dry mass [TOP591] (https://top-thesaurus.org/index)
exact match: Litter (leaf) magnesium (Mg) content per leaf litter dry mass [TRY:952] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Mg_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000132

Senesced leaf manganese (Mn) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000134
alternative label leaf_senesced_Mn_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of manganese in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 1
maxAllowedValue 10000
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword manganese atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter manganese content per litter dry mass [TOP592] (https://top-thesaurus.org/index)
exact match: Litter (leaf) manganese (Mn) content per leaf litter dry mass [TRY:953] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Mn_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000134

Senesced leaf molybdenum (Mo) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000136
alternative label leaf_senesced_Mo_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of molybdenum [CHEBI:28685] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of molybdenum in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0
maxAllowedValue 30
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword molybdenum atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter molybdenum content per litter dry mass [TOP593] (https://top-thesaurus.org/index)
exact match: Litter molybdenium (Mo) content per litter dry mass [TRY:999] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Mo_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000136

Senesced leaf nitrogen (N) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000138
alternative label leaf_senesced_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of nitrogen in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 100
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
senescent
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter nitrogen content per litter dry mass [TOP596] (https://top-thesaurus.org/index)
exact match: Litter (leaf) nitrogen (N) content per leaf litter dry mass [TRY:954] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_N_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000138

Senesced leaf sodium (Na) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000140
alternative label leaf_senesced_Na_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of sodium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 10
maxAllowedValue 20000
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword sodium atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter sodium content per litter dry mass [TOP594] (https://top-thesaurus.org/index)
exact match: Litter sodium (Na) content per litter dry mass [TRY:1004] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Na_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000140

Senesced leaf nickel (Ni) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000142
alternative label leaf_senesced_Ni_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nickel [CHEBI:28112] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of nickel in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 10
maxAllowedValue 1e+05
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nickel atom
senescent
plant litter
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Ni_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000142

Senesced leaf phosphorus (P) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000144
alternative label leaf_senesced_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of phosphorus in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.005
maxAllowedValue 15
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
senescent
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter phosphorus content per litter dry mass [TOP597] (https://top-thesaurus.org/index)
exact match: Litter (leaf) phosphorus (P) content per leaf litter dry mass [TRY:957] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_P_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000144

Senesced leaf sulphur (S) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000146
alternative label leaf_senesced_S_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of sulphur in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 100
maxAllowedValue 50000
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword sulfur atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter sulfur content per litter dry mass [TOP600] (https://top-thesaurus.org/index)
exact match: Litter (leaf) sulfur (S) content per leaf litter dry mass [TRY:961] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_S_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000146

Senesced leaf zinc (Zn) content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000152
alternative label leaf_senesced_Zn_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014].
The ratio of the mass of zinc in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/kg
milligram per kilogram
minAllowedValue 0
maxAllowedValue 100
has broader senesced leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword zinc atom
senescent
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Litter zinc content per litter dry mass [TOP601] (https://top-thesaurus.org/index)
exact match: Litter (leaf) zinc (Zn) content per leaf litter dry mass [TRY:962] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note senesced_leaf_Zn_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000152

Leaf nitrogen resorption efficiency

Property Value
URI https://w3id.org/APD/traits/trait_0022012
alternative label leaf_N_resorption
description A leaf [PO:0025034] nutrient recycling trait which is one minus the ratio [PATO:0001470] of leaf nitrogen [CHEBI:25555] content in a senescent [PATO:0001487] leaf (leaf litter [ENVO:01000628]) to leaf nitrogen content in a live [PATO:0001421] leaf, indicating what proportion of nitrogen is resorbed [NCIT:C93204] from the senescent leaf before it is shed.
A leaf nutrient recycling trait which is one minus the ratio of leaf nitrogen content in a senescent leaf (leaf litter) to leaf nitrogen content in a live leaf, indicating what proportion of nitrogen is resorbed from the senescent leaf before it is shed.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue -1.1
maxAllowedValue 1
has broader nutrient recycling trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
dimensionless ratio
keyword nitrogen atom
senescent
resorption
plant litter
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note resorption_leaf_N
is in scheme https://w3id.org/APD/traits
identifier trait_0022012

Leaf phosphorus resorption efficiency

Property Value
URI https://w3id.org/APD/traits/trait_0022013
alternative label leaf_P_resorption
description A leaf [PO:0025034] nutrient recycling trait which is one minus the ratio [PATO:0001470] of leaf phosphorus [CHEBI:28659] content in a senescent [PATO:0001487] leaf (leaf litter [ENVO:01000628]) to leaf phosphorus content in a live [PATO:0001421] leaf, indicating what proportion of phosphorus is resorbed [NCIT:C93204] from the senescent leaf before it is shed.
A leaf nutrient recycling trait which is one minus the ratio of leaf phosphorus content in a senescent leaf (leaf litter) to leaf phosphorus content in a live leaf, indicating what proportion of phosphorus is resorbed from the senescent leaf before it is shed.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue -1.1
maxAllowedValue 1
has broader nutrient recycling trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
dimensionless ratio
keyword phosphorus atom
senescent
resorption
plant litter
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note resorption_leaf_P
is in scheme https://w3id.org/APD/traits
identifier trait_0022013

Stem carbon (C) content per unit stem dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000216
alternative label stem_C_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a stem [PO:0009047] to stem dry mass.
The ratio of the mass of carbon in a stem to stem dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 100
maxAllowedValue 1000
has broader stem mineral and ion content trait
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon atom
scope note
has exact match stem carbon content
has close match
has related match
examples exact match: Stem carbon content per stem dry mass [TOP491] (https://top-thesaurus.org/index)
exact match: Stem carbon (C) content per stem dry mass [TRY:407] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000216

Stem nitrogen (N) content per unit stem dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000238
alternative label stem_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a stem [PO:0009047] to stem dry mass.
The ratio of the mass of nitrogen in a stem to stem dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 1000
has broader stem mineral and ion content trait
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
scope note
has exact match
has close match
has related match
examples exact match: Stem nitrogen content per stem dry mass [TOP501] (https://top-thesaurus.org/index)
exact match: Stem nitrogen (N) content per stem dry mass [TRY:406] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000238

Wood carbon (C) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000416
alternative label wood_C_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of carbon in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 50
maxAllowedValue 750
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon atom
wood
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood carbon content per wood dry mass trait [TOP732] (https://top-thesaurus.org/index)
close match: Stem carbon (C) content per stem dry mass [TRY:407] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000416

Wood calcium (Ca) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000418
alternative label wood_Ca_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of calcium in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
wood
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood calcium content per wood dry mass trait [TOP731] (https://top-thesaurus.org/index)
close match: Stem calcium (Ca) content per stem dry mass [TRY:623] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000418

Wood potassium (K) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000430
alternative label wood_K_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of potassium in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 100
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
wood
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood potassium content per wood dry mass trait [TOP744] (https://top-thesaurus.org/index)
close match: Stem potassium (K) content per stem dry mass [TRY:628] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000430

Wood magnesium (Mg) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000432
alternative label wood_Mg_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of magnesium in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
wood
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood magnesium content per wood dry mass trait [TOP737] (https://top-thesaurus.org/index)
close match: Stem magnesium (Mg) content per stem dry mass [TRY:624] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000432

Wood nitrogen (N) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000438
alternative label wood_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of nitrogen in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
wood
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood nitrogen content per wood dry mass trait [TOP742] (https://top-thesaurus.org/index)
exact match: Wood nitrogen (N) content per wood dry mass [TRY:1229] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000438

Wood sodium (Na) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000440
alternative label wood_Na_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of sodium in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword sodium atom
wood
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood sodium content per wood dry mass trait [TOP740] (https://top-thesaurus.org/index)
close match: Stem sodium (Na) content per stem dry mass [TRY:627] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000440

Wood phosphorus (P) content per unit wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000444
alternative label wood_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in wood [PO:0005848] to wood dry mass.
The ratio of the mass of phosphorus in wood to wood dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
wood
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Wood phosphorus content per wood dry mass trait [TOP743] (https://top-thesaurus.org/index)
close match: Stem phosphorus (P) content per stem dry mass [TRY:616] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000444

Dead wood calcium (Ca) content per unit dead wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000518
alternative label wood_dead_Ca_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.
The ratio of the mass of calcium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader senesced wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
senescent
wood
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Coarse woody debris calcium content per coarse woody debris dry mass trait [TOP750] (https://top-thesaurus.org/index)
close match: Litter (other than leaf: twigs, bark, flowers…) calcium (Ca) content per litter dry mass [TRY:2892] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note dead_wood_Ca_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000518

Dead wood potassium (K) content per unit dead wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000530
alternative label wood_dead_K_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.
The ratio of the mass of potassium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader senesced wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
senescent
wood
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Coarse woody debris potassium content per coarse woody debris dry mass trait [TOP763] (https://top-thesaurus.org/index)
close match: Litter (other than leaf: twigs, bark, flowers…) potassium (K) content per litter dry mass [TRY:2891] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note dead_wood_K_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000530

Dead wood magnesium (Mg) content per unit dead wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000532
alternative label wood_dead_Mg_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.
The ratio of the mass of magnesium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader senesced wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
senescent
wood
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Coarse woody debris magnesium content per coarse woody debris dry mass trait [TOP756] (https://top-thesaurus.org/index)
close match: Litter (other than leaf: twigs, bark, flowers…) magnesium (Mg) content per litter dry mass [TRY:2893] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note dead_wood_Mg_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000532

Dead wood nitrogen (N) content per unit dead wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000538
alternative label wood_dead_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.
The ratio of the mass of nitrogen in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader senesced wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
senescent
wood
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Coarse woody debris nitrogen content per coarse woody debris dry mass trait [TOP761] (https://top-thesaurus.org/index)
close match: Litter twig nitrogen (N) content per twig dry mass [TRY:1199] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note dead_wood_N_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000538

Dead wood sodium (Na) content per unit dead wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000540
alternative label wood_dead_Na_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.
The ratio of the mass of sodium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 1
has broader senesced wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword sodium atom
senescent
wood
micronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Coarse woody debris sodium content per coarse woody debris dry mass trait [TOP759] (https://top-thesaurus.org/index)
close match: Litter (other than leaf: twigs, bark, flowers…) sodium (Na) content per litter dry mass [TRY:2898] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note dead_wood_Na_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000540

Dead wood phosphorus (P) content per unit dead wood dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000544
alternative label wood_dead_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood.
The ratio of the mass of phosphorus in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader senesced wood mineral and ion content trait
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
senescent
wood
macronutrient
plant litter
scope note
has exact match
has close match
has related match
examples exact match: Coarse woody debris phosphorus content per coarse woody debris dry mass trait [TOP762] (https://top-thesaurus.org/index)
close match: Litter other than leaf phosphorus (P) content per litter dry mass [TRY:2890] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note dead_wood_P_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0000544

Bark aluminium (Al) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000612
alternative label bark_Al_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of aluminium in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword aluminium atom
scope note
has exact match
has close match
has related match
examples exact match: Bark aluminium content per bark dry mass [TOP373] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000612

Bark boron (B) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000614
alternative label bark_B_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of boron in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 1
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword boron atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark boron content per bark dry mass [TOP374] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000614

Bark carbon (C) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000616
alternative label bark_C_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of carbon in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 50
maxAllowedValue 750
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon atom
scope note
has exact match
has close match
has related match
examples exact match: Bark carbon content per bark dry mass [TOP376] (https://top-thesaurus.org/index)
exact match: Bark carbon (C) content per bark dry mass [TRY:617] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000616

Bark calcium (Ca) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000618
alternative label bark_Ca_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of calcium in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark calcium content per bark dry mass [TOP375] (https://top-thesaurus.org/index)
exact match: Bark calcium (Ca) content per bark dry mass [TRY:2957] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000618

Bark copper (Cu) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000626
alternative label bark_Cu_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of copper in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1e-04
maxAllowedValue 1
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword copper atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark copper content per bark dry mass [TOP379] (https://top-thesaurus.org/index)
exact match: Bark copper (Cu) content per bark dry mass [TRY:2958] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000626

Bark iron (Fe) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000628
alternative label bark_Fe_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of iron in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword iron atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark iron content per bark dry mass [TOP380] (https://top-thesaurus.org/index)
exact match: Bark iron (Fe) content per bark dry mass [TRY:2959] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000628

Bark potassium (K) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000630
alternative label bark_K_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of potassium in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 100
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
macronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark potassium content per bark dry mass [TOP388] (https://top-thesaurus.org/index)
exact match: Bark potassium (K) content per bark dry mass [TRY:2960] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000630

Bark magnesium (Mg) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000632
alternative label bark_Mg_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of magnesium in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark magnesium content per bark dry mass [TOP381] (https://top-thesaurus.org/index)
exact match: Bark magnesium (Mg) content per bark dry mass [TRY:2961] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000632

Bark manganese (Mn) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000634
alternative label bark_Mn_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of manganese in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 1
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword manganese atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark manganese content per bark dry mass [TOP382] (https://top-thesaurus.org/index)
exact match: Bark manganese (Mn) content per bark dry mass [TRY:2962] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000634

Bark nitrogen (N) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000638
alternative label bark_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of nitrogen in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark nitrogen content per bark dry mass [TOP386] (https://top-thesaurus.org/index)
exact match: Bark nitrogen (N) content per bark dry mass [TRY:618] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000638

Bark sodium (Na) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000640
alternative label bark_Na_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of sodium in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 100
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword sodium atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark sodium content per bark dry mass [TOP384] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000640

Bark phosphorus (P) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000644
alternative label bark_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of phosphorus in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
macronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark phosphorus content per bark dry mass [TOP387] (https://top-thesaurus.org/index)
exact match: Bark phosphorus (P) content per bark dry mass [TRY:619] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000644

Bark sulphur (S) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000646
alternative label bark_S_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of sulphur in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword sulfur atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark sulfur content per bark dry mass [TOP390] (https://top-thesaurus.org/index)
exact match: Bark sulfur (S) content per bark dry mass [TRY:2963] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000646

Bark zinc (Zn) content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000652
alternative label bark_Zn_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of zinc in bark to bark dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader stem mineral and ion content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword zinc atom
micronutrient
scope note
has exact match
has close match
has related match
examples exact match: Bark zinc content per bark dry mass [TOP391] (https://top-thesaurus.org/index)
exact match: Bark zinc (Zn) content per bark dry mass [TRY:2964] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000652

Root carbon (C) content per unit root dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000816
alternative label root_C_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in roots [PO:0009005] to root dry mass.
The ratio of the mass of carbon in roots to root dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 50
maxAllowedValue 750
has broader root mineral and ion content trait
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon atom
scope note
has exact match root carbon content
has close match
has related match
examples exact match: Root carbon content per root dry mass trait [TOP713] (https://top-thesaurus.org/index)
exact match: Root carbon (C) content per root dry mass [TRY:84] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000816

Root nitrogen (N) content per unit root dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000838
alternative label root_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in roots [PO:0009005] to root dry mass.
The ratio of the mass of nitrogen in roots to root dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader root mineral and ion content trait
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
scope note
has exact match root nitrogen content
has close match
has related match
examples exact match: Root nitrogen content per root dry mass trait [TOP723] (https://top-thesaurus.org/index)
exact match: Root nitrogen (N) content per root dry mass [TRY:80] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000838

Root phosphorus (P) content per unit root dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0000844
alternative label root_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in roots [PO:0009005] to root dry mass.
The ratio of the mass of phosphorus in roots to root dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 100
has broader root mineral and ion content trait
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
macronutrient
scope note
has exact match root system phosphorus content
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0000844

Flower nitrogen (N) content per unit flower dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001038
alternative label flower_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in flowers [PO:0009046] to flower dry mass.
The ratio of the mass of nitrogen in flowers to flower dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.5
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
flower
measured characteristic mass
mass fraction
ratio; proportion
keyword reproduction
nitrogen atom
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001038

Fruit calcium (Ca) content per unit fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001118
alternative label fruit_Ca_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in fruit [PO:0009001] to fruit dry mass.
The ratio of the mass of calcium in fruit to fruit dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.5
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
micronutrient
reproduction
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001118

Fruit potassium (K) content per unit fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001130
alternative label fruit_K_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in fruit [PO:0009001] to fruit dry mass.
The ratio of the mass of potassium in fruit to fruit dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.5
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
macronutrient
reproduction
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001130

Fruit magnesium (Mg) content per unit fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001132
alternative label fruit_Mg_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in fruit [PO:0009001] to fruit dry mass.
The ratio of the mass of magnesium in fruit to fruit dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.05
maxAllowedValue 10
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
micronutrient
reproduction
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001132

Fruit nitrogen (N) content per unit fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001138
alternative label fruit_N_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in fruit [PO:0009001] to fruit dry mass.
The ratio of the mass of nitrogen in fruit to fruit dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.5
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
reproduction
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001138

Fruit phosphorus (P) content per unit fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001144
alternative label fruit_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in fruit [PO:0009001] to fruit dry mass.
The ratio of the mass of phosphorus in fruit to fruit dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
macronutrient
reproduction
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001144

Fruit sulphur (S) content per unit fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001146
alternative label fruit_S_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in fruit [PO:0009001] to fruit dry mass.
The ratio of the mass of sulphur in fruit to fruit dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.05
maxAllowedValue 10
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
mass fraction
ratio; proportion
keyword sulfur atom
micronutrient
reproduction
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001146

Seed calcium (Ca) content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001218
alternative label seed_Ca_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in seeds [PO:0009010] to seed dry mass.
The ratio of the mass of calcium in seeds to seed dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
micronutrient
reproduction
scope note
has exact match
has close match
has related match
examples exact match: Seed calcium content per seed dry mass [TOP566] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_Ca_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0001218

Seed potassium (K) content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001230
alternative label seed_K_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in seeds [PO:0009010] to seed dry mass.
The ratio of the mass of potassium in seeds to seed dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword potassium atom
macronutrient
reproduction
scope note
has exact match
has close match
has related match
examples exact match: Seed potassium content per seed dry mass [TOP579] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_K_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0001230

Seed magnesium (Mg) content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001232
alternative label seed_Mg_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in seeds [PO:0009010] to seed dry mass.
The ratio of the mass of magnesium in seeds to seed dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword magnesium atom
micronutrient
reproduction
scope note
has exact match
has close match
has related match
examples exact match: Seed magnesium content per seed dry mass [TOP572] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_Mg_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0001232

Seed nitrogen (N) content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001238
alternative label seed_N_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in seeds [PO:0009010] to seed dry mass.
The ratio of the mass of nitrogen in seeds to seed dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 10
maxAllowedValue 1000
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
reproduction
scope note
has exact match concentration of nitrogen in seeds
has close match
has related match
examples exact match: Seed nitrogen content per seed dry mass [TOP577] (https://top-thesaurus.org/index)
exact match: Seed nitrogen (N) content per dry mass (or diaspore) [TRY:2945] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_N_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0001238

Seed phosphorus (P) content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001244
alternative label seed_P_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in seeds [PO:0009010] to seed dry mass.
The ratio of the mass of phosphorus in seeds to seed dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
macronutrient
reproduction
scope note
has exact match seed phosphorus content
has close match
has related match
examples exact match: Seed phosphorus content per seed dry mass [TOP578] (https://top-thesaurus.org/index)
exact match: Seed phosphorus (P) content per dry mass (or diaspore) [TRY:3000] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_P_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0001244

Seed sulphur (S) content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001246
alternative label seed_S_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in seeds [PO:0009010] to seed dry mass.
The ratio of the mass of sulphur in seeds to seed dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader reproductive shoot system mineral and ion content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword sulfur atom
micronutrient
reproduction
scope note
has exact match
has close match
has related match
examples exact match: Seed sulfur content per seed dry mass [TOP581] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_S_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0001246

Leaf cell wall nitrogen (N) per unit cell wall dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001511
alternative label leaf_cell_wall_N_per_cell_wall_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in the plant cell wall [GO:0009505] in a leaf [PO:0025034] to the cell wall dry mass in the leaf.
The ratio of the mass of nitrogen in the plant cell wall in a leaf to the cell wall dry mass in the leaf.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mmol{N}/g
micromole per gram
minAllowedValue 0.1
maxAllowedValue 10
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
cell
plant-type cell wall
scope note
has exact match
has close match
has related match
examples exact match: Leaf cell wall nitrogen (N) content per cell wall dry mass [TRY:3376] (https://www.try-db.org/de/de.php)
close match: Leaf cell wall nitrogen (N) content per leaf area [TRY:3377] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note leaf_cell_wall_N
is in scheme https://w3id.org/APD/traits
identifier trait_0001511

Leaf cell wall nitrogen (N) per unit leaf N content

Property Value
URI https://w3id.org/APD/traits/trait_0001512
alternative label leaf_cell_wall_N_per_leaf_N
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in the plant cell wall [GO:0009505] in a leaf [PO:0025034] to the mass of nitrogen in the leaf.
The ratio of the mass of nitrogen in the plant cell wall in a leaf to the mass of nitrogen in the leaf.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.01
maxAllowedValue 1
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
cell
plant-type cell wall
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note leaf_cell_wall_N_fraction
is in scheme https://w3id.org/APD/traits
identifier trait_0001512

Leaf rubisco nitrogen (N) content per unit leaf N content

Property Value
URI https://w3id.org/APD/traits/trait_0001513
alternative label leaf_rubisco_N_per_total_leaf_N
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in rubisco [MESH:D012273] in a leaf [PO:0025034] to the mass of nitrogen in a leaf.
The ratio of the mass of nitrogen in rubisco in a leaf to the mass of nitrogen in a leaf.
comments This trait is most commonly measured on leaf tissue. If the Rubisco concentration of a different tissue is measured, this should be explicitly noted in the methods or as a method context.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.05
maxAllowedValue 0.5
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
cell
protein
Ribulose-Bisphosphate Carboxylase
chloroplast
scope note
has exact match
has close match
has related match rubisco to nitrogen content ratio
examples exact match: Leaf nitrogen (N) fraction in Rubisco [TRY:187] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note cell_rubisco_N_per_total_N
is in scheme https://w3id.org/APD/traits
identifier trait_0001513

Leaf thylakoid protein nitrogen (N) content per unit leaf N content

Property Value
URI https://w3id.org/APD/traits/trait_0001514
alternative label leaf_thylakoid_N_per_total_leaf_N
description The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in thylakoid [GO:0009579] proteins [CHEBI:36080] in a leaf [PO:0025034] to the mass of nitrogen in a leaf.
The ratio of the mass of nitrogen in thylakoid proteins in a leaf to the mass of nitrogen in a leaf.
comments This trait is most commonly measured on leaf tissue. If the Rubisco concentration of a different tissue is measured, this should be explicitly noted in the methods or as a method context.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.05
maxAllowedValue 0.3
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword nitrogen atom
macronutrient
cell
protein
thylakoid
chloroplast
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note cell_thylakoid_N_per_total_N
is in scheme https://w3id.org/APD/traits
identifier trait_0001514

Leaf epidermis calcium (Ca) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001611
alternative label leaf_epidermis_Ca_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in epidermal cells [PO:0004013] to epidermal cell fresh mass [APD:glossary/40044].
The ratio of the mass of calcium in epidermal cells to epidermal cell fresh mass.
comments
value type continuous variable
unit umol{Ca}/g
micromole per gram
minAllowedValue 1
maxAllowedValue 500
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
epidermal cell
cell
micronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_epidermis_Ca_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001611

Leaf hypodermis calcium (Ca) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001612
alternative label leaf_hypodermis_Ca_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in the hypodermis [PO:0005051] to hypodermis fresh mass [APD:glossary/40044].
The ratio of the mass of calcium in the hypodermis to hypodermis fresh mass.
comments
value type continuous variable
unit umol{Ca}/g
micromole per gram
minAllowedValue 1
maxAllowedValue 500
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
hypodermis
cell
micronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_hypodermis_Ca_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001612

Leaf internal parenchyma cell calcium (Ca) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001613
alternative label leaf_internal_parenchyma_Ca_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in parenchyma cells [PO:0000074] to parenchyma cell fresh mass [APD:glossary/40044].
The ratio of the mass of calcium in parenchyma cells to parenchyma cell fresh mass.
comments
value type continuous variable
unit umol{Ca}/g
micromole per gram
minAllowedValue 1
maxAllowedValue 500
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
parenchyma cell
cell
micronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_internal_parenchyma_Ca_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001613

Leaf palisade mesophyll cell calcium (Ca) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001614
alternative label leaf_palisade_mesophyll_Ca_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in palisade mesophyll cells [PO:0006206] to palisade mesophyll cell fresh mass [APD:glossary/40044].
The ratio of the mass of calcium in palisade mesophyll cells to palisade mesophyll cell fresh mass.
comments
value type continuous variable
unit umol{Ca}/g
micromole per gram
minAllowedValue 1
maxAllowedValue 100
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
palisade mesophyll cell
cell
micronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_palisade_mesophyll_Ca_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001614

Leaf sclerenchyma cell calcium (Ca) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001615
alternative label leaf_sclerenchyma_Ca_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in sclerenchyma cells [PO:0000077] to sclerenchyma cell fresh mass [APD:glossary/40044].
The ratio of the mass of calcium in sclerenchyma cells to sclerenchyma cell fresh mass.
comments
value type continuous variable
unit umol{Ca}/g
micromole per gram
minAllowedValue 1
maxAllowedValue 100
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
sclerenchyma cell
cell
micronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_sclerenchyma_Ca_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001615

Leaf spongy mesophyll cell calcium (Ca) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001616
alternative label leaf_spongy_mesophyll_Ca_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in spongy mesophyll cells [PO:0006205] to spongy mesophyll cell fresh mass [APD:glossary/40044].
The ratio of the mass of calcium in spongy mesophyll cells to spongy mesophyll cell fresh mass.
comments
value type continuous variable
unit umol{Ca}/g
micromole per gram
minAllowedValue 1
maxAllowedValue 1000
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword calcium atom
spongy mesophyll cell
cell
micronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_spongy_mesophyll_Ca_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001616

Leaf epidermis phosphorus (P) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001661
alternative label leaf_epidermis_P_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in epidermal cells [PO:0004013] to epidermal cell fresh mass [APD:glossary/40044].
The ratio of the mass of phosphorus in epidermal cells to epidermal cell fresh mass.
comments
value type continuous variable
unit umol{P}/g
micromole per gram
minAllowedValue 0
maxAllowedValue 10
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
epidermal cell
cell
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_epidermis_P_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001661

Leaf hypodermis phosphorus (P) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001662
alternative label leaf_hypodermis_P_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in the hypodermis [PO:0005051] to hypodermis fresh mass [APD:glossary/40044].
The ratio of the mass of phosphorus in the hypodermis to hypodermis fresh mass.
comments
value type continuous variable
unit umol{P}/g
micromole per gram
minAllowedValue 0
maxAllowedValue 10
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
hypodermis
cell
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_hypodermis_P_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001662

Leaf internal parenchyma cell phosphorus (P) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001663
alternative label leaf_internal_parenchyma_P_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in parenchyma cells [PO:0000074] to parenchyma cell fresh mass [APD:glossary/40044].
The ratio of the mass of phosphorus in parenchyma cells to parenchyma cell fresh mass.
comments
value type continuous variable
unit umol{P}/g
micromole per gram
minAllowedValue 0
maxAllowedValue 10
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
parenchyma cell
cell
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_internal_parenchyma_P_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001663

Leaf palisade mesophyll cell phosphorus (P) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001664
alternative label leaf_palisade_mesophyll_P_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in palisade mesophyll cells [PO:0006206] to palisade mesophyll cell fresh mass [APD:glossary/40044].
The ratio of the mass of phosphorus in palisade mesophyll cells to palisade mesophyll cell fresh mass.
comments
value type continuous variable
unit umol{P}/g
micromole per gram
minAllowedValue 0
maxAllowedValue 30
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
palisade mesophyll cell
cell
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_palisade_mesophyll_P_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001664

Leaf sclerenchyma cell phosphorus (P) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001665
alternative label leaf_sclerenchyma_P_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in sclerenchyma cells [PO:0000077] to sclerenchyma cell fresh mass [APD:glossary/40044].
The ratio of the mass of phosphorus in sclerenchyma cells to sclerenchyma cell fresh mass.
comments
value type continuous variable
unit umol{P}/g
micromole per gram
minAllowedValue 0
maxAllowedValue 10
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
sclerenchyma cell
cell
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_sclerenchyma_P_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001665

Leaf spongy mesophyll cell phosphorus (P) content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0001666
alternative label leaf_spongy_mesophyll_P_per_fresh_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in spongy mesophyll cells [PO:0006205] to spongy mesophyll cell fresh mass [APD:glossary/40044].
The ratio of the mass of phosphorus in spongy mesophyll cells to spongy mesophyll cell fresh mass.
comments
value type continuous variable
unit umol{P}/g
micromole per gram
minAllowedValue 0
maxAllowedValue 30
has broader cell or tissue mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
spongy mesophyll cell
cell
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note cell_spongy_mesophyll_P_per_fresh_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0001666

Lipid fraction of leaf P content per dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001667
alternative label leaf_lipid_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to lipids [CHEBI:18059] to leaf total dry mass [TO:0001014].
The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
lipid
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001667

Nucleic acid fraction of leaf P content per dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001668
alternative label leaf_nucleic_acid_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of the fraction of total phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to nucleic acids [CHEBI:33696] to leaf total dry mass [TO:0001014].
The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.001
maxAllowedValue 10
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
nucleic acid
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001668

Residual P fraction of leaf P content per dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001669
alternative label leaf_residual_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to residual P to leaf total dry mass [TO:0001014].
The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0
maxAllowedValue 1
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001669

Inorganic P fraction of leaf P content per dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001670
alternative label leaf_inorganic_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] that is inorganic [NCIT:C66829] P to leaf total dry mass [TO:0001014].
The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.01
maxAllowedValue 20
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
inorganic
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001670

Metabolite fraction of leaf P content per dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0001671
alternative label leaf_metabolite_P_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to metabolites [NCIT:C61154] to leaf total dry mass [TO:0001014].
The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1e-04
maxAllowedValue 10
has broader live leaf mineral and ion content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phosphorus atom
metabolite
macronutrient
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0001671

Leaf total non-structural carbohydrate content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002021
alternative label leaf_total_non-structural_carbohydrates_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of non-structural carbohydrates [CHEBI:16646] (sugars [APD:glossary/40106]) in a leaf [PO:0025034] to leaf area [PATO:0001323], where non-structural carbohydrates are carbohydrates that do not comprise the cell wall [GO:0009505].
The ratio of the mass of non-structural carbohydrates (sugars) in a leaf to leaf area, where non-structural carbohydrates are carbohydrates that do not comprise the cell wall.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 50
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword carbohydrate
starch
sugar
non-structural carbohydrates
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002021

Leaf total non-structural carbohydrate content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002022
alternative label leaf_total_non-structural_carbohydrates_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of non-structural carbohydrates [CHEBI:16646] (sugars [APD:glossary/40106]) in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where non-structural carbohydrates are carbohydrates that do not comprise the cell wall [GO:0009505].
The ratio of the mass of non-structural carbohydrates (sugars) in a leaf to leaf dry mass, where non-structural carbohydrates are carbohydrates that do not comprise the cell wall.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 10
maxAllowedValue 200
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword carbohydrate
starch
sugar
non-structural carbohydrates
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002022

Leaf cellulose content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002024
alternative label leaf_cellulose_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of cellulose [CHEBI:62967] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of cellulose in a leaf to leaf dry mass.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.01
maxAllowedValue 1
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword amorphous cellulose
carbohydrate
scope note
has exact match
has close match
has related match cellulose content
examples exact match: Leaf cellulose content per leaf dry mass [TOP819] (https://top-thesaurus.org/index)
exact match: Leaf cellulose content per leaf dry mass [TRY:92] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002024

Leaf starch content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002025
alternative label leaf_starch_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of total starch [CHEBI:28017] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of total starch in a leaf to leaf area.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 50
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword starch
carbohydrate
scope note
has exact match
has close match
has related match starch content
examples exact match: Leaf starch content per leaf dry mass [TRY:505] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note starch_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002025

Leaf soluble starch content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002027
alternative label leaf_soluble_starch_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of water soluble starch in a leaf to leaf area.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 100
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword starch
carbohydrate
scope note
has exact match
has close match
has related match
examples exact match: Leaf starch content per leaf dry mass [TRY:505] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note soluble_starch_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002027

Leaf soluble starch content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002028
alternative label leaf_soluble_starch_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of water soluble starch in a leaf to leaf dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 200
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword starch
carbohydrate
scope note
has exact match
has close match
has related match
examples exact match: Leaf starch content per leaf dry mass [TRY:505] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002028

Leaf soluble sugar content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002031
alternative label leaf_soluble_sugars_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a leaf [PO:0025034] to leaf area [PATO:0001323], where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].
The ratio of the mass of sugar in a leaf to leaf area, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 100
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword carbohydrate
monosaccharide
disaccharide
sugar
scope note
has exact match
has close match
has related match total soluble sugar content
examples exact match: Leaf sugar content per leaf area [TOP920] (https://top-thesaurus.org/index)
exact match: Leaf sugar content per leaf dry mass [TRY:506] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note soluble_sugars_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002031

Leaf soluble sugar content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002032
alternative label leaf_soluble_sugars_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].
The ratio of the mass of sugar in a leaf to leaf dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 200
has broader carbohydrate content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword carbohydrate
monosaccharide
disaccharide
sugar
scope note
has exact match
has close match
has related match total soluble sugar content
examples exact match: Leaf sugar content per leaf dry mass [TOP919] (https://top-thesaurus.org/index)
exact match: Leaf sugar content per leaf dry mass [TRY:506] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002032

Leaf soluble protein content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002035
alternative label leaf_soluble_protein_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] protein [CHEBI:36080] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of water soluble protein in a leaf to leaf area.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 20
has broader protein content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword protein
scope note
has exact match Leaf soluble protein content
has close match
has related match soluble protein content
examples close match: Leaf protein content per leaf dry mass [TRY:503] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note soluble_protein_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002035

Leaf insoluble protein content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002037
alternative label leaf_insoluble_protein_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] insoluble [PATO:0001538] protein [CHEBI:36080] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of water insoluble protein in a leaf to leaf area.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 20
has broader protein content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword protein
scope note
has exact match
has close match
has related match
examples close match: Leaf protein content per leaf dry mass [TRY:503] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note insoluble_protein_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002037

Leaf lignin content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002050
alternative label leaf_lignin_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of lignin [CHEBI:6457] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of lignin in a leaf to leaf dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 10
maxAllowedValue 1000
has broader phenolic compound content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword lignin
scope note
has exact match
has close match
has related match lignin content
examples exact match: Leaf lignin content per leaf dry mass [TOP844] (https://top-thesaurus.org/index)
exact match: Leaf lignin content per leaf dry mass [TRY:87] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed 13/02/2023
reviewed by Félix de Tombeur
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002050

Total leaf phenolic content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002052
alternative label leaf_phenol_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of phenol [CHEBI:33853] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where phenol content is measured as gallic acid [CHEBI:30778] equivalents.
The ratio of the mass of phenol in a leaf to leaf dry mass, where phenol content is measured as gallic acid equivalents.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 1000
has broader phenolic compound content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phenol
gallic acid
scope note
has exact match
has close match
has related match
examples exact match: Leaf phenols content per leaf dry mass [TRY:147] (https://www.try-db.org/de/de.php)
references Salminen & Karonen 2011
date created 14/07/2021
date modified 05/02/2024
date reviewed 13/02/2023
reviewed by Félix de Tombeur
change note leaf_phenol_content_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002052

Leaf tannin content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002054
alternative label leaf_tannin_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of tannins [CHEBI:26848] (a class of polyphenols [CHEBI:26195]) in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of tannins (a class of polyphenols) in a leaf to leaf dry mass.
comments Tannins are a specific class of polyphenolic compounds found in plants.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 1000
has broader phenolic compound content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword phenol
tannin
polyphenol
scope note
has exact match
has close match
has related match
examples exact match: Leaf tannin content per leaf dry mass [TRY:148] (https://www.try-db.org/de/de.php)
references Salminen & Karonen 2011
date created 14/07/2021
date modified 06/06/2025
date reviewed 13/02/2023
reviewed by Félix de Tombeur
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002054

Leaf carotenoid content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002055
alternative label leaf_carotenoid_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carotenoids [CHEBI:23044] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of carotenoids in a leaf to leaf area.
comments
value type continuous variable
unit umol{carotenoid}/m2
micromole per square metre
minAllowedValue 10
maxAllowedValue 1000
has broader pigment content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword carotenoid
scope note
has exact match
has close match
has related match carotenoid content
examples exact match: Leaf carotenoid content per leaf area [TRY:491] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note carotenoid_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002055

Leaf carotenoid content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002056
alternative label leaf_carotenoid_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of carotenoids [CHEBI:23044] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of carotenoids in a leaf to leaf dry mass.
comments
value type continuous variable
unit mmol{carotenoid}/kg
millimole per kilogram
minAllowedValue 0.1
maxAllowedValue 10
has broader pigment content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword carotenoid
scope note
has exact match
has close match
has related match carotenoid content
examples exact match: Leaf carotenoid content per leaf dry mass [TRY:809] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note carotenoid_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002056

Leaf total chlorophyll content (chlorophyll A + B) per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002081
alternative label leaf_chlorophyll_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll [CHEBI:28966] in a leaf [PO:0025034] to leaf area [PATO:0001323], where chlorophyll content is the sum of chlorophyll a [CHEBI:18230] and chlorophyll b [CHEBI:27888] content.
The ratio of the mass of chlorophyll in a leaf to leaf area, where chlorophyll content is the sum of chlorophyll a and chlorophyll b content.
comments
value type continuous variable
unit umol{chlorophyll}/m2
micromole per square metre
minAllowedValue 10
maxAllowedValue 2000
has broader pigment content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword chlorophyll
chlorophyll a
chlorophyll b
chloroplast
scope note
has exact match Leaf chlorophyll content
has close match leaf chlorophyll content
has related match chlorophyll content
examples exact match: Leaf chlorophyll content per leaf area [TRY:413] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002081

Leaf total chlorophyll content (chlorophyll A + B) per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002082
alternative label leaf_chlorophyll_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll [CHEBI:28966] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where chlorophyll content is the sum of chlorophyll a [CHEBI:18230] and chlorophyll b [CHEBI:27888] content.
The ratio of the mass of chlorophyll in a leaf to leaf dry mass, where chlorophyll content is the sum of chlorophyll a and chlorophyll b content.
comments
value type continuous variable
unit mmol{chlorophyll}/kg
millimole per kilogram
minAllowedValue 0.1
maxAllowedValue 15
has broader pigment content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword chlorophyll
chlorophyll a
chlorophyll b
chloroplast
scope note
has exact match Leaf chlorophyll content
has close match leaf chlorophyll content
has related match chlorophyll content
examples exact match: Leaf chlorophyll content per leaf dry mass [TRY:164] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002082

Leaf chlorophyll A content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002083
alternative label leaf_chlorophyll_A_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of chlorophyll a in a leaf to leaf area.
comments
value type continuous variable
unit umol{chlorophyllA}/m2
micromole per square metre
minAllowedValue 50
maxAllowedValue 2000
has broader pigment content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword chlorophyll a
chlorophyll
chloroplast
scope note
has exact match
has close match Chlorophyll a
chlorophyll a content
has related match
examples exact match: Leaf chlorophyll content per leaf area (Chl a) [TRY:3476] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_A_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002083

Leaf chlorophyll A content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002084
alternative label leaf_chlorophyll_A_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of chlorophyll a in a leaf to leaf dry mass.
comments
value type continuous variable
unit mmol{chlorophyllA}/kg
millimole per kilogram
minAllowedValue 0.1
maxAllowedValue 10
has broader pigment content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword chlorophyll a
chlorophyll
chloroplast
scope note
has exact match Chlorophyll a
has close match chlorophyll a content
has related match
examples exact match: Leaf chlorophyll content per leaf dry mass (Chl a) [TRY:3474] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_A_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002084

Leaf chlorophyll B content per unit leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0002085
alternative label leaf_chlorophyll_B_per_area
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll b [CHEBI:27888] in a leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of chlorophyll b in a leaf to leaf area.
comments
value type continuous variable
unit umol{chlorophyllB}/m2
micromole per square metre
minAllowedValue 10
maxAllowedValue 1000
has broader pigment content trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword chlorophyll b
chlorophyll
chloroplast
scope note
has exact match
has close match Chlorophyll b
chlorophyll b content
has related match
examples exact match: Leaf chlorophyll content per leaf area (Chl b) [TRY:3477] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_B_per_area
is in scheme https://w3id.org/APD/traits
identifier trait_0002085

Leaf chlorophyll B content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002086
alternative label leaf_chlorophyll_B_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll b [CHEBI:27888] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of chlorophyll b in a leaf to leaf dry mass.
comments
value type continuous variable
unit mmol{chlorophyllB}/kg
millimole per kilogram
minAllowedValue 0.01
maxAllowedValue 10
has broader pigment content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword chlorophyll b
chlorophyll
chloroplast
scope note
has exact match Chlorophyll b
has close match chlorophyll b content
has related match
examples exact match: Leaf chlorophyll content per leaf dry mass (Chl b) [TRY:3475] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_B_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002086

Ratio of leaf chlorophyll A content to leaf chlorophyll B content

Property Value
URI https://w3id.org/APD/traits/trait_0002087
alternative label leaf_chlorophyll_A_B_ratio
description The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to the mass of chlorophyll b [CHEBI:27888] in the leaf.
The ratio of the mass of chlorophyll a in a leaf to the mass of chlorophyll b in the leaf.
comments
value type continuous variable
unit umol{chlorophyllA}/umol{chlorophyllB}
micromole per micromole
minAllowedValue 0.1
maxAllowedValue 6
has broader pigment content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
dimensionless ratio
keyword chlorophyll a
chlorophyll b
chlorophyll
chloroplast
scope note
has exact match Chlorophyll a to b ratio
chlorophyll ratio
has close match
has related match
examples exact match: Leaf chlorophyll a/b ratio [TRY:416] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note chlorophyll_A_B_ratio
is in scheme https://w3id.org/APD/traits
identifier trait_0002087

Leaf rubisco content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002090
alternative label leaf_rubisco_per_leaf_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of rubisco [MESH:D012273] in a leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of rubisco in a leaf to leaf dry mass.
comments This trait is most commonly measured on leaf tissue. If the Rubisco concentration of a different tissue is measured, this should be explicitly noted in the methods or as a method context.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 5
maxAllowedValue 50
has broader protein content trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword Ribulose-Bisphosphate Carboxylase
chloroplast
protein
scope note
has exact match Leaf rubisco content
has close match rubisco content
has related match
examples close match: Leaf Rubisco content per leaf area [TRY:504] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note cell_rubisco_concentration
is in scheme https://w3id.org/APD/traits
identifier trait_0002090

Stem soluble starch content per unit stem dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002127
alternative label stem_soluble_starch_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a stem [PO:0009047] to stem dry mass.
The ratio of the mass of water soluble starch in a stem to stem dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader carbohydrate content trait
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword starch
carbohydrate
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002127

Stem soluble sugar content per unit stem dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002131
alternative label stem_soluble_sugars_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a stem [PO:0009047] to stem dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].
The ratio of the mass of sugar in a stem to stem dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 200
has broader carbohydrate content trait
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword carbohydrate
monosaccharide
disaccharide
sugar
scope note
has exact match
has close match
has related match sugar content
examples exact match: Stem sugar content per stem dry mass [TOP914] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002131

Bark cellulose content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002224
alternative label bark_cellulose_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of cellulose [CHEBI:62967] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of cellulose in bark to bark dry mass.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.1
maxAllowedValue 1
has broader carbohydrate content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword amorphous cellulose
scope note
has exact match
has close match
has related match
examples exact match: Bark cellulose content per bark dry mass [TOP813] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002224

Bark lignin content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002250
alternative label bark_lignin_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of lignin [CHEBI:6457] in bark [PO:0004518] to bark dry mass.
The ratio of the mass of lignin in bark to bark dry mass.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.01
maxAllowedValue 1
has broader phenolic compound content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword lignin
scope note
has exact match
has close match
has related match
examples exact match: Bark lignin content per bark dry mass [TOP838] (https://top-thesaurus.org/index)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002250

Bark tannin content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002255
alternative label bark_tannin_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of tannins [CHEBI:26848] (polyphenols [CHEBI:26195]) in bark [PO:0004518] to bark dry mass.
The ratio of the mass of tannins (polyphenols) in bark to bark dry mass.
comments Tannins are a specific class of polyphenolic compounds found in plants.
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 1e-05
maxAllowedValue 1
has broader phenolic compound content trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword tannin
polyphenol
scope note
has exact match
has close match
has related match
examples
references Salminen & Karonen 2011
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002255

Root soluble starch content per unit root dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002327
alternative label root_soluble_starch_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in roots [PO:0009005] to root dry mass.
The ratio of the mass of water soluble starch in roots to root dry mass.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 0.1
maxAllowedValue 100
has broader carbohydrate content trait
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword starch
carbohydrate
scope note
has exact match
has close match
has related match
examples exact match: Root starch content per dry mass [TRY:3147] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002327

Root soluble sugar content per unit root dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002331
alternative label root_soluble_sugars_per_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in roots [PO:0009005] to root dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233].
The ratio of the mass of sugar in roots to root dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 1
maxAllowedValue 200
has broader carbohydrate content trait
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword carbohydrate
monosaccharide
disaccharide
sugar
scope note
has exact match
has close match
has related match
examples exact match: Root sugar content per root dry mass trait [TOP921] (https://top-thesaurus.org/index)
exact match: Root sugar content per root dry mass [TRY:662] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0002331

Seed protein content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002534
alternative label seed_protein_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of protein [CHEBI:36080] in a seed [PO:0009010] to seed dry mass [TO:0000181].
The ratio of the mass of protein in a seed to seed dry mass.
comments It is best practice for the denominator to be seed dry mass. If data are reported on a per seed fresh mass basis, this should be explicitly noted in the methods or as a method context.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 1e-05
maxAllowedValue 1
has broader protein content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword protein
seed weight, seed mass
scope note
has exact match seed protein content
has close match
has related match
examples exact match: Seed protein content per seed mass [TRY:97] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_protein_content
is in scheme https://w3id.org/APD/traits
identifier trait_0002534

Seed oil content per unit seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002544
alternative label seed_oil_per_seed_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of lipids [CHEBI:18059] in a seed [PO:0009010] to seed dry mass [TO:0000181].
The ratio of the mass of lipids in a seed to seed dry mass.
comments It is best practice for the denominator to be seed dry mass. If data are reported on a per seed fresh mass basis, this should be explicitly noted in the methods or as a method context.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 1e-04
maxAllowedValue 1
has broader lipid content trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
mass fraction
ratio; proportion
keyword lipid
seed weight, seed mass
scope note
has exact match
has close match
has related match
examples exact match: Seed lipid content per seed dry mass [TOP880] (https://top-thesaurus.org/index)
exact match: Seed oil content per seed mass [TRY:96] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seed_oil_content
is in scheme https://w3id.org/APD/traits
identifier trait_0002544

Leaf ash content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002822
alternative label leaf_ash_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of leaf [PO:0025034] ash [ENVO:02000090] remaining after a combustion process [ENVO:01000839] to the leaf dry mass before the combustion process.
The ratio of the mass of leaf ash remaining after a combustion process to the leaf dry mass before the combustion process.
comments Includes measurements on leaf analogs, including cladodes and phylloclades.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 1e-04
maxAllowedValue 1
has broader live leaf mineral and ion content trait
fire response trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword ash
combustion process
fire
scope note
has exact match
has close match
has related match
examples exact match: Leaf ash content per leaf dry mass [TRY:333] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note leaf_ash_content_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002822

Bark ash content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0002824
alternative label bark_ash_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of bark [PO:0004518] ash [ENVO:02000090] remaining after a combustion process [ENVO:01000839] to the bark dry mass before the combustion process.
The ratio of the mass of bark ash remaining after a combustion process to the bark dry mass before the combustion process.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 1e-04
maxAllowedValue 1
has broader stem mineral and ion content trait
fire response trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword ash
combustion process
fire
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note bark_ash_content_per_dry_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0002824

Bark stable carbon isotope composition (delta13C)

Property Value
URI https://w3id.org/APD/traits/trait_0003011
alternative label bark_delta13C
description Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of bark [PO:0004518].
Carbon stable isotope ratio, d13C, of bark.
comments delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -50
maxAllowedValue 0
has broader stable isotope ratio determination
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon stable isotope ratio
d13C
carbon-13 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Bark carbon (C) isotope signature (delta 13C) [TRY:829] (https://www.try-db.org/de/de.php)
related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003011

Leaf stable carbon isotope composition (delta13C)

Property Value
URI https://w3id.org/APD/traits/trait_0003012
alternative label leaf_delta13C
description Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of leaves [PO:0025034].
Carbon stable isotope ratio, d13C, of leaves.
comments delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -50
maxAllowedValue 0
has broader stable isotope ratio determination
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon stable isotope ratio
d13C
carbon-13 atom
isotope
scope note
has exact match Delta C13
has close match
has related match
examples exact match: Leaf carbon isotope signature (delta 13C) [TRY:89] (https://www.try-db.org/de/de.php)
related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003012

Stem stable carbon isotope composition (delta13C)

Property Value
URI https://w3id.org/APD/traits/trait_0003013
alternative label stem_delta13C
description Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of stems [PO:0009047].
Carbon stable isotope ratio, d13C, of stems.
comments delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -50
maxAllowedValue 0
has broader stable isotope ratio determination
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon stable isotope ratio
d13C
carbon-13 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Stem carbon isotope ratio (delta 13C) [TRY:4049] (https://www.try-db.org/de/de.php)
related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 29/03/2023
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003013

Root stable carbon isotope composition (delta13C)

Property Value
URI https://w3id.org/APD/traits/trait_0003014
alternative label root_delta13C
description Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of roots [PO:0009005].
Carbon stable isotope ratio, d13C, of roots.
comments delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -50
maxAllowedValue 0
has broader stable isotope ratio determination
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon stable isotope ratio
d13C
carbon-13 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Root carbon isotope signature (delta 13C) [TRY:90] (https://www.try-db.org/de/de.php)
related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003014

Wood stable carbon isotope composition (delta13C)

Property Value
URI https://w3id.org/APD/traits/trait_0003015
alternative label wood_delta13C
description Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of wood [NCIT:C93062].
Carbon stable isotope ratio, d13C, of wood.
comments delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -50
maxAllowedValue 0
has broader stable isotope ratio determination
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword carbon stable isotope ratio
d13C
wood
carbon-13 atom
isotope
scope note
has exact match Wood Carbon content
has close match
has related match
examples exact match: Wood carbon (C) isotope signature (delta 13C) [TRY:1213] (https://www.try-db.org/de/de.php)
related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003015

Bark stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003031
alternative label bark_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of bark [PO:0004518].
Nitrogen stable isotope ratio, d15N, of bark.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
nitrogen-15 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Bark nitrogen (N) isotope signature (delta 15N) [TRY:832] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003031

Leaf stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003032
alternative label leaf_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of leaves [PO:0025034].
Nitrogen stable isotope ratio, d15N, of leaves.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
nitrogen-15 atom
isotope
scope note
has exact match Delta 15N
has close match
has related match
examples exact match: Leaf nitrogen (N) isotope signature (delta 15N) [TRY:78] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003032

Stem stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003033
alternative label stem_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of stems [PO:0009047].
Nitrogen stable isotope ratio, d15N, of stems.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
nitrogen-15 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Root nitrogen (N) isotope signature (delta 15N) [TRY:81] (https://www.try-db.org/de/de.php)
references no linked references
date created 29/03/2023
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003033

Root stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003034
alternative label root_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of roots [PO:0009005].
Nitrogen stable isotope ratio, d15N, of roots.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
nitrogen-15 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Root nitrogen (N) isotope signature (delta 15N) [TRY:81] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003034

Wood stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003035
alternative label wood_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of wood [NCIT:C93062].
Nitrogen stable isotope ratio, d15N, of wood.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure plant structure stem
wood, secondary xylem
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
wood
nitrogen-15 atom
isotope
scope note
has exact match
has close match
has related match
examples exact match: Wood nitrogen (N) isotope signature (delta 15N) [TRY:1230] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003035

Leaf xylem stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003052
alternative label leaf_xylem_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of xylem sap [PO:0025539] from leaf [PO:0025034] xylem vessels [PO:0025417].
Nitrogen stable isotope ratio, d15N, of xylem sap from leaf xylem vessels.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
xylem vessel
nitrogen-15 atom
isotope
xylem sap
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003052

Root xylem stable nitrogen isotope composition (delta15N)

Property Value
URI https://w3id.org/APD/traits/trait_0003053
alternative label root_xylem_delta15N
description Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of xylem sap [PO:0025539] from root [PO:0009005] xylem vessels [PO:0025417].
Nitrogen stable isotope ratio, d15N, of xylem sap from root xylem vessels.
comments delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -25
maxAllowedValue 75
has broader stable isotope ratio determination
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword stable nitrogen isotope ratio
d15N
xylem vessel
nitrogen-15 atom
isotope
xylem sap
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003053

Leaf stable oxygen isotope composition (delta18O)

Property Value
URI https://w3id.org/APD/traits/trait_0003072
alternative label leaf_delta18O
description Oxygen stable isotope ratio [ECSO:00001740], d18O [ECSO:00001677], of leaves [PO:0025034].
Oxygen stable isotope ratio, d18O, of leaves.
comments delta18O is calculated as the ratio of, 18O/16O in a sample to 18O/16O in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta18O indicates the sample has a lower ratio of 18O/16O relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -10
maxAllowedValue 50
has broader stable isotope ratio determination
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword stable oxygen isotope ratio
d18O
oxygen-18 atom
scope note
has exact match
has close match
has related match
examples exact match: Leaf oxygen (O) isotope signature (delta 18O) [TRY:3358] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003072

Stem water stable oxygen isotope composition (delta18O)

Property Value
URI https://w3id.org/APD/traits/trait_0003092
alternative label stem_water_delta18O
description Oxygen stable isotope ratio [ECSO:00001740], d18O [ECSO:00001677], of water [CHEBI:15377] in stems [PO:0009047].
Oxygen stable isotope ratio, d18O, of water in stems.
comments delta18O is calculated as the ratio of, 18O/16O in a sample to 18O/16O in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta18O indicates the sample has a lower ratio of 18O/16O relative to the standard.
value type continuous variable
unit {delta}[ppth]
parts per thousand
minAllowedValue -10
maxAllowedValue 50
has broader stable isotope ratio determination
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword stable oxygen isotope ratio
d18O
water
oxygen-18 atom
d18O in water
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0003092

Plant canopy width

Property Value
URI https://w3id.org/APD/traits/trait_0010021
alternative label plant_width
description A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] width [PATO:0000600] dimension of a plant canopy [ENVO:01001242] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341]. This is a plant’s larger [NCIT:C49508] width dimension and is orthogonal to its height and breadth.
Maximum width dimension of a plant’s vegetative canopy for a taxon in a given location; this is the larger width dimension and is orthogonal to its height and breadth.
comments This trait is not meant to document the width of all individuals sampled, but instead the maximum width for a typical mature individual within a population or across a taxon.
value type continuous variable
unit m
metre
minAllowedValue 0.001
maxAllowedValue 20
has broader whole plant morphology trait
plant structure whole plant
measured characteristic width
size
keyword canopy
vegetation
scope note
has exact match plant width
has close match Crown size
has related match
examples exact match: Crown (canopy) width [TRY:325] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010021

Plant canopy breadth

Property Value
URI https://w3id.org/APD/traits/trait_0010022
alternative label plant_breadth
description A whole plant morphology trait [TO:0000398] which is the smaller [NCIT:C25376] width [PATO:0000600] dimension of a plant canopy [ENVO:01001242].
Shortest width dimension of a plant’s vegetative canopy; orthogonal to its height and width.
comments This trait is not meant to document the breadth of all individuals sampled, but instead the maximum breadth for a typical mature individual within a population or across a taxon.
value type continuous variable
unit m
metre
minAllowedValue 0.001
maxAllowedValue 20
has broader whole plant morphology trait
plant structure whole plant
measured characteristic width
size
keyword canopy
vegetation
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010022

Plant vegetative height

Property Value
URI https://w3id.org/APD/traits/trait_0010023
alternative label plant_height
description A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the vegetative shoot system [PO:0025607] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341].
Maximum vertical height of the vegetative shoot system for a taxon in a given location.
comments This trait is not meant to document the height of all individuals sampled, but instead the maximum height for a typical mature individual within a population or across a taxon. The trait should only be scored for self-supporting plants, not climbers whose height is dependent on the surrounding vegetation. The documented height should be the maximum height achieved by vegetative shoots, not the height reached by reproductive shoots, as it represents the plant’s access to light.
value type continuous variable
unit m
metre
minAllowedValue 0.001
maxAllowedValue 130
has broader whole plant morphology trait
plant structure plant structure whole plant
stem
measured characteristic height
size
keyword canopy
vegetation
vegetative shoot system
scope note self-supporting plants
has exact match Tree height
Tree height
plant height
has close match plant height
has related match
examples exact match: Plant height vegetative [TOP69] (https://top-thesaurus.org/index)
exact match: Plant height vegetative [TRY:3106] (https://www.try-db.org/de/de.php)
exact match: Plant_height_max [GIFT:1.6.2] (https://gift.uni-goettingen.de)
exact match: maximum whole plant height [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: whole plant height [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
related match: minimum whole plant height [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Average height [BROT:10] (http://doi.org/10.1038/sdata.2018.135)
exact match: MaxStemHeight_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references Pérez-Harguindeguy 2013
Falster 2003a
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010023

Stem diameter at breast height

Property Value
URI https://w3id.org/APD/traits/trait_0010024
alternative label plant_diameter_breast_height
description A stem morphology trait [TO:0000361] which is the maximum [NCIT:C25564] diameter at breast height [ECSO:00000549] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341].
Maximum diameter at breast height (DBH) for a taxon in a given location. It is commonly referred to as ‘DBH’.
comments This trait is always meant to be measured at ‘breast height’, 1.4 m above the ground surface. However, for short plants this will not always be the case, so it is advised to read the methods to confirm at what height plant diameters have been recorded. This trait is not meant to document the DBH of all individuals sampled, but instead the maximum DBH for a typical mature individual within a population or across a taxon.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 5000
has broader stem morphology trait
plant structure plant structure stem
whole plant
measured characteristic diameter
size
keyword tree diameter at breast height
vegetation
scope note
has exact match Stem diameter
tree breast height diameter
has close match Stem diameter
stem diameter
has related match
examples exact match: Stem diameter [TRY:21] (https://www.try-db.org/de/de.php)
exact match: DBH_max [GIFT:1.8.2] (https://gift.uni-goettingen.de)
exact match: diameter at breast height (1.3 m) [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: MaxStemDia_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note basal_diameter
is in scheme https://w3id.org/APD/traits
identifier trait_0010024

Stem count

Property Value
URI https://w3id.org/APD/traits/trait_0010025
alternative label stem_count
description A shoot system morphology trait [TO:0000077] which is the count [NCIT:C25463] of stems [PO:0009047] on multi-stemmed plants [PO:0000003].
Count of stems on multi-stemmed plants.
comments
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 5
has broader stem morphology trait
plant structure stem
measured characteristic has number of
count
keyword branchiness
scope note
has exact match stem number
has close match
has related match
examples close match: Plant biomass and allometry: Shoots per plant [TRY:1034] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stem_count_categorical
is in scheme https://w3id.org/APD/traits
identifier trait_0010025

Height of a climbing plant

Property Value
URI https://w3id.org/APD/traits/trait_0010026
alternative label plant_height_climbing_plant
description A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the vegetative shoot system [PO:0025607] of a climbing plant that is not self-supporting and therefore has a height that depends on the surrounding vegetation.
Maximum vertical height of the vegetative shoot system of a non-self-supporting taxon (i.e. a vine, liana, or other climbing plant) in a given location.
comments While this trait is intended to indicate the height of climbing plants, the height of an epiphyte relative to the soil surface could also be documented here. All heights reported for this trait will be dependent on the height of the surrounding vegetation.
value type continuous variable
unit m
metre
minAllowedValue 0.1
maxAllowedValue 100
has broader whole plant morphology trait
plant structure whole plant
measured characteristic height
size
keyword canopy
vegetation
vegetative shoot system
scope note applicable to plants that are not self-supporting
has exact match
has close match
has related match plant height
examples related match: Plant height [TRY:18] (https://www.try-db.org/de/de.php)
related match: Average height [BROT:10] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010026

Plant reproductive height

Property Value
URI https://w3id.org/APD/traits/trait_0010027
alternative label plant_height_reproductive
description A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the reproductive shoot system [PO_0025082].
Maximum vertical height of the reproductive shoot system for a taxon in a given location.
comments This trait can be scored for any species, but is intended as an alternative to plant_height (maximum vegetative height) for species that do not have vegetative stems, but instead only leaves and reproductive stems that extend past the soil surface.
value type continuous variable
unit m
metre
minAllowedValue 0.1
maxAllowedValue 100
has broader whole plant morphology trait
plant structure whole plant
measured characteristic height
size
keyword vegetation
reproductive shoot system
scope note
has exact match
has close match
has related match
examples related match: Plant height [TRY:18] (https://www.try-db.org/de/de.php)
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010027

Stem length

Property Value
URI https://w3id.org/APD/traits/trait_0010028
alternative label stem_length
description A shoot system morphology trait [TO:0000077] which is the maximum [NCIT:C25564] length [PATO_0000122] of a plant’s stems [PO_0009047].
Maximum length or extent of a plant’s stems.
comments This trait can be scored for any species, but is intended as an alternative to plant_height (maximum vegetative height) for species that are prostrate, crawling, creeping or similar.
value type continuous variable
unit m
metre
minAllowedValue 0.01
maxAllowedValue 10
has broader stem morphology trait
plant structure plant structure stem
whole plant
measured characteristic length
size
keyword vegetation
vegetative shoot system
scope note
has exact match
has close match
has related match
examples exact match: Stem length [TRY:326] (https://www.try-db.org/de/de.php)
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010028

Plant spinescence

Property Value
URI https://w3id.org/APD/traits/trait_0010070
alternative label plant_spinescence
description A plant structure morphology trait [TO:0000839] which summarises the extent of physical structures a plant uses for defence [NCIT:C61556], including the size [PATO:0000117], strength [PATO:0001230], and density [PATO:0001019] of spine leaves [PO:0025173], spine stipules [PO:0025174], thorns [PO:0025172], and/or prickles [PO:0025169] and the pain [HP:0012531] inflicted by these structures.
Degree to which a plant is defended by spines, thorns and/or prickles, as defined by P<U+00C3><U+00A9>rez-Harguindeguy et al. 2013.
comments The values for this trait are exactly those defined by P<U+00C3><U+00A9>rez-Harguindeguy et al. 2013 (doi:10.1071/BT12225). See the trait plant_physical_defence for individual plant structures a plant uses for physical defence.
value type categorical variable
has narrower plant_spinescence_no_spines No spines. (level 0)
plant_spinescence_low_density_soft_spines Low or very local density of soft spines <5mm long; plant may sting or prickle when hit carelessly but not impart strong pain. (level 1)
plant_spinescence_high_density_soft_spines High density of soft spines, intermediate density of spines of intermediate hardness, or low density of hard, sharp spines >5mm long; plant causes actual pain when hit carelessly. (level 2)
plant_spinescence_high_density_hard_spines_to_5mm Intermediate or high density of hard, sharp spines >5mm long; plant causes strong pain when hit carelessly. (level 3)
plant_spinescence_high_density_hard_spines_to_20mm Intermediate or high density of hard, sharp spines >20mm long; plant may cause significant wounds when hit carelessly. (level 4)
plant_spinescence_high_density_hard_spines_to_100mm Intermediate or high density of hard, sharp spines >100mm long; plant is dangerous to careless large mammals, including humans. (level 5)
has broader leaf shape trait
whole plant phenotype trait
plant structure morphology trait
plant structure plant structure plant structure leaf
stem
whole plant
measured characteristic structure
surface feature shape
keyword spine leaf
spine stipule
thorn
prickle
defense
pain
spiny
scope note none
has exact match
has close match
has related match
examples close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
close match: Spinescence [BROT:21] (http://doi.org/10.1038/sdata.2018.135)
close match: Armature [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note spinescence
is in scheme https://w3id.org/APD/traits
identifier trait_0010070

Embryo colour

Property Value
URI https://w3id.org/APD/traits/trait_0010110
alternative label embryo_colour
description A plant embryo cotyledon morphology trait [TO:0000775] which is the colour [PATO:0000014] of the plant embryo cotyledon [PO:0025470].
Binary variable distinguishing between embryos that are green versus colourless.
comments
value type categorical variable
has narrower embryo_colour_colourless Colourless embryo.
embryo_colour_green Green embryo.
has broader plant embryo morphology trait
plant structure whole plant
measured characteristic colour / color
keyword plant embryo cotyledon
plant embryo
scope note
has exact match plant embryo cotyledon color
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010110

Cotyledon function

Property Value
URI https://w3id.org/APD/traits/trait_0010111
alternative label cotyledon_function
description A plant embryo morphology trait [TO:0000064] which indicates if the cotyledons [PO:0020030] have a photosynthesis [GO:0015979] or energy [PATO:0001021] storage [OBI:0302893] function [NCIT:C25518].
Binary variable distinguishing between cotyledons that function as leaves and can photosynthesise versus ones that function as energy reserve organs.
comments
value type categorical variable
has narrower cotyledon_function_foliaceous Cotyledons are photosynthetic and function as leaves.
cotyledon_function_reserve Cotyledons function as energy reserve organs.
has broader plant embryo morphology trait
leaf morphology trait
plant structure plant structure whole plant
leaf
measured characteristic organismal quality
function
keyword cotyledon
photosynthesis
energy
storage
plant embryo
seedling
plant embryo cotyledon
storage organ
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010111

Cotyledon position at germination

Property Value
URI https://w3id.org/APD/traits/trait_0010112
alternative label cotyledon_position
description A plant embryo morphology trait [TO:0000064] which indicates if the cotyledons [PO:0020030] emerge from the seed coat [PO:0009088] at germination [GO:0009845].
Binary variable distinguishing between seedlings where the cotyledon remains within the seed coat versus emerges from the seed coat at germination.
comments
value type categorical variable
has narrower cotyledon_position_cryptocotylar A type of seed germination in which the cotyledons remain within the seed coat at germination.
cotyledon_position_phanerocotylar A type of seed germination in which the cotyledons emerge from the seed coat.
has broader plant embryo morphology trait
plant structure plant structure whole plant
leaf
measured characteristic organismal quality
structure
position
keyword cotyledon
seed coat
seed germination
plant embryo
seedling
plant embryo cotyledon
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010112

Cotyledon hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0010113
alternative label cotyledon_hairs
description A plant embryo cotyledon morphology trait [TO:0000775] which is the pilosity [PATO:0000066] of the cotyledons [PO:0020030].
Binary variable distinguishing between glabrous versus hairy cotyledons.
comments
value type categorical variable
has narrower cotyledon_hairs_glabrous Cotyledons lack hairs.
cotyledon_hairs_hairy Cotyledons have hairs.
has broader plant embryo morphology trait
plant structure plant structure whole plant
leaf
measured characteristic texture
pilosity
keyword cotyledon
plant embryo
hairy
seedling
plant embryo cotyledon
trichome
scope note
has exact match cotyledon pilosity
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010113

Hypocotyl hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0010114
alternative label seedling_hypocotyl_hairs
description A seedling hypocotyl morphology trait [TO:0000769] which is the pilosity [PATO:0000066] of the hypocotyl [PO:0020100].
Binary variable distinguishing between glabrous versus hairy hypocotyls, where the hypocotyl is the embryonic axis to which the cotyledons are attached.
comments
value type categorical variable
has narrower seedling_hypocotyl_hairs_glabrous Hypocotyl lacks hairs.
seedling_hypocotyl_hairs_hairy Hypocotyl has hairs.
has broader plant embryo morphology trait
plant structure plant structure whole plant
leaf
measured characteristic texture
pilosity
keyword seedling hypocotyl morphology trait
hypocotyl
plant embryo
hairy
seedling
trichome
scope note
has exact match hypocotyl pilosity
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note hypocotyl_type
is in scheme https://w3id.org/APD/traits
identifier trait_0010114

Seedling first true leaf type

Property Value
URI https://w3id.org/APD/traits/trait_0010160
alternative label seedling_first_node_leaf_type
description A plant embryo true leaf formation stage [PO:0001095] morphology trait which is whether the leaf [PO:0025034] at the first stem node [PO:0020141] above the epicotyl [PO:0020035] in a true leaf or a scale leaf.
Binary variable distinguishing between seedlings where the first leaf is scale-like (cataphyll) versus leaf-like.
comments
value type categorical variable
has narrower seedling_first_node_leaf_type_scale-like The leaf at the first stem node above the epicotyl is scale-like, also referred to as a cataphyll.
seedling_first_node_leaf_type_leaf-like The leaf at the first stem node above the epicotyl is leaf-like (broad).
has broader plant embryo morphology trait
plant structure plant structure whole plant
leaf
measured characteristic organismal quality
structure
keyword plant embryo true leaf formation stage
stem node
epicotyl
seedling
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seedling_first_leaf
is in scheme https://w3id.org/APD/traits
identifier trait_0010160

Seedling first node leaf count

Property Value
URI https://w3id.org/APD/traits/trait_0010161
alternative label seedling_first_node_leaf_count
description A plant embryo true leaf formation stage [PO:0001095] morphology trait which is the leaf count [NCIT:C25463] at the first stem node [PO:0020141] above the epicotyl [PO:0020035].
Binary variable distinguishing between seedlings where the leaves at the first node are single versus paired.
comments
value type categorical variable
has narrower seedling_first_node_leaf_count_single There is a single leaf at the first stem node above the epicotyl.
seedling_first_node_leaf_count_paired There is a pair of leaves at the first stem node above the epicotyl.
has broader plant embryo morphology trait
plant structure plant structure whole plant
leaf
measured characteristic has number of
count
keyword plant embryo true leaf formation stage
stem node
epicotyl
seedling
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note seedling_first_node
is in scheme https://w3id.org/APD/traits
identifier trait_0010161

Seedling germination location

Property Value
URI https://w3id.org/APD/traits/trait_0010162
alternative label seedling_germination_location
description A plant embryo cotyledon morphology trait [TO:0000775] which is the position [PATO:0000140] of seed germination [GO:0009845] relative to the soil surface.
Binary variable distinguishing between seedlings that germinate above versus below ground.
comments
value type categorical variable
has narrower seedling_germination_location_epigeal Germinant with one or more cotyledons emerging aboveground.
seedling_germination_location_hypogeal Germinant with all cotyledons remaining belowground.
has broader plant embryo morphology trait
plant structure plant structure whole plant
leaf
measured characteristic organismal quality
position
keyword seed germination
plant embryo
seedling
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0010162

Leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0011211
alternative label leaf_area
description A leaf area trait [TO:0000540] which is the 2-D [PATO:0001709] area [PATO:0001323] of the leaf lamina [PO:0020039] and petiole [PO:0020038]. For some species the area of a leaf analogue is reported, possibly a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335].
The 2-dimensional projected (scanned) area of the surface of a leaf or a leaf analogue (leaflet, cladode, phyllode).
comments This trait includes measurements of leaves and leaf analogues, including leaflets, cladodes, and phyllodes. When a measurement is made on a leaf analogue, the tissue measured should be documented as part of the methods or noted as a context or under measurement_remarks. Leaflet area can alternatively be mapped to the trait leaflet_area. It is most common for leaf area measurements to include the petiole; the measurement methods should indicate if the petiole is included in or excluded from the leaf area measurements. For terete leaves, leaf area should still be recorded as the 2-dimensional projected area.
value type continuous variable
unit mm2
square millimetre
minAllowedValue 0.1
maxAllowedValue 1e+07
has broader leaf size trait
plant structure leaf
measured characteristic area
2-D extent
size
keyword leaf lamina
petiole
leaflet
cladode
phyllode
scope note Plants with leaves.
has exact match leaf area trait
Leaf area
has close match
has related match leaf size
examples exact match: Leaf area [TOP25] (https://top-thesaurus.org/index)
exact match: Leaf area (in case of compound leaves: leaf, undefined if petiole in- or excluded) [TRY:3112] (https://www.try-db.org/de/de.php)
close match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards)
exact match: Leaf size [GIFT:4.4] (https://gift.uni-goettingen.de)
exact match: leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Average leaf area [BROT:6] (http://doi.org/10.1038/sdata.2018.135)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011211

Leaflet area

Property Value
URI https://w3id.org/APD/traits/trait_0011212
alternative label leaflet_area
description A leaf area trait [TO:0000540] which is the 2-D [PATO:0001709] area [PATO:0001323] of a single leaflet [PO:0020049] lamina [PO:0025060]. This measurement will not include the petiole [PO:0020038].
The 2-dimensional projected (scanned) area of the surface of the lamina of a single leaflet.
comments
value type continuous variable
unit mm2
square millimetre
minAllowedValue 0.1
maxAllowedValue 1e+06
has broader leaf size trait
plant structure leaf
measured characteristic area
2-D extent
size
keyword leaflet
leaf lamina
scope note
has exact match leaflet area
has close match
has related match leaf size
examples close match: leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
close match: Average leaf area [BROT:6] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011212

Leaf length

Property Value
URI https://w3id.org/APD/traits/trait_0011213
alternative label leaf_length
description Length [PATO:0000122] of the leaf [PO:0025034] (including the petiole [PO:0020038]), from the leaf apex [PO:0020137] to the leaf axil [PO:0009023] (leaf base [PO:0020040]).
Length of a leaf or a leaf analogue, from the leaf apex to the leaf axil (base), including the petiole.
comments If a leaf analogue (leaflet, cladode, or phyllode) is measured instead of a true leaf, this should be noted as a context or under measurement_remarks.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.1
maxAllowedValue 1e+05
has broader leaf size trait
plant structure leaf
measured characteristic length
size
keyword petiole
leaf apex
leaf base
leaf lamina
scope note Plants with leaves.
has exact match leaf length
has close match
has related match leaf size
examples exact match: Leaf length [TOP13] (https://top-thesaurus.org/index)
exact match: Leaf length [TRY:144] (https://www.try-db.org/de/de.php)
exact match: Leaf length [GIFT:4.6] (https://gift.uni-goettingen.de)
close match: maximum leaf length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
close match: minimum leaf length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
close match: Max_Blade_Length_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
close match: Max_Rachis_Length_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
close match: Max_Petiole_length_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011213

Leaf width

Property Value
URI https://w3id.org/APD/traits/trait_0011214
alternative label leaf_width
description The maximum [NCIT:C25564] width [PATO:0000921] axis of a leaf [PO:0025034] measured perpendicular to [PATO:0002434] the leaf length [PATO:0000122].
Maximum width axis of a leaf or a leaf analogue (a leaflet, cladode, or phyllode), measured perpendicular to the leaf length.
comments This trait refers to the single line marking the maximum width of the leaf lamina. For a compound leaf the correct measure is the maximum width of the total leaf blade. It is the width measure that will be reported in taxonomic treatments. It is not the effective_leaf_width, a trait that records the diameter of the largest circle that can be drawn within the leaf lamina. If a leaf analogue (leaflet, cladode, or phyllode) is measured instead of a true leaf, this should be noted as a context or under measurement_remarks.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 1e+05
has broader leaf size trait
plant structure leaf
measured characteristic width
size
keyword leaf lamina
scope note Plants with leaves.
has exact match leaf width
has close match
has related match leaf size
examples exact match: Leaf width [TOP17] (https://top-thesaurus.org/index)
exact match: Leaf width [TRY:145] (https://www.try-db.org/de/de.php)
exact match: Leaf width [GIFT:4.7] (https://gift.uni-goettingen.de)
close match: maximum leaf width [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
close match: minimum leaf width [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011214

Leaf thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011215
alternative label leaf_thickness
description The thickness [PATO:0000915] of the leaf lamina [PO:0020039], measured away from the midrib [PO:0020139].
Thickness of the leaf lamina, generally measured away from the midrib.
comments Leaf thickness measurements submitted from the ecological literature will generally avoid the midrib, but not avoid lower order leaf veins. In contrast, leaf thickness measurements from the taxonomic literature will usually include any veins or midribs, because in this literature leaf thickness is generally measured on succulent plants. The methods field should document whether the midrib was explicitly included, excluded, or ignored.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 51
has broader leaf size trait
plant structure leaf
measured characteristic thickness
size
keyword leaf lamina
scope note Plants with leaves.
has exact match leaf thickness
has close match
has related match
examples exact match: Leaf thickness [TOP20] (https://top-thesaurus.org/index)
exact match: Leaf thickness [TRY:46] (https://www.try-db.org/de/de.php)
exact match: Leaf thickness [GIFT:4.8] (https://gift.uni-goettingen.de)
exact match: leaf thickness [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011215

Leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0011216
alternative label leaf_dry_mass
description A leaf mass trait [TO:0000505] which is the dry [PATO:0001824] mass [PATO:0000125] of a leaf [PO:0025034] or leaf analogue (a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335]).
The oven dry mass of a leaf or leaf analogue (leaflet, cladode, phyllode).
comments This trait includes measurements of leaves and leaf analogues, including leaflets, cladodes, and phyllodes. When a measurement is made on a leaf analogue, the tissue measured should be documented as part of the methods or noted as a context or under measurement_remarks. Leaflet dry mass can alternatively be mapped to the trait leaflet_dry_mass. It is most common for leaf dry mass measurements to include the petiole; the measurement methods should indicate if the petiole is included in or excluded from the leaf dry mass measurements.
value type continuous variable
unit mg
milligram
minAllowedValue 0.01
maxAllowedValue 15000
has broader leaf mass trait
plant structure leaf
measured characteristic mass
keyword leaf weight, leaf mass
leaflet
cladode
phyllode
scope note
has exact match leaf dry weight, leaf dry mass
Leaf dry mass
has close match
has related match leaf size
examples exact match: Leaf dry mass [TOP40] (https://top-thesaurus.org/index)
exact match: Leaf dry mass (single leaf) [TRY:55] (https://www.try-db.org/de/de.php)
close match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards)
exact match: leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011216

Leaflet dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0011217
alternative label leaflet_dry_mass
description A leaf dry mass trait [TO:0001014] which is the dry mass of a single leaflet [PO:0020049]. This measurement will not include the petiole [PO:0020038].
The oven dry mass of a single leaflet.
comments
value type continuous variable
unit mg
milligram
minAllowedValue 0.005
maxAllowedValue 15000
has broader leaf mass trait
plant structure leaf
measured characteristic mass
keyword leaflet
leaf weight, leaf mass
scope note
has exact match
has close match leaflet weight
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011217

Leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0011218
alternative label leaf_fresh_mass
description A leaf mass trait [TO:0000505] which is the mass [PATO:0000125] of a fresh [EnvThes:21976] leaf [PO:0025034], indicating its mass at the time it is picked. For some species the mass of a leaf analogue is reported, possibly a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335].
The fresh mass of a leaf or leaf analogue (leaflet, cladode, phyllode).
comments This trait includes measurements of leaves and leaf analogues, including leaflets, cladodes, and phyllodes. When a measurement is made on a leaf analogue, the tissue measured should be documented as part of the methods or noted as a context or under measurement_remarks. It is most common for leaf fresh mass measurements to include the petiole; the measurement methods should indicate if the petiole is included in or excluded from the leaf fresh mass measurements. A source of variation for this trait will be how leaves are stored and if they are partially rehydrated before weighing; these details should be documented in the methods.
value type continuous variable
unit mg
milligram
minAllowedValue 0.1
maxAllowedValue 30000
has broader leaf mass trait
plant structure leaf
measured characteristic mass
keyword leaf weight, leaf mass
leaflet
cladode
phyllode
scope note Plants with leaves
has exact match leaf wet weight
has close match
has related match
examples exact match: Leaf fresh mass [TOP35] (https://top-thesaurus.org/index)
exact match: Leaf fresh mass [TRY:163] (https://www.try-db.org/de/de.php)
exact match: leaf fresh mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011218

Petiole length

Property Value
URI https://w3id.org/APD/traits/trait_0011219
alternative label petiole_length
description Length [PATO:0000122] of the petiole [PO:0020038]), from the leaf base [PO:0020040] to the leaf axil [PO:0009023].
Length of the petiole from the leaf base to the leaf axil.
comments
value type continuous variable
unit cm
minAllowedValue 0
maxAllowedValue 1000
has broader leaf size trait
plant structure leaf
measured characteristic length
size
keyword petiole
scope note
has exact match petiole length
Petiole length
has close match
has related match
examples
references no linked references
date created 31/05/2023
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011219

Petiole width

Property Value
URI https://w3id.org/APD/traits/trait_0011220
alternative label petiole_width
description Width [PATO:0000921] of the petiole [PO:0020038].
Width of the petiole.
comments
value type continuous variable
unit cm
minAllowedValue 0
maxAllowedValue 200
has broader leaf size trait
plant structure leaf
measured characteristic width
size
keyword petiole
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 31/05/2023
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011220

Number of leaflets per leaf

Property Value
URI https://w3id.org/APD/traits/trait_0011221
alternative label leaflet_count
description A leaf morphology trait [TO:0000748] which is the count [NCIT:C25463] of leaflets [PO:0020049] in a compound leaf [PO:0020043].
The count of leaflets that comprise a single leaf .
comments
value type continuous variable
unit {count}
minAllowedValue 1
maxAllowedValue 1000
has broader leaf morphology trait
plant structure plant structure leaf
leaflet
measured characteristic has number of
count
keyword leaflet
compound leaf morphology trait
compound leaf
scope note
has exact match leaflet number
has close match
has related match
examples exact match: Leaflet number per leaf [TRY:759] (https://www.try-db.org/de/de.php)
references no linked references
date created 31/03/2025
date modified 31/03/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011221

Leaflet length

Property Value
URI https://w3id.org/APD/traits/trait_0011222
alternative label leaflet_length
description A leaf morphology trait [TO:0000748] which is length [PATO:0000122] of a leaflet [PO:0020049], from the leaflet apex [PO:0020137] to where the leaflet attaches to the leaf [PO:0025034] rachis [PO:020055].
The length of a single leaflet comprising a botanical leaf.
comments This trait is intended to only apply to compound leaves, those with multiple leaflets per botanical leaf.
value type continuous variable
unit cm
minAllowedValue 0.1
maxAllowedValue 10
has broader leaf size trait
plant structure plant structure leaf
leaflet
measured characteristic length
size
keyword leaflet
compound leaf morphology trait
compound leaf
scope note applicable to plants with compound leaves.
has exact match leaflet length
has close match
has related match
examples exact match: Leaflet_length [TOP14] (https://top-thesaurus.org/index)
exact match: Leaflet length [TRY:985] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011222

Leaflet width

Property Value
URI https://w3id.org/APD/traits/trait_0011223
alternative label leaflet_width
description A leaf morphology trait [TO:0000748] which is the maximum [NCIT:C25564] width [PATO:0000921] axis of a single leaflet [PO:0020049] measured perpendicular to [PATO:0002434] the leaflet length [PATO:0000122].
Maximum width axis of a single leaflet, measured perpendicular to the leaflet length.
comments This trait is intended to only apply to compound leaves, those with multiple leaflets per botanical leaf.
value type continuous variable
unit cm
minAllowedValue 0.1
maxAllowedValue 10
has broader leaf size trait
plant structure plant structure leaf
leaflet
measured characteristic width
size
keyword leaflet
compound leaf morphology trait
compound leaf
scope note applicable to plants with compound leaves.
has exact match leaflet width
has close match
has related match
examples exact match: Leaflet_width [TOP18] (https://top-thesaurus.org/index)
exact match: Leaflet width [TRY:986] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011223

Leaf mass per area

Property Value
URI https://w3id.org/APD/traits/trait_0011230
alternative label leaf_mass_per_area
description A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of the leaf dry [PATO:0001824] mass [PATO:0000125] to the leaf [PO:0025034] area [PATO:0001323], measured on an entire leaf, including the entire leaf lamina [PO:0020039] and the petiole [PO:0020038].
The ratio of leaf dry mass to leaf area, measured on an entire leaf including the petiole; LMA. (The inverse of specific leaf area, SLA.)
comments Leaf mass per area includes measurements submitted as specific_leaf_area (SLA); they are simply inverted. Leaf mass per area is explicitly measured on an entire leaf, including the petiole. Measurements made on a piece of the leaf lamina, excluding petiole and midrib, are a separate trait leaf_lamina_mass_per_area. If measurements are made on the entire leaf lamina, but the petiole is excluded, this should be noted in the methods or under measurement_remarks.
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 1
maxAllowedValue 3000
has broader leaf size trait
leaf mass trait
plant structure leaf
measured characteristic area
ratio; proportion
mass
per unit area
area density
keyword leaf weight, leaf mass
biomass allocation
leaf lamina
petiole
scope note
has exact match specific leaf area
Leaf mass per leaf area
has close match
has related match
examples exact match: Leaf mass per area [TOP53] (https://top-thesaurus.org/index)
close match: Specific leaf area [TOP50] (https://top-thesaurus.org/index)
exact match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): undefined if petiole is in- or exclu [TRY:3117] (https://www.try-db.org/de/de.php)
exact match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole included [TRY:3116] (https://www.try-db.org/de/de.php)
close match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards)
exact match: Specific Leaf Area (SLA) [GIFT:4.1] (https://gift.uni-goettingen.de)
exact match: leaf area per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Average specific leaf area [BROT:8] (http://doi.org/10.1038/sdata.2018.135)
references Pérez-Harguindeguy 2013
Wright 2004
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note specific_leaf_area
is in scheme https://w3id.org/APD/traits
identifier trait_0011230

Leaf lamina mass per area

Property Value
URI https://w3id.org/APD/traits/trait_0011231
alternative label leaf_lamina_mass_per_area
description A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of the leaf dry [PATO:0001824] mass [PATO:0000125] to leaf [PO:0025034] area [PATO:0001323], measured on a piece of the leaf lamina [PO:0020039] located away from major leaf lamina veins [PO:0020138].
The ratio of leaf dry mass to leaf area, measured on a piece of the leaf lamina that does not include the petiole or major veins.
comments This trait only includes data made on subsamples of a leaf, usually a leaf punch sample associated with leaf strength measurements or from the herbivory literature. For whole-leaf LMA/SLA data, see the trait leaf_mass_per_area
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 0.1
maxAllowedValue 500
has broader leaf size trait
leaf mass trait
plant structure leaf
measured characteristic area
ratio; proportion
mass
per unit area
area density
keyword leaf weight, leaf mass
biomass allocation
leaf lamina
scope note Plants with leaves.
has exact match
has close match
has related match specific leaf area
examples related match: Leaf mass per area [TOP53] (https://top-thesaurus.org/index)
related match: Specific leaf area [TOP50] (https://top-thesaurus.org/index)
exact match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole excluded [TRY:3115] (https://www.try-db.org/de/de.php)
close match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA) petiole, rhachis and midrib excluded [TRY:3086] (https://www.try-db.org/de/de.php)
related match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): undefined if petiole is in- or exclu [TRY:3117] (https://www.try-db.org/de/de.php)
related match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole included [TRY:3116] (https://www.try-db.org/de/de.php)
related match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards)
references Pérez-Harguindeguy 2013
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011231

Leaf tissue density

Property Value
URI https://w3id.org/APD/traits/trait_0011232
alternative label leaf_density
description The ratio [PATO:0001470] of leaf [PO:0025034] dry [PATO:0001824] mass [PATO:0000125] to leaf fresh [EnvThes:21976] volume [PATO:0000918].
The ratio of leaf dry mass to leaf fresh volume.
comments
value type continuous variable
unit mg/mm3
milligram per cubic millimetre
minAllowedValue 0.05
maxAllowedValue 1.1
has broader leaf mass trait
plant structure leaf
measured characteristic mass density, density
mass
ratio; proportion
keyword leaf weight, leaf mass
biomass allocation
scope note
has exact match leaf mass density
has close match
has related match
examples exact match: Leaf density [TOP56] (https://top-thesaurus.org/index)
exact match: Leaf density (leaf tissue density, leaf dry mass per leaf volume) [TRY:48] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011232

Leaf area ratio (LAR)

Property Value
URI https://w3id.org/APD/traits/trait_0011260
alternative label leaf_area_ratio
description A leaf area trait [TO:0000540] which is the ratio [PATO:0001470] of a plant’s [PO:0000003] total [NCIT:C25304] leaf [PO:0025034] area [PATO:0001323] to total plant dry mass [TO_0000352].
The ratio of total plant leaf area to total plant dry mass; LAR.
comments Since the total plant dry mass is required for leaf area ratio measurements, this is standardly measured only on young plants or plants grown for experiments where the entire plant can be harvested. It is best practice for leaf area measurements to include the petiole.
value type continuous variable
unit mm2/mg
square millimetre per milligram
minAllowedValue 0.05
maxAllowedValue 100
has broader leaf size trait
plant structure plant structure leaf
whole plant
measured characteristic area
size
ratio; proportion
keyword whole plant dry mass
biomass allocation
scope note Plants with leaves.
has exact match leaf area ratio
has close match Leaf area
has related match
examples exact match: Leaf area ratio [TOP88] (https://top-thesaurus.org/index)
exact match: Leaf area per plant dry mass (leaf area ratio, LAR) [TRY:109] (https://www.try-db.org/de/de.php)
references Poorter & Remkes 1990
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011260

Leaf mass fraction

Property Value
URI https://w3id.org/APD/traits/trait_0011261
alternative label leaf_mass_fraction
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of a plant’s total leaf dry mass [APD:glossary/40080] to total plant dry mass [TO:0000352].
The ratio of leaf dry mass to total plant dry mass.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 1
has broader leaf mass trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword leaf weight, leaf mass
biomass allocation
whole plant dry mass
whole plant stem dry mass
scope note
has exact match
has close match
has related match
examples exact match: Leaf mass ratio [TOP85] (https://top-thesaurus.org/index)
exact match: Plant biomass and allometry: Leaf dry mass per plant dry mass (leaf weight ratio, LWR) [TRY:110] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011261

Leaf dry matter content (LDMC)

Property Value
URI https://w3id.org/APD/traits/trait_0011262
alternative label leaf_dry_matter_content
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of leaf dry mass [TO:0001014] to leaf [PO:0025034] fresh mass [APD:glossary/40044].
The ratio of leaf dry mass to leaf fresh mass; LDMC.
comments Two similar traits are leaf_water_content_per_dry_mass, the ratio of leaf water content to leaf dry mass and leaf_water_content_per_fresh_mass, the ratio of leaf water content to leaf fresh mass.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.001
maxAllowedValue 2
has broader leaf mass trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword leaf weight, leaf mass
biomass allocation
fresh mass
scope note
has exact match
has close match
has related match
examples exact match: Leaf dry matter content [TOP45] (https://top-thesaurus.org/index)
exact match: Leaf dry mass per leaf fresh mass (leaf dry matter content, LDMC) [TRY:47] (https://www.try-db.org/de/de.php)
exact match: Leaf dry matter content (LDMC) [GIFT:4.9] (https://gift.uni-goettingen.de)
exact match: leaf dry mass per leaf fresh mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Leaf dry matter content [BROT:14] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011262

Leaf fresh mass per leaf area

Property Value
URI https://w3id.org/APD/traits/trait_0011263
alternative label leaf_fresh_mass_per_area
description A leaf area trait [TO:0000540] which is the ratio [PATO:0001470] of the leaf [PO:0025034] fresh mass [APD:glossary/40044]. to the leaf area [PATO:0001323], measured on an entire leaf, including the entire leaf lamina [PO:0020039] and the petiole [PO:0020038].
The ratio of leaf fresh mass to leaf area, measured on an entire leaf including the petiole.
comments Leaf fresh mass per area is explicitly measured on an entire leaf, including the petiole.
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 200
maxAllowedValue 2000
has broader leaf size trait
leaf mass trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword leaf weight, leaf mass
biomass allocation
fresh mass
leaf lamina
petiole
scope note Plants with leaves.
has exact match
has close match
has related match
examples exact match: Leaf area per leaf fresh mass (specific leaf area (SLA or 1/LMA) based on leaf fresh mass) [TRY:125] (https://www.try-db.org/de/de.php)
references Pérez-Harguindeguy 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011263

Leaf water content per unit leaf area (leaf succulence)

Property Value
URI https://w3id.org/APD/traits/trait_0011264
alternative label leaf_water_content_per_area
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf area [PATO:0001323].
The ratio of the mass of water in a fresh leaf (leaf water content) to leaf surface area; leaf succulence.
comments
value type continuous variable
unit g/m2
gram per square metre
minAllowedValue 10
maxAllowedValue 5000
has broader leaf mass trait
plant structure leaf
measured characteristic mass
per unit area
ratio; proportion
keyword leaf weight, leaf mass
water
leaf water content
scope note
has exact match
has close match tissue succulence
has related match leaf moisture content
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011264

Leaf water content per unit leaf dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0011265
alternative label leaf_water_content_per_dry_mass
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf dry mass [TO:0001014].
The ratio of the mass of water in a fresh leaf (leaf water content) to leaf dry mass.
comments A similar trait that is recorded by a greater number of studies is leaf_dry_matter_content, the ratio of a leaf dry mass to leaf fresh mass.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 20
has broader leaf mass trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword leaf weight, leaf mass
water
leaf water content
scope note
has exact match
has close match
has related match leaf moisture content
examples exact match: Leaf water content per leaf dry mass (not saturated) [TRY:3120] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011265

Leaf water content per unit leaf fresh mass

Property Value
URI https://w3id.org/APD/traits/trait_0011266
alternative label leaf_water_content_per_fresh_mass
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf fresh mass [APD:glossary/40044].
The ratio of the mass of water in a fresh leaf (leaf water content) to leaf fresh mass
comments A similar trait that is recorded by a greater number of studies is leaf_dry_matter_content, the ratio of a leaf dry mass to leaf fresh mass.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader leaf mass trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword leaf weight, leaf mass
water
leaf water content
fresh mass
scope note
has exact match
has close match
has related match leaf moisture content
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011266

Leaf water content per unit saturated leaf mass

Property Value
URI https://w3id.org/APD/traits/trait_0011267
alternative label leaf_water_content_per_saturated_mass
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a saturated leaf [PO:0025034] to the mass of a saturated leaf. A saturated leaf is one that has been hydrated to its maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620].
The ratio of the mass of water in a leaf (leaf water content) to the mass of a saturated leaf. A saturated leaf is one that has been hydrated to its maximal water holding capacity and is at full turgor.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.1
maxAllowedValue 10
has broader leaf mass trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword leaf weight, leaf mass
water
leaf water content
fresh mass
water holding capacity
turgor
scope note
has exact match
has close match
has related match leaf moisture content
examples close match: Leaf water content per leaf dry mass (at saturation) [TRY:3122] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011267

Leaf cell wall fraction

Property Value
URI https://w3id.org/APD/traits/trait_0011268
alternative label leaf_cell_wall_fraction
description A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of cell wall [GO:0009505] dry mass to leaf dry mass [TO:0001014].
The ratio of leaf cell wall dry mass to leaf dry mass. This is the fraction of cell wall material that can be recovered from total leaf biomass.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.1
maxAllowedValue 1
has broader leaf mass trait
plant structure leaf
measured characteristic mass
mass fraction
ratio; proportion
keyword leaf weight, leaf mass
plant-type cell wall
biomass allocation
cell
scope note
has exact match
has close match
has related match
examples exact match: Leaf cell wall mass per leaf dry mass [TRY:3375] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011268

Leaf type

Property Value
URI https://w3id.org/APD/traits/trait_0011310
alternative label leaf_type
description A leaf shape trait [TO:0000492] which is inclusive [NCIT:C166400] categories describing the shape [PATO:0000052] of a leaf [PO:0025034] as broad, needle-like, or scale-like.
Variable distinguishing between general leaf form (shape) categories.
comments Terms identifying the plant organ responsible for photosynthesis are mapped to the trait plant_photosynthetic_organ, while leaf_form_general only includes terms pertaining to the general form of the leaf. The trait leaf_shape provides detailed vocabulary to describe a leaf’s shape.
value type categorical variable
has narrower leaf_type_broadleaf Flat leaf lamina, regardless of whether the leaves are or are not toothed, lobed, or compound.
leaf_type_needle Needle or awl-shaped leaf that is generally 3-dimensional.
leaf_type_scale Leaf reduced to a small scale.
leaf_type_leafless An adult stage plant that lacks leaves throughout the year.
has broader leaf shape trait
plant structure leaf
measured characteristic shape
structure
keyword leaf shape
scope note none
has exact match
has close match
has related match
examples related match: Leaf form [TOP225] (https://top-thesaurus.org/index)
related match: Leaf type [TRY:43] (https://www.try-db.org/de/de.php)
exact match: Leaf_form_1 [GIFT:4.11.1] (https://gift.uni-goettingen.de)
close match: Basic leaf shape [BROT:5] (http://doi.org/10.1038/sdata.2018.135)
references Pérez-Harguindeguy 2013
Burrows 2001
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011310

Leaf shape

Property Value
URI https://w3id.org/APD/traits/trait_0011311
alternative label leaf_shape
description A leaf shape trait [TO:0000492] which is the shape [PATO:0000052] and form of a leaf [PO:0025034], considering the ratio [PATO:0001470] of leaf length [PATO:0000122] to leaf width [PATO:0000921], the position [PATO:0000140] of the maximum [NCIT:C25564] width dimension relative to the length axis [NCIT:C25154], and the curvature [PATO:0001591] of the leaf margin [PO:0020128].
Shapes of a leaf or leaflet.
comments This trait includes terms to describe the shape of leaves, regardless of if they are broad leaves, scale-like leaves, or needle-like leaves. For compound leaves, it is generally the shape of the leaflet that is recorded. See the trait leaf_type for broad terms referencing leaf form (broadleaf, needle, scale) and plant_photosynthetic_organ for terms indicating which plant organ or tissue functions as the plant’s primary photosynthetic organ (e.g. phyllode, cladode, article).
value type categorical variable
has narrower leaf_shape_linear A 2-dimensional shape that is very long and narrow, has parallel sides and a length:width ratio between 12:1 and 6:1.
leaf_shape_narrowly_oblong A 2-dimensional shape that is long and narrow, has parallel sides and a length:width ratio between 6:1 and 3:1. (Synonyms, lorate)
leaf_shape_oblong A 2-dimensional shape that is somewhat elongated with parallel sides and a length:width ratio between 2:1 and 3:2.
leaf_shape_linear_elliptical A 2-dimensional shape that is a very narrow oval, with curved margins, lacking a distinct point, and a length:width ratio between 12:1 and 6:1.
leaf_shape_narrowly_elliptical A 2-dimensional shape that is a narrow oval, with curved margins, lacking a distinct point, and a length:width ratio between 6:1 and 3:1.
leaf_shape_elliptical A 2-dimensional shape that is an oval, with curved margins, lacking a distinct point, and a length:width ratio between 2:1 and 3:2. (Synonyms, oval)
leaf_shape_widely_elliptical A 2-dimensional shape that is a broad oval, with curved margins, lacking a distinct point, and a length:width ratio of approximately 6:5.
leaf_shape_orbicular A 2-dimensional shape that is approximately circular. (Synonyms, circular)
leaf_shape_oblate A 2-dimensional shape that is a broad oval, with curved margins, lacking a distinct point, and a length:width ratio of approximately 5:6. The petiole attaches along the broader side.
leaf_shape_narrowly_lanceolate A 2-dimensional shape that is very narrowly egg-shaped or teardrop-shaped, with a length:width ratio between 12:1 and 6:1. The petiole attaches near the wide, curved portion of the leaf lamina.
leaf_shape_lanceolate A 2-dimensional shape that is narrowly egg-shaped or teardrop-shaped, with a length:width ratio between 6:1 and 3:1. The petiole attaches near the wide, curved portion of the leaf lamina.
leaf_shape_narrowly_ovate A 2-dimensional shape that is narrowly egg-shaped, with a length:width ratio between 3:1 and 2:1. The petiole attaches near the wide, curved portion of the leaf lamina. (Synonyms, lance_ovate, widely_lanceolate)
leaf_shape_ovate A 2-dimensional shape that is egg-shaped, with a length:width ratio between 2:1 and 3:2. The petiole attaches near the wide, curved portion of the leaf lamina.
leaf_shape_widely_ovate A 2-dimensional shape that is broadly egg-shaped with a length:width ratio of approximately 6:5. The petiole attaches near the wide, curved portion of the leaf lamina.
leaf_shape_narrowly_oblanceolate A 2-dimensional shape that is very narrowly teardrop-shaped, with a length:width ratio between 12:1 and 6:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. (Synonyms, linear_oblanceolate)
leaf_shape_oblanceolate A 2-dimensional shape that is narrowly teardrop-shaped, with a length:width ratio between 6:1 and 3:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole.
leaf_shape_narrowly_obovate A 2-dimensional shape that is narrowly egg-shaped or teardrop-shaped with a length:width ratio between 3:1 and 2:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. (Synonyms, oblance-obovate)
leaf_shape_obovate A 2-dimensional shape that is egg-shaped with a length:width ratio between 2:1 and 3:2. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole.
leaf_shape_widely_obovate A 2-dimensional shape that is broadly egg-shaped with a length:width ratio of approximately 6:5. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole.
leaf_shape_narrowly_triangular A 2-dimensional shape that is narrowly triangular to wedge-shaped, with a length:width ratio 6:1 to 3:1 and the petiole attaching near the widest portion of the leaf. (Synonyms, triangular)
leaf_shape_triangular A 2-dimensional shape that is triangular to wedge-shaped, with a length:width ratio 2:1 to 3:2 and the petiole attaching near the widest portion of the leaf.
leaf_shape_deltate A 2-dimensional shape that is triangular to wedge-shaped, with a length:width ratio of 1:1 and the petiole attaching along one of the broad sides.
leaf_shape_obtriangular A 2-dimensional shape that is triangular to fan-shaped or wedge-shaped, with a length:width ratio 2:1 to 3:2 and the petiole attaching near the narrowest portion of the leaf. (Synonyms, flabelliform, fan-shaped, wedge-shaped, cuneate)
leaf_shape_narrowly_rhomboidal A 2-dimensional shape that is like a narrow diamond, with approximately straight edges, widest near the middle, and a length:width ratio 6:1 to 3:1. The petiole attaches near one of the points.
leaf_shape_rhomboidal A 2-dimensional shape that is approximately diamond or parallelogram-shaped, with approximately straight edges, widest near the middle, and a length:width ratio 2:1 to 3:2. The petiole attaches near one of the points. (Synonyms, rhombic, diamond)
leaf_shape_trullate A 2-dimensional shape that is approximately parallelogram-shaped, widest near where the petiole attaches (base), and with a length:width ratio 2:1 to 3:2.
leaf_shape_narrowly_obtrullate A 2-dimensional shape that is narrowly parallelogram-shaped, widest near the tip, and with a length:width ratio 3:1 to 6:1.
leaf_shape_obtrullate A 2-dimensional shape that is approximately parallelogram-shaped, widest near the tip, and with a length:width ratio 2:1 to 3:2.
leaf_shape_widely_obtrullate A 2-dimensional shape that is broadly parallelogram-shaped, widest near the tip, and with a length:width ratio of approximately 6:5.
leaf_shape_acicular A 2-dimensional or 3-dimensional shape with a needlelike tip, often round in cross-section, with margins straight and parallel, and a length:width ratio greater than 12:1. (Synonyms, needle-shaped)
leaf_shape_terete A 3-dimensional shape that is a long, narrow cylinder approximately circular in cross-section. Terete leaves often taper toward the tip. (Synonym, cylindrical)
leaf_shape_cordate A 2-dimensional heart shape, approximately ovate with a cordate base, with the petiole attaching at the notch between the lobes. (Synonyms, cordiform)
leaf_shape_obcordate A 2-dimensional heart shape, approximately ovate with a cordate apex, and the petiole attaching along the broad, curve side.
leaf_shape_falcate A 2-dimensional sickle-shape, approximately lanceolate to linear and curved on one side. (Synonyms, scimitar-shaped, sickle-shaped, falciform, crescent-shaped)
leaf_shape_reniform A 2-dimensional shape that is roughly kidney-shaped, wider than long with a rounded apex and two rounded, basal lobes that are smoothly concave at their intersection. (Synonyms, kidney-shaped)
leaf_shape_spathulate A 2-dimensional shape that can be approximately oblong, obovate, or oblanceolate in outline with a long attenuate base, like a spoon. (Synonyms, spoon-shaped, clavate)
leaf_shape_strap-shaped A 2-dimensional shape that is strap-shaped with a rounded tip and a length:width ratio greater than 12:1.
leaf_shape_ensiform A 2-dimensional shape that is sword-shaped with a pointed tip and a length:width ratio greater than 12:1. (Synonyms, sword-shaped)
leaf_shape_subulate A 2-dimensional shape that is awl-shaped, approximately narrowly oblong to narrowly triangular with a broad base. (Synonyms, awl-shaped)
leaf_shape_peltate A 2-dimensional orbicular shape, distinguished by the petiole attaching near the center of the leaf lamina.
leaf_shape_filiform A 2-dimensional shape that is long, thin, and typically flexuous, threadlike, filamentous.
has broader leaf shape trait
plant structure leaf
measured characteristic shape
length
curvature
structure
keyword leaf shape
leaf margin
scope note Plants with leaves.
has exact match leaf shape
has close match
has related match leaflet shape
examples close match: Leaf form [TOP225] (https://top-thesaurus.org/index)
close match: Leaf shape [TRY:154] (https://www.try-db.org/de/de.php)
references Simpson 2019
Ellis 2009
Systematics Association Committee for Descriptive Terminology 1962
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011311

Leaf base shape

Property Value
URI https://w3id.org/APD/traits/trait_0011312
alternative label leaf_base_shape
description A leaf shape trait [TO:0000492] which is the variation in shapes and forms of a leaf base [PO:0020040], especially considering the curvature [PATO:0001591] of the leaf margin [PO:0020128] near where it attaches to the petiole [PO:0020038].
The shape of the basal margin of a leaf.
comments
value type categorical variable
has narrower leaf_base_shape_acute Two sides of the leaf meet at an angle that is less than 90<U+00B0>.
leaf_base_shape_attenuate Basal leaf margin abruptly incurved (concave), the two sides intersecting at an angle of less than 45<U+00B0>.
leaf_base_shape_auriculate Basal leaf margin with two rounded lobes. The leaf margin above the lobes is concave. The petiole attaches in the incised region between the lobes.
leaf_base_shape_cordate Basal leaf margin with two rounded lobes and the petiole attaches in the incised region between the lobes. The leaf margin above the lobes is rounded.
leaf_base_shape_cuneate Basal leaf margin approximately straight, the two sides intersecting at an angle between 45<U+00B0> to 90<U+00B0>.
leaf_base_shape_hastate Basal leaf margin with two lobes oriented outward, such that they are at approximately 90<U+00B0> relative to the central axis and the leaf margin is fairly flat at the base.
leaf_base_shape_oblique The leaf base is asymmetrical, with the midrib not lying in the middle of the leaf where it attaches to the petiole.
leaf_base_shape_obtuse Two sides of the leaf meet at an angle that is greater than 90<U+00B0>.
leaf_base_shape_rounded Basal leaf margin is rounded.
leaf_base_shape_sagittate Basal leaf margin with two lobes oriented downward, away from apex. The petiole attaches in the concave region between the lobes.
leaf_base_shape_sheathing The leaf base is broad and envelops the stem.
leaf_base_shape_truncate The basal leaf margin is cut straight across with the two sides intersecting at approximately 180<U+00B0>.
has broader leaf shape trait
plant structure leaf
measured characteristic shape
curvature
keyword leaf shape
leaf base
leaf margin
petiole
scope note Plants with leaves.
has exact match Lamina shape
has close match
has related match
examples
references Simpson 2019
Ellis 2009
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011312

Leaf margin

Property Value
URI https://w3id.org/APD/traits/trait_0011313
alternative label leaf_margin
description A leaf shape trait [TO:0000492] which is the edge shape [PATO:0002267] of the leaf margin [PO:0020128], in particular the presence or absence of leaf lamina teeth [PO:0025518] and the shape [PATO:0000052] of the leaf lamina teeth (serrate, dentate, or crenate).
Shape of the leaf margin, referencing incisions or projections in the outermost 1/8 of the leaf lamina.
comments Terms describing leaf lamina projections and incisions that are greater than 1/8 of the distance to the midrib are captured under the traits leaf_lobation and leaf_division_detailed.
value type categorical variable
has narrower leaf_margin_entire Leaf margin lacks projections or indentations that extend less than 1/8 of the distance to the midrib.
leaf_margin_toothed Leaf margin has small projections or lobes along the edge of the leaf lamina. This is a broad term that includes serrate, dentate, and crenulate.
leaf_margin_toothed_serrate Type of toothed leaf margin with sharp teeth that are angled toward the tip of the leaf. (Synonyms, saw-toothed)
leaf_margin_toothed_dentate Type of toothed leaf margin with sharp teeth that point outwards at right angles to the leaf margin.
leaf_margin_toothed_crenate Type of toothed leaf margin with teeth that are rounded or obtuse, rather than having a sharp apex. [PO:0025519]
has broader leaf shape trait
plant structure leaf
measured characteristic shape
edge shape
curvature
keyword leaf shape
leaf margin
leaf lamina tooth
scope note Plants with broad leaves.
has exact match leaf margin morphology trait
has close match leaf margin serrated
has related match
examples exact match: Leaf margin type [TRY:963] (https://www.try-db.org/de/de.php)
exact match: Leaf_margin_1 [GIFT:4.12.1] (https://gift.uni-goettingen.de)
references Simpson 2019
Ellis 2009
Hickey 1973
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011313

Leaf margin posture

Property Value
URI https://w3id.org/APD/traits/trait_0011314
alternative label leaf_margin_posture
description A leaf shape trait [TO:0000492] which is the degree of curvature [PATO:0001591] or curling [PATO:0000405] of the leaf margin [PO:0020128] along the side of the leaf, at the leaf apex [PO:0020137], and at the leaf base [PO:0020040].
The posture (curvature or curling) of the leaf margin.
comments This trait captures the posture of the leaf margin, where posture refers to the placement of different parts of the leaf’s margin relative to a flat plane. This trait’s definition and allowable levels will be further refined once more data are available to map onto possible trait values.
value type categorical variable
has narrower leaf_margin_posture_involute Leaf margins along the side of the leaf are rolled upward, such that the margin of the adaxial (upper) leaf surface is rolled inward. (Synonym, inrolled, incurved)
leaf_margin_posture_revolute Leaf margins along the side of the leaf are rolled under or downward, such that the margin of the abaxial (lower) leaf surface is rolled inward. [PATO:0001963] (Synonym, recurved)
leaf_margin_posture_flat Leaf lamina, including margins, is a 2-dimensional plane. [PATO:0000407]
leaf_margin_posture_undulate Leaf margins are wavy in their 2-dimensional plane. (Synonyms, wavy, rippled, sinuate)
has broader leaf shape trait
plant structure leaf
measured characteristic shape
curvature
keyword leaf shape
curled
leaf margin
leaf apex
leaf base
scope note Plants with leaves
has exact match
has close match
has related match
examples
references Simpson 2019
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011314

Leaf lobation

Property Value
URI https://w3id.org/APD/traits/trait_0011315
alternative label leaf_lobation
description A leaf shape trait [TO:0000492] which is the presence or absence of leaf lamina lobes [PO:0025517] along the leaf margin [PO:0020128] and how far the lobes extend toward the midrib [PO:0020139].
A trait that indicates whether a leaf lamina or leaflets display any divisions or dissections on a 2-dimensional plane that extend greater than 1/8 of the distance to the midrib.
comments For terms pertaining to 2-dimensional leaf lamina divisions and incisions that are less than 1/8 of the distance to the midrib, see the trait leaf_margin. See also the trait leaf_division for more detailed leaf lobation (and leaf compoundness) trait values. Note that there might be some species that grade from deeply lobed simple leaves to compound leaves.
value type categorical variable
has narrower leaf_lobation_unlobed Leaf margin lacks projections or indentations that extend more than 1/8 of the distance to the midrib.
leaf_lobation_lobed Leaf margin has incisions, either rounded or sharp-tipped, that extend more than 1/8 of the distance to the midrib.
leaf_lobation_lobed_deep Leaf margin with lobes that extend more than halfway to the midrib. (Synonyms, parted, divided)
leaf_lobation_lobed_shallow Leaf margin with lobes that extend less than halfway to the midrib. (Synonyms, cleft, sinuate)
has broader leaf shape trait
plant structure leaf
measured characteristic shape
edge shape
curvature
keyword leaf shape
leaf lamina lobes
leaf margin
leaf midvein, midrib
scope note Plants with well-developed leaves.
has exact match leaf lamina lobes
has close match
has related match
examples
references Simpson 2019
Ellis 2009
Hickey 1973
Wikipedia 2022 Glossary of leaf morphology
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011315

Leaf compoundness

Property Value
URI https://w3id.org/APD/traits/trait_0011316
alternative label leaf_compoundness
description A leaf shape trait [TO:0000492] which is whether a leaf [PO:0025034] is a simple leaf [PO:0020042] or is divided into leaflets [PO:0020049] making it a compound leaf [PO:0020043].
A binary trait that indicates whether a leaf lamina is simple or divided into discontinuous leaflets (compound).
comments See also the trait ‘leaf_division’ for more detailed leaf compoundness trait values. Note that there might be some species that grade from deeply lobed simple leaves (see trait leaf_lobation) to compound leaves.
value type categorical variable
has narrower leaf_compoundness_simple A leaf with a single undivided blade. [PO:0020042]
leaf_compoundness_compound A leaf that is divided into multiple leaflets. [PO:0020043]
has broader leaf shape trait
plant structure leaf
measured characteristic shape
spatial pattern, distribution
keyword leaf shape
simple leaf
leaflet
compound leaf
leaf midvein, midrib
scope note Plants with well-developed leaves.
has exact match
has close match
has related match
examples exact match: Leaf compoundness [TOP222] (https://top-thesaurus.org/index)
exact match: Leaf compoundness [TRY:17] (https://www.try-db.org/de/de.php)
exact match: leaf compoundness [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Leaf division degree [BROT:2] (http://doi.org/10.1038/sdata.2018.135)
references Ellis 2009
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011316

Leaf divisions

Property Value
URI https://w3id.org/APD/traits/trait_0011317
alternative label leaf_lamina_division
description A leaf shape trait [TO:0000492] which indicates types [NCIT:C25284] of lobed leaves [PO:0025517] and compound leaves [PO:0020043], including the arrangement [NCIT:C61532] of the lobes or leaflets [PO:0020049] with respect to the midrib [PO:0020139] or other primary leaf veins [PO:0025413] and the number of orders of axes [NCIT:C25154] across which there are incisions or divisions.
A trait that captures the types of lobed leaves and compound leaves, including the arrangement of the lobes or leaflets relative to the midrib or other primary veins and the number of orders of axes across which there are incisions or divisions.
comments This trait provides detailed description of leaf divisions, beyond what is captured in leaf_compoundness (simple versus compound) and leaf_lobation (entire versus lobed). The term ‘divided’ is meant to indicate lobing, such as ‘trichotomously divided’, but as ‘divided’ is also used in some context to indicate the divisions of leaves into leaflets, it is avoided as a trait value for this trait.
value type categorical variable
has narrower leaf_lamina_division_pinnately_lobed Leaf with divisions that are symmetrical across a central axis, usually the midvein.
leaf_lamina_division_palmately_lobed Leaf with divisions that radiate from a single point, usually the basal junction of major veins.
leaf_lamina_division_pinnatifid Leaf with lobes that extend less than half-way to the midrib.
leaf_lamina_division_bipinnatifid Leaf that is twice pinnatifid, with two series of lobes that extend less than half-way to the midrib.
leaf_lamina_division_pinnatipartite Leaf with lobes that extend more than half-way to the midrib.
leaf_lamina_division_pinnatisect Leaf with lobes that extend nearly to the midrib but the divisions are not separated into distinct leaflets.
leaf_lamina_division_bipinnatipartite Leaf that is twice pinnatipartite, with two series of lobes that extend more than half-way to the midrib.
leaf_lamina_division_bipinnatisect Leaf that is twice pinnatisect, with two series of lobes that extend nearly to the midrib but the divisions are not separated into distinct leaflets.
leaf_lamina_division_trichotomously_lobed Leaf divided repeatedly into three lobes.
leaf_lamina_division_dichotomously_lobed Leaf divided repeatedly into two lobes.
leaf_lamina_division_bipartite Leaf divided into two lobes.
leaf_lamina_division_tripartite Leaf divided into three lobes.
leaf_lamina_division_pinnately_compound Leaf divided into leaflets that attach along a central axis. [PATO_0000410]
leaf_lamina_division_palmately_compound Leaf divided into leaflets that attach to a central point. (Synonyms, digitate)
leaf_lamina_division_bipinnate Leaf that is twice pinnate, having pinnate leaflets that are themselves pinnately divided.
leaf_lamina_division_tripinnate Leaf that is three times pinnate, with pinnate leaflets that are twice more pinnately divided.
leaf_lamina_division_trifoliate Leaf divided into three leaflets. (Synonyms, ternate, trifoliolate)
has broader leaf shape trait
plant structure leaf
measured characteristic spatial pattern, distribution
shape
arrangement
keyword leaf shape
leaf lamina lobes
compound leaf
leaflet
leaf midvein, midrib
leaf margin
pinnate
scope note Plants with broad leaves.
has exact match
has close match
has related match
examples related match: Leaf compoundness [TOP222] (https://top-thesaurus.org/index)
related match: Leaf compoundness [TRY:17] (https://www.try-db.org/de/de.php)
references Simpson 2019
Ellis 2009
Hickey 1973
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note leaf_division
is in scheme https://w3id.org/APD/traits
identifier trait_0011317

Leaf lamina posture (leaf 3-dimensionality)

Property Value
URI https://w3id.org/APD/traits/trait_0011318
alternative label leaf_posture_numeric
description A leaf shape trait [TO:0000492] which is the maximum [NCIT:C25564] perpendicular [PATO:0002434] height [PATO:0000119] of a leaf [PO:0025034] placed on a flat surface, indicating the 3-D extent [PATO:0001710] of the leaf. This generally corresponds with whether the leaf is curled [FLOPO:0010468].
The maximum perpendicular height of a leaf when positioned on a flat surface.
comments A measure of how planar versus 3-dimensional a leaf is. The term posture refers to the placement of different parts of the leaf’s lamina relative to a flat plane.
value type continuous variable
unit mm
millimetre
minAllowedValue 0
maxAllowedValue 100
has broader leaf shape trait
plant structure leaf
measured characteristic height
shape
3-D extent
keyword leaf shape
leaf lamina
scope note Plants with leaves
has exact match
has close match
has related match
examples
references Simpson 2019
Grootemaat 2017
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note leaf_curliness
is in scheme https://w3id.org/APD/traits
identifier trait_0011318

Leaf lamina posture (leaf 3-dimensional shape)

Property Value
URI https://w3id.org/APD/traits/trait_0011319
alternative label leaf_lamina_posture
description A leaf shape trait [TO:0000492] which is the 3-D shape [PATO:0002266] of the surface [NCIT:C25245] of the leaf lamina [PO:0020039], including its degree of curvature [PATO:0001591] and whether it is bent [PATO:0000617].
The 3-dimension shape of the surface of the leaf lamina, including its degree of curvature and bending.
comments
value type categorical variable
has narrower leaf_lamina_posture_concave A longitudinal posture term indicating the leaf lamina is curved upward along the central axis, such that the upper leaf lamina surface is concave.
leaf_lamina_posture_conduplicate A longitudinal posture term indicating the leaf lamina is folded upward along the central axis. (Synonym, folded)
leaf_lamina_posture_convex A longitudinal posture term indicating the leaf lamina is curved downward along the central axis, such that the lower leaf lamina surface is concave. (Synonym, concavoconvex, planoconvex)
leaf_lamina_posture_convolute Leaf lamina is contorted. (Synonym, twisted)
leaf_lamina_posture_flat The leaf lamina is a 2-dimensional plane.
leaf_lamina_posture_incurved Leaf tip is curved upward.
leaf_lamina_posture_involute A longitudinal posture term indicating the outer portions of the leaf lamina are rolled inward or upward.
leaf_lamina_posture_plicate Leaf margin is pleated, with a series of longitudinal folds.
leaf_lamina_posture_recurved Leaf tip is curved outward or downward. [PATO:0002211]
leaf_lamina_posture_revolute A longitudinal posture term indicating the outer portions of the leaf lamina are rolled outward or downward.
leaf_lamina_posture_undulate The leaf lamina surface is rippled or wavy in its 2-dimensional plane. [PATO:0000967] (Synonyms, wavy, rippled, sinuate)
has broader leaf shape trait
plant structure leaf
measured characteristic shape
3-D shape
3-D extent
keyword leaf shape
leaf lamina
scope note Plants with leaves
has exact match
has close match
has related match
examples
references no linked references
date created 07/06/2023
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011319

Leaf glaucousness

Property Value
URI https://w3id.org/APD/traits/trait_0011360
alternative label leaf_glaucousness
description A leaf morphology trait [TO:0000748] which is the presence or absence of epicuticular wax [PO:0025388] on the leaf lamina [PO:0020039] of a fully expanded adult phase leaf [PO:0006340], giving the leaf lamina surface a white-coloured [PATO:0000014] or blue-coloured waxy [PATO:0002384] appearance, termed glaucous.
Variable describing whether mature adult phase leaves have a glaucous appearance, the colour due to being covered with a smooth, usually whitish to bluish epicuticular wax that can easily be rubbed off.
comments The accompanying methods should specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type categorical variable
has narrower leaf_glaucousness_glaucous Mature (fully expanded) leaves on adult plants have a waxy coating, giving them a glaucous appearance. (Synonyms, waxy bloom, pruinose)
leaf_glaucousness_not_glaucous Mature (fully expanded) leaves on adult plants do not have a glaucous appearance.
has broader leaf morphology trait
plant structure leaf
measured characteristic colour / color
texture
keyword epicuticular wax
leaf lamina
waxy
scope note Plants with leaves.
has exact match leaf gloss
has close match
has related match
examples
references Simpson 2019
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note glaucous
is in scheme https://w3id.org/APD/traits
identifier trait_0011360

Mature leaf hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0011361
alternative label leaf_hairs_adult_leaves
description A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of a fully expanded adult phase leaf [PO:0006340].
Variable describing whether mature (fully expanded) adult phase leaves on adult plants have hairs (trichomes).
comments This trait captures the presence or absence of hairs on mature (fully expanded), adult phase leaves on adult plants. Best practice is to sample leaves that are recently fully expanded, since older leaves have often shed many of their hairs. The accompanying methods should specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type categorical variable
has narrower leaf_hairs_adult_leaves_glabrous Mature (fully expanded) leaves on adult plants do not have hairs (trichomes).
leaf_hairs_adult_leaves_hairy Mature (fully expanded) leaves on adult plants have hairs (trichomes).
has broader leaf morphology trait
plant structure leaf
measured characteristic texture
pilosity
keyword leaf pubescence
leaf trichome
leaf lamina
vestiture
hairy
adult vascular leaf
pilosity
trichome
scope note Plants with leaves.
has exact match leaf lamina pubescence
has close match
has related match leaf trichome density
examples close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
references Hanley 2007
Simpson 2019
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note leaf_hairs_adult
is in scheme https://w3id.org/APD/traits
identifier trait_0011361

Juvenile phase leaf hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0011362
alternative label leaf_hairs_juvenile_leaves
description A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of a juvenile vascular leaf [PO:0006339] (on adult or juvenile plants).
Variable describing whether fully expanded juvenile phase leaves (on adult or juvenile plants) have hairs (trichomes).
comments This trait captures the presence or absence of hairs on fully-expanded juvenile phase leaves. It is best practice for the methods to specify whether these leaves were sampled from adult or juvenile plants. The accompanying methods should also specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type categorical variable
has narrower leaf_hairs_juvenile_leaves_glabrous Juvenile phase leaves do not have hairs (trichomes).
leaf_hairs_juvenile_leaves_hairy Juvenile phase leaves have hairs (trichomes).
has broader leaf morphology trait
plant structure leaf
measured characteristic texture
pilosity
keyword leaf pubescence
leaf trichome
leaf lamina
vestiture
hairy
juvenile vascular leaf
pilosity
trichome
scope note Plants with leaves.
has exact match
has close match leaf lamina pubescence
has related match leaf trichome density
examples close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
references Hanley 2007
Simpson 2019
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note leaf_hairs_juvenile
is in scheme https://w3id.org/APD/traits
identifier trait_0011362

Immature leaf hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0011363
alternative label leaf_hairs_immature_leaves
description A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of an adult phase leaf [PO:0006340] still in the vascular leaf expansion stage [PO:0001052].
Variable describing whether immature (still expanding) adult phase leaves on adult plants have hairs (trichomes).
comments This trait captures the presence or absence of hairs on immature (still expanding) leaves on adult plants. The accompanying methods should specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type categorical variable
has narrower leaf_hairs_immature_leaves_glabrous Immature (still expanding) leaves do not have hairs (trichomes).
leaf_hairs_immature_leaves_hairy Immature (still expanding) leaves have hairs (trichomes).
has broader leaf morphology trait
plant structure leaf
measured characteristic texture
pilosity
keyword leaf pubescence
leaf trichome
leaf lamina
vestiture
hairy
adult vascular leaf
vascular leaf expansion stage
trichome
scope note Plants with leaves.
has exact match
has close match leaf lamina pubescence
has related match leaf trichome density
examples close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
references Hanley 2007
Simpson 2019
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011363

Leaf surface colour

Property Value
URI https://w3id.org/APD/traits/trait_0011364
alternative label leaf_surface_colour
description A leaf morphology trait [TO:0000748] which is which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the leaf lamina [PO:0020039].
Human visible dominant colour of the surface of a leaf.
comments
value type categorical variable
has narrower leaf_surface_colour_green Leaf lamina surface is green.
leaf_surface_colour_grey_green Leaf lamina surface is grey-green.
leaf_surface_colour_dark_green Leaf lamina surface is dark green.
leaf_surface_colour_pale_green Leaf lamina surface is pale green.
leaf_surface_colour_pale_grey_green Leaf lamina surface is pale grey-green.
leaf_surface_colour_dark_grey_green Leaf lamina surface is dark grey-green.
leaf_surface_colour_red Leaf lamina surface is red.
leaf_surface_colour_purple Leaf lamina surface is purple.
has broader leaf morphology trait
plant structure leaf
measured characteristic colour / color
keyword leaf lamina
adult vascular leaf
scope note
has exact match leaf lamina color
has close match
has related match
examples exact match: Leaf color [TRY:582] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011364

Leaf surface reflectivity

Property Value
URI https://w3id.org/APD/traits/trait_0011365
alternative label leaf_surface_reflectivity
description A leaf morphology trait [TO:0000748] which is the reflectivity [PATO:0001297] of the surface [NCIT:C25245] of a mature [PATO:0001701] leaf lamina [PO:0020039].
Surface reflectivity of a leaf, indicating whether the leaf lamina is shiny or dull.
comments There will be overlap between this trait and the trait leaf_glaucousness. This trait can be used to document surface reflectivity more broadly, not only distinguishing between leaf lamina that are glaucous versus not glaucous.
value type categorical variable
has narrower leaf_surface_reflectivity_dull Leaf surface appears dull, indicating it has a low reflectivity.
leaf_surface_reflectivity_shiny Leaf surface appears shiny or glossy, indicating it has a high reflectivity. (Synonyms, glossy, lustrous, polished)
has broader leaf morphology trait
plant structure leaf
measured characteristic texture
morphology
reflectivity
keyword leaf lamina
adult vascular leaf
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011365

Leaf phyllotaxis

Property Value
URI https://w3id.org/APD/traits/trait_0011410
alternative label leaf_phyllotaxis
description A leaf [PO:0025034] spatial pattern [PATO:0000060] trait which is the phyllotaxy [TO:0006014] of leaves, indicating a leaf’s position [TO:0000541] relative to other leaves at a stem node [PO:0020141].
Leaf arrangement at a stem node, indicating the relative number and position of leaves at a single stem note.
comments This trait only includes true phyllotaxis, the arrangement of leaves at a single node, while the trait leaf arrangement indicates how the leaves are presented along the entire shoot system. See the trait stem_growth_habit for whole-plant leaf pattern terms, such as rosette.
value type categorical variable
has narrower leaf_phyllotaxis_alternate A single leaf is attached at each stem node.
leaf_phyllotaxis_opposite A pair of leaves are attached opposite to one another at each stem node.
leaf_phyllotaxis_whorled Three or more leaves are attached at each stem node, arranged in a symmetrical pattern around the stem. (Synonyms, verticillate)
has broader leaf position trait
plant structure leaf
measured characteristic spatial pattern, distribution
arrangement
keyword phyllotaxy
stem node
scope note Plants with leaves.
has exact match phyllotaxy
has close match
has related match
examples close match: Leaf distribution along the shoot axis (arrangement type) [TRY:16] (https://www.try-db.org/de/de.php)
close match: Leaf_arrangement [GIFT:4.15.1] (https://gift.uni-goettingen.de)
references Simpson 2019
Ellis 2009
Wikipedia 2022 Leaf arrangement on the stem
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011410

Leaf arrangement

Property Value
URI https://w3id.org/APD/traits/trait_0011411
alternative label leaf_arrangement
description A leaf [PO:0025034] spatial pattern [PATO:0000060] trait which is the position [TO:0000541] and arrangement [NCIT:C61532] of leaves between stem nodes [PO:0020141], indicating how leaves align and cluster along the vegetative shoot system [PO:0025607].
Leaf arrangement across stem nodes, indicating the 3-dimensional arrangement of leaves along the length of the stem.
comments Leaf arrangement, as defined for this trait, indicates patterns of leaf arrangement across nodes, including both formal terms (decussate, distichous, spiral) as well as indications of leaf clustering due to variation in internode length. See the trait leaf phyllotaxis for terms describing leaf arrangement at a single node. See the trait stem_growth_habit for whole-plant leaf pattern terms, such as rosette.
value type categorical variable
has narrower leaf_arrangement_decussate When sequential pairs of opposite leaves attach at right angles to the pair above or below.
leaf_arrangement_distichous When alternate or opposite leaves are arranged in two opposite rows.
leaf_arrangement_scattered When leaves are attached along the entire length of stem.
leaf_arrangement_spiral When alternate leaves are attached in a progressive spiral pattern along the stem.
leaf_arrangement_fasciculate A tight bundle or cluster of leaves attaches to the stem in a single location, generally due to the shortening of internodes. (Synonym, tufted)
leaf_arrangement_clustered When leaves are in any type of cluster, where the type of cluster is not further specified. The leaves may be clustered toward the base of the plant (a rosette), toward the top of the plant, toward the end of branches, or in distinct tuft (fascicles), but more information is not provided.
leaf_arrangement_clustered_basal When leaves are clustered or crowded to the base of the stem, arranged in a rosette or other basal clustering. (Synonym, rosette)
leaf_arrangement_crowded When internode length varies considerably along a stem, such that leaves appear crowded or tufted. (Synonym, congested)
leaf_arrangement_rosette When leaves radiate from a central point at or near ground level.
leaf_arrangement_imbricate When leaves lay tightly against the stem and are overlapping.
has broader leaf position trait
plant structure leaf
measured characteristic spatial pattern, distribution
arrangement
keyword stem node
vegetative shoot system
phyllotaxy
scope note Plants with leaves.
has exact match
has close match
has related match
examples close match: Leaf distribution along the shoot axis (arrangement type) [TRY:16] (https://www.try-db.org/de/de.php)
close match: Leaf_arrangement [GIFT:4.15.1] (https://gift.uni-goettingen.de)
references Simpson 2019
Ellis 2009
Wikipedia 2022 Leaf arrangement on the stem
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011411

Leaf axil angle

Property Value
URI https://w3id.org/APD/traits/trait_0011412
alternative label leaf_axil_angle
description A leaf position trait [TO:0000541] which is the angle [PATO:0002326] of a leaf axil [PO:0009023], the space between a leaf [PO:0025034] and the stem [PO:0009047] to which it is attached.
The angle between a shoot and a leaf that connects to the shoot axis.
comments This trait records the angle between the upper surface of a leaf and the stem to which it is attached. The slope of the leaf lamina relative to the solar zenith is a separate trait, leaf_inclination_angle.
value type continuous variable
unit deg
plane angle degree
minAllowedValue 0
maxAllowedValue 180
has broader leaf position trait
plant structure leaf
measured characteristic angle
orientation
spatial pattern, distribution
keyword leaf axil
leaf attitude
leaf lamina
leaf angle
scope note Plants with leaves
has exact match leaf angle
has close match
has related match
examples related match: Leaf angle (inclination, orientation) [TRY:3] (https://www.try-db.org/de/de.php)
references no linked references
date created 31/10/2022
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011412

Leaf inclination angle

Property Value
URI https://w3id.org/APD/traits/trait_0011413
alternative label leaf_inclination_angle
description A leaf position trait [TO:0000541] which is the inclination angle [PATO:0002326] of the adaxial [PATO:0002047] leaf lamina [PO:0020039] surface relative to a horizontal [PATO:0001855] plane, such that an adaxial leaf surface facing the solar zenith [ECSO:00002723] has an angle of 0.
The inclination angle of the upper (adaxial) leaf lamina surface relative to a horizontal plane, such that a leaf surface facing the solar zenith has an angle of 0.
comments This trait records the slope of the leaf lamina relative to the ground surface. The angle between a stem and the leaf lamina is a separate trait, leaf_axil_angle. Complanate leaves can be scored as having a leaf_inclination_angle of 0.
value type continuous variable
unit deg
plane angle degree
minAllowedValue -90
maxAllowedValue 90
has broader leaf position trait
plant structure leaf
measured characteristic angle
orientation
spatial pattern, distribution
keyword leaf lamina
solar zenith angle
leaf attitude
scope note Plants with leaves
has exact match
has close match leaf attitude
has related match
examples exact match: Leaf angle (inclination, orientation) [TRY:3] (https://www.try-db.org/de/de.php)
references Niinemets 2010
Falster 2003b
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note leaf_angle
is in scheme https://w3id.org/APD/traits
identifier trait_0011413

Leaf pendulousness

Property Value
URI https://w3id.org/APD/traits/trait_0011414
alternative label leaf_pendulousness
description A leaf position trait [TO:0000541] which indicates whether leaves [PO:0025034] are pendulous [APD:glossary/40066], hanging loosely downward and possessing the ability to flutter.
Binary variable describing whether leaves are pendulous, hanging loosely downward and possessing the ability to flutter.
comments This is a binary categorical trait, scoring whether leaves are pendulous and therefore hanging loosely downward and able to flutter. Two additional traits, leaf_inclination_angle and leaf_axil_angle document the actual angle of the leaf lamina relative to, respectively, the solar zenith and the stem.
value type categorical variable
has narrower leaf_pendulousness_pendulous Leaves are pendulous.
leaf_pendulousness_not_pendulous Leaves are not pendulous.
has broader leaf position trait
plant structure leaf
measured characteristic spatial pattern, distribution
orientation
angle
keyword leaf angle
pendulous
scope note Plants with leaves
has exact match
has close match leaf attitude
has related match
examples close match: Leaf angle (inclination, orientation) [TRY:3] (https://www.try-db.org/de/de.php)
references Curtis 2012
date created 14/07/2021
date modified 05/02/2024
date reviewed 31/10/2022
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Ian J. Wright
Peter Vesk
Gregory J. Jordan
Andy Leigh
Patricia Lu-Irving
Lily Dun
David Coleman
change note pendulous_leaves
is in scheme https://w3id.org/APD/traits
identifier trait_0011414

Cuticle thickness on the lower leaf surface

Property Value
URI https://w3id.org/APD/traits/trait_0011510
alternative label leaf_cuticle_thickness_abaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of cuticle [PO:0000022] on the abaxial [PATO:0002046] (lower) leaf lamina [PO:0020039].
Thickness of the cuticle on the lower leaf surface.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 0.1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword plant cuticle
abaxial
leaf lamina
scope note
has exact match
has close match plant cuticle thickness
has related match
examples close match: Leaf cuticula thickness [TRY:716] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note lower_cuticle_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011510

Cuticle thickness on the upper leaf surface

Property Value
URI https://w3id.org/APD/traits/trait_0011511
alternative label leaf_cuticle_thickness_adaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of cuticle [PO:0000022] on the adaxial [PATO:0002047] (upper) leaf lamina [PO:0020039].
Thickness of the cuticle on the upper leaf surface.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 0.1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword plant cuticle
adaxial
leaf lamina
scope note
has exact match
has close match plant cuticle thickness
has related match
examples close match: Leaf cuticula thickness [TRY:716] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note upper_cuticle_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011511

Leaf epidermis thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011512
alternative label leaf_epidermis_thickness
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] , in samples where the leaf lamina [PO:0020039] side (abaxial [PATO:0002046] versus adaxial [PATO:0002047]) is not specified.
Thickness of the epidermis, where the leaf lamina side is not specified.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
epidermis
leaf lamina
abaxial
adaxial
scope note
has exact match
has close match epidermis thickness
has related match
examples exact match: Leaf epidermis thickness [TRY:101] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note epidermis_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011512

Lower leaf side epidermis thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011513
alternative label leaf_epidermis_thickness_abaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039].
Thickness of the epidermis on the lower side of the leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
epidermis
abaxial
leaf lamina
scope note
has exact match
has close match epidermis thickness
has related match
examples close match: Leaf epidermis thickness [TRY:101] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note epidermis_thickness_lower_leaf_surface
is in scheme https://w3id.org/APD/traits
identifier trait_0011513

Upper leaf side epidermis thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011514
alternative label leaf_epidermis_thickness_adaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039].
Thickness of the epidermis on the upper side of the leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
epidermis
adaxial
leaf lamina
scope note
has exact match
has close match epidermis thickness
has related match
examples close match: Leaf epidermis thickness [TRY:101] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note epidermis_thickness_upper_leaf_surface
is in scheme https://w3id.org/APD/traits
identifier trait_0011514

Average leaf epidermal cell density

Property Value
URI https://w3id.org/APD/traits/trait_0011515
alternative label leaf_epidermal_cell_density_both_sides
description A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] to leaf lamina [PO:0020039] area [PATO_0001323], averaged across the abaxial [PATO:0002046] (lower) and adaxial [PATO:0002047] (upper) sides of the leaf lamina.
Epidermal cell density averaged across the upper and lower sides of the leaf.
comments
value type continuous variable
unit {count}/mm2
reciprocal square millimetre
minAllowedValue 100
maxAllowedValue 10000
has broader plant cell morphology trait
leaf morphology trait
plant structure leaf
measured characteristic has number of
count density
ratio; proportion
count
per unit area
keyword cell
epidermal cell
abaxial
adaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note epidermal_cell_density_both_sides
is in scheme https://w3id.org/APD/traits
identifier trait_0011515

Lower leaf side epidermal cell density

Property Value
URI https://w3id.org/APD/traits/trait_0011516
alternative label leaf_epidermal_cell_density_abaxial
description A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039] to leaf lamina area [PATO_0001323].
Epidermal cell density on the lower side of the leaf.
comments
value type continuous variable
unit {count}/mm2
reciprocal square millimetre
minAllowedValue 100
maxAllowedValue 10000
has broader plant cell morphology trait
leaf morphology trait
plant structure leaf
measured characteristic has number of
count density
ratio; proportion
count
per unit area
keyword cell
epidermal cell
abaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note epidermal_cell_density_abaxial
is in scheme https://w3id.org/APD/traits
identifier trait_0011516

Upper leaf side epidermal cell density

Property Value
URI https://w3id.org/APD/traits/trait_0011517
alternative label leaf_epidermal_cell_density_adaxial
description A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039] to leaf lamina area [PATO_0001323].
Epidermal cell density on the upper side of the leaf.
comments
value type continuous variable
unit {count}/mm2
reciprocal square millimetre
minAllowedValue 100
maxAllowedValue 10000
has broader plant cell morphology trait
leaf morphology trait
plant structure leaf
measured characteristic has number of
count density
ratio; proportion
count
per unit area
keyword cell
epidermal cell
adaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note epidermal_cell_density_adaxial
is in scheme https://w3id.org/APD/traits
identifier trait_0011517

Lower leaf side hypodermis thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011518
alternative label leaf_hypodermis_thickness_abaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of hypodermis [PO:0005051] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039].
Thickness of the hypodermis on the lower side of the leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
hypodermis
abaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples close match: Leaf hypodermis thickness [TRY:806] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note lower_hypodermis_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011518

Upper leaf side hypodermis thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011519
alternative label leaf_hypodermis_thickness_adaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of hypodermis [PO:0005051] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039].
Thickness of the hypodermis on the upper side of the leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 1
maxAllowedValue 100
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
hypodermis
adaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples close match: Leaf hypodermis thickness [TRY:806] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note upper_hypodermis_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011519

Lower palisade mesophyll thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011520
alternative label leaf_palisade_tissue_thickness_abaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of palisade mesophyll [PO:0005648] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039].
Thickness of palisade mesophyll tissue on the lower side of the leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 10
maxAllowedValue 1000
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
palisade mesophyll
abaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note lower_palisade_cell_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011520

Upper palisade mesophyll thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011521
alternative label leaf_palisade_tissue_thickness_adaxial
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of palisade mesophyll [PO:0005648] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039].
Thickness of palisade mesophyll tissue on the upper side of the leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 10
maxAllowedValue 1000
has broader portion of plant tissue morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
palisade mesophyll
adaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note upper_palisade_cell_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011521

Palisade cell length

Property Value
URI https://w3id.org/APD/traits/trait_0011522
alternative label leaf_palisade_cell_length
description A plant cell morphology trait [TO:0002683] which is the length [PATO:0000122] of individual palisade mesophyll cells [PO:0006206].
Length of individual leaf palisade cells.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 10
maxAllowedValue 1000
has broader plant cell morphology trait
leaf morphology trait
plant structure leaf
measured characteristic length
keyword cell
palisade mesophyll cell
leaf lamina
scope note
has exact match
has close match
has related match plant cell length
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note palisade_cell_length
is in scheme https://w3id.org/APD/traits
identifier trait_0011522

Palisade cell width

Property Value
URI https://w3id.org/APD/traits/trait_0011523
alternative label leaf_palisade_cell_width
description A plant cell morphology trait [TO:0002683] which is the width [PATO:0000921] of individual palisade mesophyll cells [PO:0006206].
Width of individual leaf palisade cells.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 1
maxAllowedValue 50
has broader plant cell morphology trait
leaf morphology trait
plant structure leaf
measured characteristic width
keyword cell
palisade mesophyll cell
leaf lamina
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note palisade_cell_width
is in scheme https://w3id.org/APD/traits
identifier trait_0011523

Number of layers of palisade cells

Property Value
URI https://w3id.org/APD/traits/trait_0011524
alternative label leaf_palisade_layer_number
description A portion of plant tissue morphology trait [TO:0000843] which is the count [NCIT:C25463] of cell layers [UBERON:0000119] of palisade mesophyll cells [PO:0006206].
The number of layers of palisade cells in a leaf.
comments
value type continuous variable
unit {count}
minAllowedValue 1
maxAllowedValue 10
has broader portion of plant tissue morphology trait
leaf morphology trait
plant structure leaf
measured characteristic has number of
count
keyword cell
palisade mesophyll cell
palisade mesophyll
leaf lamina
tissue
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note palisade_layer_number
is in scheme https://w3id.org/APD/traits
identifier trait_0011524

Spongy mesophyll cell thickness

Property Value
URI https://w3id.org/APD/traits/trait_0011525
alternative label leaf_spongy_mesophyll_thickness
description A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of spongy mesophyll [PO:0005647] tissue [CARO:0000043] within the leaf lamina [PO:0020039].
Thickness of spongy mesophyll tissue in a leaf.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 10
maxAllowedValue 1000
has broader portion of plant tissue morphology trait
leaf morphology trait
plant structure leaf
measured characteristic thickness
keyword tissue
spongy mesophyll
leaf lamina
scope note
has exact match
has close match
has related match
examples exact match: Leaf mesophyll spongiophyll thickness [TRY:805] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note spongy_mesophyll_thickness
is in scheme https://w3id.org/APD/traits
identifier trait_0011525

Cell cross-sectional area

Property Value
URI https://w3id.org/APD/traits/trait_0011526
alternative label cell_cross-sectional_area
description A plant cell size [TO:0002684] trait which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a cell [CL:0000000].
Cell cross-sectional area.
comments This trait can be used to map the area of cell types that are not already explicitly listed as their own trait. The cell type and tissue/organ being measured needs to be specified under measurement_remarks or as a method_context.
value type continuous variable
unit um2
square micrometre
minAllowedValue 100
maxAllowedValue 10000
has broader plant cell morphology trait
plant structure whole plant
measured characteristic area
2-D extent
size
keyword cell
plant cell size
cross section
scope note
has exact match
has close match
has related match
examples close match: Leaf epidermis cell area [TRY:338] (https://www.try-db.org/de/de.php)
close match: Leaf mesophyll cell area [TRY:573] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011526

Stomatal density on the lower leaf surface

Property Value
URI https://w3id.org/APD/traits/trait_0011610
alternative label leaf_stomatal_density_abaxial
description A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] on the abaxial [PATO:0002046] (lower) leaf lamina [PO:0020039] to the leaf lamina area [PATO:0001323].
Density of stomata on the lower leaf surface.
comments
value type continuous variable
unit {count}/mm2
reciprocal square millimetre
minAllowedValue 1
maxAllowedValue 1000
has broader leaf stomatal complex morphology trait
plant structure leaf
measured characteristic has number of
count density
ratio; proportion
count
per unit area
keyword leaf stomatal complex frequency
stomata; stomatal pore
abaxial
leaf lamina
scope note
has exact match abaxial stomatal frequency
has close match Stomatal density
has related match
examples exact match: Stomata density abaxial [TOP371] (https://top-thesaurus.org/index)
close match: Stomata density [TRY:63] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stomatal_density_abaxial
is in scheme https://w3id.org/APD/traits
identifier trait_0011610

Stomatal density on the upper leaf surface

Property Value
URI https://w3id.org/APD/traits/trait_0011611
alternative label leaf_stomatal_density_adaxial
description A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] on the adaxial [PATO:0002047] (upper) leaf lamina [PO:0020039] to the leaf lamina area [PATO:0001323].
Density of stomata on the upper leaf surface.
comments
value type continuous variable
unit {count}/mm2
reciprocal square millimetre
minAllowedValue 0
maxAllowedValue 1000
has broader leaf stomatal complex morphology trait
plant structure leaf
measured characteristic has number of
count density
ratio; proportion
count
per unit area
keyword leaf stomatal complex frequency
stomata; stomatal pore
adaxial
leaf lamina
scope note
has exact match adaxial stomatal frequency
has close match Stomatal density
has related match
examples exact match: Stomata density adaxial [TOP372] (https://top-thesaurus.org/index)
close match: Stomata density [TRY:63] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stomatal_density_adaxial
is in scheme https://w3id.org/APD/traits
identifier trait_0011611

Stomatal density averaged across both leaf surfaces

Property Value
URI https://w3id.org/APD/traits/trait_0011612
alternative label leaf_stomatal_density_average
description A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] to leaf lamina [PO:0020039] area [PATO:0001323], averaged the across the abaxial [PATO:0002046] and adaxial [PATO:0002047] sides of the leaf lamina.
Density of stomata averaged across both leaf surfaces.
comments
value type continuous variable
unit {count}/mm2
reciprocal square millimetre
minAllowedValue 0
maxAllowedValue 1000
has broader leaf stomatal complex morphology trait
plant structure leaf
measured characteristic has number of
count density
ratio; proportion
count
per unit area
keyword leaf stomatal complex frequency
stomata; stomatal pore
abaxial
adaxial
leaf lamina
scope note
has exact match Stomatal density
has close match
has related match
examples exact match: Stomata density trait [TOP370] (https://top-thesaurus.org/index)
exact match: Stomata density [TRY:63] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stomatal_density_average
is in scheme https://w3id.org/APD/traits
identifier trait_0011612

Stomatal distribution

Property Value
URI https://w3id.org/APD/traits/trait_0011613
alternative label leaf_stomatal_distribution
description A leaf stomatal complex frequency trait [TO:0000566] which indicates whether stomata [PO:0008032] occur on the abaxial [PATO:0002046] and/or adaxial [PATO:0002047] leaf lamina [PO:0020039].
Distribution of stomatal across the two leaf surfaces.
comments
value type categorical variable
has narrower leaf_stomatal_distribution_amphistomatic Leaves with stomata on both leaf surfaces.
leaf_stomatal_distribution_amphistomatic_uneven Leaves with stomata on both leaf surfaces but more stomata on the lower leaf surface.
leaf_stomatal_distribution_epistomatic Leaves that only have stomata on the upper (adaxial) leaf surface.
leaf_stomatal_distribution_hypostomatic Leaves that only have stomata on the lower (abaxial) leaf surface.
has broader leaf stomatal complex morphology trait
plant structure leaf
measured characteristic spatial pattern, distribution
keyword leaf stomatal complex frequency
stomata; stomatal pore
abaxial
adaxial
leaf lamina
scope note
has exact match
has close match
has related match
examples exact match: Stomata distribution to ab- and adaxial leaf surfaces [TRY:781] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stomatal_distribution
is in scheme https://w3id.org/APD/traits
identifier trait_0011613

Stomatal hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0011614
alternative label leaf_stomatal_hairs
description A leaf stomatal complex frequency trait [TO:0000566] which is the pilosity [PATO:0000066] (hairiness) of the stomatal pores [PO:0008032].
Variable indicating if stomata are covered by hairs and, if hairy, the density of hairs present.
comments
value type categorical variable
has narrower leaf_stomatal_hairs_stomata_covered_with_dense_hairs Stomata densely covered by hairs.
leaf_stomatal_hairs_stomata_not_covered_by_hairs Stomata not hair-covered.
has broader leaf stomatal complex morphology trait
plant structure leaf
measured characteristic texture
pilosity
keyword leaf stomatal complex frequency
stomata; stomatal pore
hairy
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stomatal_hairs
is in scheme https://w3id.org/APD/traits
identifier trait_0011614

Guard cell length

Property Value
URI https://w3id.org/APD/traits/trait_0011615
alternative label leaf_guard_cell_length
description A leaf stomatal complex frequency trait [TO:0000566] which indicates the length [PATO:0000122] of a guard cell [PO:0000293].
Length of guard cells.
comments
value type continuous variable
unit um
micrometre
minAllowedValue 10
maxAllowedValue 200
has broader leaf stomatal complex morphology trait
plant structure leaf
measured characteristic length
keyword cell
leaf stomatal complex frequency
guard cell
stomata; stomatal pore
scope note
has exact match
has close match
has related match plant cell length
examples exact match: Stomata guard cell size [TRY:733] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note guard_cell_length
is in scheme https://w3id.org/APD/traits
identifier trait_0011615

Leaf visible light transmission

Property Value
URI https://w3id.org/APD/traits/trait_0011710
alternative label leaf_transmission
description A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is transmitted [SWEET:procWave/Transmission] through a leaf.
Proportion of incoming visible light (between 400-700 nm) that is transmitted through a leaf.
comments
value type continuous variable
unit umol/umol
micromole per micromole
minAllowedValue 0.001
maxAllowedValue 1
has broader leaf optical properties trait
plant structure leaf
measured characteristic dimensionless ratio
ratio; proportion
electromagnetic property
radiation
keyword optics
visible light
transmission
photosynthetically active radiation
scope note
has exact match
has close match
has related match
examples exact match: Leaf light transmittance [TRY:535] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011710

Leaf visible light absorption

Property Value
URI https://w3id.org/APD/traits/trait_0011711
alternative label leaf_absorption
description A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is absorbed [SWEET:procWave/Absorption] by a leaf.
Proportion of incoming visible light (between 400-700 nm) that is absorbed by a leaf.
comments
value type continuous variable
unit umol/umol
micromole per micromole
minAllowedValue 0.01
maxAllowedValue 1
has broader leaf optical properties trait
plant structure leaf
measured characteristic dimensionless ratio
ratio; proportion
electromagnetic property
radiation
keyword optics
visible light
absorption
photosynthetically active radiation
scope note
has exact match
has close match
has related match
examples exact match: Leaf light absorption (extinction) [TRY:10] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011711

Leaf visible light reflection

Property Value
URI https://w3id.org/APD/traits/trait_0011712
alternative label leaf_reflectance
description A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is reflected [SWEET:procWave/Reflection] by a leaf.
Proportion of incoming visible light (between 400-700 nm) that is reflected by a leaf.
comments
value type continuous variable
unit umol/umol
micromole per micromole
minAllowedValue 0.01
maxAllowedValue 1
has broader leaf optical properties trait
plant structure leaf
measured characteristic dimensionless ratio
ratio; proportion
electromagnetic property
radiation
keyword optics
visible light
reflection
photosynthetically active radiation
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011712

Leaf infra-red light reflection

Property Value
URI https://w3id.org/APD/traits/trait_0011713
alternative label leaf_reflectance_near_infrared
description A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming infrared radiation [ENVO:21001214] that is reflected [SWEET:procWave/Reflection] by a leaf.
Proportion of incoming near infra-red light (between 750-10500 nm) that is reflected by a leaf.
comments
value type continuous variable
unit umol/umol
micromole per micromole
minAllowedValue 0.1
maxAllowedValue 1
has broader leaf optical properties trait
plant structure leaf
measured characteristic dimensionless ratio
ratio; proportion
electromagnetic property
radiation
keyword infrared radiation
optics
reflection
radiation
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011713

Stem cross-sectional area

Property Value
URI https://w3id.org/APD/traits/trait_0011811
alternative label stem_cross_sectional_area
description A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a stem [PO:0009047] segment which includes the bark [PO:0004518].
Cross-sectional area of a stem, including both wood and bark.
comments This is not a trait itself, since it is dependent on the location sampled. However, values might be reported that are effectively contexts for another trait, such as bark thickness or Huber value.
value type continuous variable
unit mm2
square millimetre
minAllowedValue 1
maxAllowedValue 1000
has broader stem morphology trait
plant structure stem
measured characteristic area
size
keyword cross section
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stem_cross_section_area
is in scheme https://w3id.org/APD/traits
identifier trait_0011811

Wood cross-sectional area

Property Value
URI https://w3id.org/APD/traits/trait_0011812
alternative label sapwood_cross_sectional_area
description A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a stem [PO:0009047] segment from which the bark [PO:0004518] has been removed; this includes both sapwood [PO:0004513] and, if present heartwood [PO:0004512].
Cross-sectional area of a stem without bark.
comments This is not a trait itself, since it is dependent on the location sampled. However, values might be reported that are effectively contexts for another trait, such as bark thickness or Huber value. Although it is defined as the diameter of the entire stem without bark, it is generally measured on narrow branches that have not yet developed heartwood and therefore is the cross-section of sapwood.
value type continuous variable
unit mm2
square millimetre
minAllowedValue 1
maxAllowedValue 1000
has broader stem morphology trait
plant structure stem
measured characteristic area
size
keyword sapwood
cross section
wood, secondary xylem
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note stem_cross_sectional_area_no_bark
is in scheme https://w3id.org/APD/traits
identifier trait_0011812

Terminal twig cross-sectional area

Property Value
URI https://w3id.org/APD/traits/trait_0011813
alternative label branch_terminal_twig_cross_sectional_area
description A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of the most distal [PATO:0001234] branch [PO:0025073].
Cross-sectional area of the terminal twig, measured just beyond the final branching node.
comments
value type continuous variable
unit mm2
square millimetre
minAllowedValue 0.05
maxAllowedValue 50
has broader stem morphology trait
plant structure stem
measured characteristic area
size
keyword branch
cross section
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note twig_area
is in scheme https://w3id.org/APD/traits
identifier trait_0011813

Terminal twig length

Property Value
URI https://w3id.org/APD/traits/trait_0011814
alternative label branch_terminal_twig_length
description A stem morphology trait [TO:0000361] which is the length [PATO:0000122] of the most distal [PATO:0001234] branch [PO:0025073].
Length of the terminal twig, the length of stem beyond the final branching node.
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 1
maxAllowedValue 1000
has broader stem morphology trait
plant structure stem
measured characteristic length
keyword branch
scope note
has exact match
has close match
has related match
examples exact match: Twig length [TRY:1202] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note twig_length
is in scheme https://w3id.org/APD/traits
identifier trait_0011814

Wood density

Property Value
URI https://w3id.org/APD/traits/trait_0011815
alternative label wood_density
description A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of a unit of wood [NCIT:C93062] from a stem [PO:0009047] assessed after drying to its volume [PATO:0000918] assessed without drying.
The ratio of wood dry mass to wood fresh volume, measured on a stem segment without bark.
comments
value type continuous variable
unit mg/mm3
milligram per cubic millimetre
minAllowedValue 0.05
maxAllowedValue 1.4
has broader stem morphology trait
plant structure stem
measured characteristic mass
volume density
ratio; proportion
mass density, density
keyword wood
wood, secondary xylem
biomass allocation
scope note
has exact match secondary xylem volumetric density, wood volumetric density
Wood density
has close match
has related match
examples exact match: Wood density [TOP284] (https://top-thesaurus.org/index)
exact match: Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density [TRY:4] (https://www.try-db.org/de/de.php)
exact match: Stem specific density (SSD) [GIFT:4.3] (https://gift.uni-goettingen.de)
exact match: stem wood density [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Stem specific density [BROT:11] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011815

Herbaceous stem density

Property Value
URI https://w3id.org/APD/traits/trait_0011816
alternative label stem_density
description A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of a unit of herbaceous [PATO:0002352] stem [PO:0009047] assessed after drying to its volume [PATO:0000918] assessed without drying.
The ratio of stem dry mass to stem fresh volume, measured on an herbaceous stem segment.
comments This trait is specifically for non-woody or partially woody stems that otherwise are outliers for wood density
value type continuous variable
unit mg/mm3
milligram per cubic millimetre
minAllowedValue 0.1
maxAllowedValue 1.4
has broader stem morphology trait
plant structure stem
measured characteristic mass
volume density
ratio; proportion
mass density, density
keyword herbaceous
biomass allocation
scope note
has exact match
has close match
has related match
examples exact match: Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density [TRY:4] (https://www.try-db.org/de/de.php)
exact match: Stem specific density (SSD) [GIFT:4.3] (https://gift.uni-goettingen.de)
exact match: stem wood density [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Stem specific density [BROT:11] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011816

Stem hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0011817
alternative label stem_hairs
description A stem morphology trait [TO:0000361] which is the presence or absence of trichomes [PO:0006504] (hairs) on a stem [PO_0009047].
Variable describing whether plant stems have hairs (trichomes).
comments As stems often lose hairs as they age, the context field should indicate the stem age.
value type categorical variable
has narrower stem_hairs_glabrous Adult plant stems do not have hairs (trichomes).
stem_hairs_hairy Adult plant stems have hairs (trichomes).
has broader stem morphology trait
plant structure stem
measured characteristic texture
pilosity
keyword trichome
scope note
has exact match stem hairiness
has close match
has related match
examples close match: Stem emergences (pubescence, hairs, spines, thorns) [TRY:1256] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011817

Xylem to leaf area ratio (XLA)

Property Value
URI https://w3id.org/APD/traits/trait_0011910
alternative label xylem_to_leaf_area_ratio
description A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of xylem vessel [PO:0025417] area [PATO:0001323] in a petiole [PO:0020038] to the attached leaf lamina [PO:0020039] area.
For a single leaf, the ratio of xyem cross-sectional area at the base of the petiole to the leaf lamina area.
comments This trait is similar to Huber value, but at the level of a single leaf, not the entire branch.
value type continuous variable
unit cm2{xylem}/mm2{leaf}
minAllowedValue 0.01
maxAllowedValue 100
has broader leaf morphology trait
plant structure leaf
measured characteristic area
size
dimensionless ratio
ratio; proportion
keyword biomass allocation
xylem
water transport
leaf lamina
scope note
has exact match
has close match
has related match
examples
references Blackman 2023
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011910

Huber value

Property Value
URI https://w3id.org/APD/traits/trait_0011911
alternative label huber_value
description A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of sapwood [PO:0004513] area [PATO:0001323] to the total distal [PATO:0001234] leaf [PO:0025034] area.
For a single branch, the ratio of sapwood cross-sectional area at the base of the branch to the branch’s total leaf area.
comments
value type continuous variable
unit mm2{sapwood}/mm2{leaf}
square millimetre per square millimetre
minAllowedValue 1e-06
maxAllowedValue 0.2
has broader stem morphology trait
plant structure plant structure leaf
stem
measured characteristic area
size
dimensionless ratio
ratio; proportion
keyword sapwood
biomass allocation
scope note
has exact match
has close match
has related match
examples exact match: Stem, branch or twig sapwood cross-sectional area per supported leaf area (Huber value) [TRY:171] (https://www.try-db.org/de/de.php)
references Tyree & Ewers 1991
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011911

Leaf dry mass to stem dry mass ratio

Property Value
URI https://w3id.org/APD/traits/trait_0011912
alternative label leaf_mass_to_stem_mass_ratio
description A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of a plant’s total leaf [PO:0025034] dry [PATO:0001824] mass [PATO:0000125] to its total stem [PO:0009047] and branch [PO:0025073] dry mass.
The ratio of whole plant leaf dry mass to whole plant stem dry mass.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 10
has broader leaf mass trait
stem mass trait
plant structure plant structure plant structure leaf
stem
whole plant
measured characteristic mass
mass fraction
ratio; proportion
keyword biomass allocation
whole plant stem dry mass
whole plant leaf dry mass
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note leaf_mass_to_stem_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0011912

Stem dry mass to vegetative shoot dry mass ratio (support fraction)

Property Value
URI https://w3id.org/APD/traits/trait_0011913
alternative label stem_mass_to_shoot_mass_ratio
description A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of stem [PO:0009047] and branch [PO:0025073] dry [PATO:0001824] mass [PATO:0000125] to vegetative shoot system [PO:0025607] dry mass.
The ratio of whole plant stem dry mass to whole plant vegetative shoot dry mass. That is, the proportion of aboveground vegetative biomass that is comprised of stems and branches, rather than leaves; support fraction.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0
maxAllowedValue 1
has broader leaf mass trait
stem mass trait
plant structure plant structure plant structure leaf
stem
whole plant
measured characteristic mass
mass fraction
ratio; proportion
keyword branch
vegetative shoot system
biomass allocation
whole plant stem dry mass
whole plant leaf dry mass
scope note
has exact match
has close match
has related match
examples close match: Plant biomass and allometry: Supportive structure (stem, twig, leaf petiole, leaf sheeth) dry mass p [TRY:3059] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note support_fraction
is in scheme https://w3id.org/APD/traits
identifier trait_0011913

Side branch dry mass to whole plant dry mass ratio

Property Value
URI https://w3id.org/APD/traits/trait_0011914
alternative label branch_mass_fraction
description A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of side branch [PO:0025073] dry [PATO:0001824] mass [PATO:0000125] to whole plant [PO:0000003] dry mass.
The ratio of side branch dry mass to whole plant dry mass. That is, the proportion of total plant biomass that is comprised of all branches off the main shoot axis.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0
maxAllowedValue 1
has broader stem mass trait
plant structure plant structure stem
whole plant
measured characteristic mass
mass fraction
ratio; proportion
keyword branch
biomass allocation
whole plant dry mass
scope note
has exact match
has close match
has related match
examples exact match: Plant biomass and allometry: Branch dry mass per plant dry mass [TRY:137] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011914

Stem dry matter content (SDMC)

Property Value
URI https://w3id.org/APD/traits/trait_0011915
alternative label stem_dry_matter_content
description A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of stem [PO:0009047] (or branch [PO:0025073]) dry [PATO:0001824] mass [PATO:0000125] to stem (or branch) fresh mass [APD:glossary/40044].
The ratio of stem (or branch) dry mass to stem (or branch) fresh mass; SDMC.
comments A related trait is stem_water_content_per_saturated_mass.
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 100
maxAllowedValue 1000
has broader stem mass trait
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword branch
fresh mass
water
scope note
has exact match
has close match
has related match
examples exact match: Stem dry mass per stem fresh mass (stem dry matter content, StDMC) [TRY:1181] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011915

Stem water content per unit saturated stem mass

Property Value
URI https://w3id.org/APD/traits/trait_0011916
alternative label stem_water_content_per_saturated_mass
description A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a stem [PO:0009047] to stem fresh mass [APD:glossary/40044] in a stem that is at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620].
The ratio of the mass of water in a stem (stem water content) to total stem mass, measured in a saturated stem. A saturated stem is one that has been hydrated to its maximal water holding capacity and is at full turgor.
comments A related trait is stem_dry_matter_content.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.1
maxAllowedValue 10
has broader stem mass trait
plant structure stem
measured characteristic mass
mass fraction
ratio; proportion
keyword water
water holding capacity
turgor
fresh mass
scope note
has exact match
has close match
has related match stem moisture content
examples close match: Wood (sapwood) water content [TRY:1097] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011916

Stem mass fraction

Property Value
URI https://w3id.org/APD/traits/trait_0011917
alternative label stem_mass_fraction
description A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of whole plant [PO:0000003] stem [PO:0009047] dry [PATO:0001824] mass [PATO:0000125] to whole plant dry mass [APD:glossary/40080].
The ratio of stem dry mass to whole plant dry mass. That is, the proportion of total plant biomass that is comprised of stems and branches.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 1
has broader stem mass trait
plant structure plant structure stem
whole plant
measured characteristic mass
mass fraction
ratio; proportion
keyword biomass allocation
whole plant stem dry mass
whole plant dry mass
scope note
has exact match
has close match
has related match
examples exact match: Plant biomass and allometry: Stem dry mass per plant dry mass [TRY:136] (https://www.try-db.org/de/de.php)
related match: stem dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011917

Stem saturated water content per unit stem dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0011918
alternative label stem_saturated_water_content_per_dry_mass
description A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a saturated stem [PO:0009047] to stem dry [PATO:0001824] mass [PATO:0000125], where a saturated stem is one at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091].
The ratio of the mass of water in a saturated stem to stem dry mass.
comments Similar traits include stem_dry_matter_content, the ratio of a stem dry mass to stem fresh mass and stem_water_content_per_saturated_mass the ratio of the mass of water in a stem to saturated stem mass.
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 20
has broader stem mass trait
plant structure plant structure stem
whole plant
measured characteristic mass
mass fraction
ratio; proportion
keyword biomass allocation
whole plant stem dry mass
whole plant dry mass
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0011918

Bark morphology, Eucalyptus

Property Value
URI https://w3id.org/APD/traits/trait_0012010
alternative label bark_morphology_eucalyptus
description A bark morphology trait which is the texture [PATO:0000150] of the bark [PO:0004518] of Eucalyptus taxa.
Broad categories that describe the texture of Eucalyptus bark.
comments This trait just records the morphology of taxa in the genera Eucalyptus, Corymbia, and Angophora.
value type categorical variable
has narrower bark_morphology_eucalyptus_eucalypt_box Eucalyptus, Corymbia or Angophora taxa with ‘Box’ type bark.
bark_morphology_eucalyptus_eucalypt_gum Eucalyptus, Corymbia or Angophora taxa with ‘Gum’ type bark.
bark_morphology_eucalyptus_eucalypt_stocking Eucalyptus, Corymbia or Angophora taxa with ‘Gum’ type bark above a rough barked stocking.
bark_morphology_eucalyptus_eucalypt_ironbark Eucalyptus, Corymbia or Angophora taxa with ‘Ironbark’ type bark.
bark_morphology_eucalyptus_eucalypt_peppermint Eucalyptus, Corymbia or Angophora taxa with ‘Peppermint’ type bark.
bark_morphology_eucalyptus_eucalypt_stringybark Eucalyptus, Corymbia or Angophora taxa with ‘Stringy’ type bark.
bark_morphology_eucalyptus_eucalypt_ribbonbark Eucalyptus, Corymbia or Angophora taxa with ‘Gum’ type bark that sheds in ribbons.
has broader bark morphology trait
plant structure bark
measured characteristic structure
texture
scope note taxa in the genera Eucalyptus, Corymbia, and Angophora
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012010

Bark thickness

Property Value
URI https://w3id.org/APD/traits/trait_0012011
alternative label bark_thickness
description A bark morphology trait which is the thickness [PATO:0000915] of the bark [PO:0004518]. The corresponding stem [PO:0009047] diameter [PATO:0001334] should also be recorded so that a scaled measure, the bark thickness to stem diameter ratio, can also be calculated.
Thickness of the bark on the stem.
comments The corresponding stem diameter should also be recorded so that a scaled measure, the bark thickness to stem diameter ratio, can also be calculated.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 200
has broader bark morphology trait
plant structure bark
measured characteristic thickness
scope note
has exact match Bark thickness
has close match
has related match
examples exact match: Bark thickness [TOP283] (https://top-thesaurus.org/index)
exact match: Bark thickness [TRY:24] (https://www.try-db.org/de/de.php)
exact match: Bark thickness (inner bark) [TRY:3355] (https://www.try-db.org/de/de.php)
exact match: Bark thickness (outer bark) [TRY:3356] (https://www.try-db.org/de/de.php)
exact match: Average bark thickness [BROT:9] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012011

Scaled bark thickness

Property Value
URI https://w3id.org/APD/traits/trait_0012012
alternative label bark_thickness_index
description A bark morphology trait which is the ratio [PATO:0001470] of two times bark [PO:0004518] thickness [PATO:0000915] to the stem [PO:0009047] diameter [PATO:0001334].
The ratio of two times a single bark thickness measure to stem diameter, to adjust for increasing bark thickness with increasing stem diameter.
comments
value type continuous variable
unit mm/mm
millimetre per millimetre
minAllowedValue 0.001
maxAllowedValue 1
has broader bark morphology trait
plant structure plant structure bark
stem
measured characteristic thickness
index
keyword biomass allocation
scope note
has exact match
has close match
has related match
examples close match: Bark thickness [TOP283] (https://top-thesaurus.org/index)
close match: Bark thickness [TRY:24] (https://www.try-db.org/de/de.php)
close match: Bark thickness (inner bark) [TRY:3355] (https://www.try-db.org/de/de.php)
close match: Bark thickness (outer bark) [TRY:3356] (https://www.try-db.org/de/de.php)
references Lawes 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012012

Bark density

Property Value
URI https://w3id.org/APD/traits/trait_0012013
alternative label bark_density
description The ratio [PATO:0001470] of bark [PO:0004518] dry [PATO:0001824] mass [PATO:0000125] to the corresponding fresh [EnvThes:21976] bark volume [PATO:0000918].
The ratio of bark dry mass to bark fresh volume.
comments
value type continuous variable
unit mg/mm3
milligram per cubic millimetre
minAllowedValue 0.1
maxAllowedValue 1.1
has broader bark morphology trait
plant structure bark
measured characteristic mass
volume density
ratio; proportion
mass density, density
keyword biomass allocation
scope note
has exact match
has close match
has related match
examples exact match: Bark density [TOP282] (https://top-thesaurus.org/index)
exact match: Bark density (bark dry mass per bark volume) [TRY:248] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012013

Bark dry mass per unit bark surface area

Property Value
URI https://w3id.org/APD/traits/trait_0012014
alternative label bark_dry_mass_per_surface_area
description The ratio [PATO:0001470] of bark [PO:0004518] dry [PATO:0001824] mass [PATO:0000125] to the corresponding bark surface area [EnvThes:21564].
The ratio of bark dry mass to bark surface area.
comments
value type continuous variable
unit g/cm2
gram per square centimetre
minAllowedValue 0.01
maxAllowedValue 1
has broader bark morphology trait
plant structure bark
measured characteristic mass
area density
per unit area
ratio; proportion
mass density, density
keyword biomass allocation
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note bark_mass_area
is in scheme https://w3id.org/APD/traits
identifier trait_0012014

Bark water content per unit bark dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0012015
alternative label bark_water_content_per_dry_mass
description The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in bark [PO:0004518] (bark water content [APD:glossary/40014]) to bark dry [PATO:0001824] mass.
The ratio of the mass of water in bark (bark water content) to bark dry mass.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.01
maxAllowedValue 1
has broader bark morphology trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword water
bark water content
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012015

Bark water content per unit saturated bark mass

Property Value
URI https://w3id.org/APD/traits/trait_0012016
alternative label bark_water_content_per_saturated_mass
description A bark mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in bark [PO:0004518] to the fresh mass [APD:glossary/40044] of bark that is at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620].
The ratio of the mass of water in bark (bark water content) to total bark mass, measured in saturated bark. Saturated bark is bark that has been hydrated to its maximal water holding capacity and is at full turgor.
comments
value type continuous variable
unit g/g
gram per gram
minAllowedValue 0.1
maxAllowedValue 10
has broader bark morphology trait
plant structure bark
measured characteristic mass
mass fraction
ratio; proportion
keyword water
water holding capacity
turgor
fresh mass
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012016

Root diameter

Property Value
URI https://w3id.org/APD/traits/trait_0012111
alternative label root_diameter
description A root morphology trait [TO:0000043] which is the maximum [NCIT:C25564] root [PO:0009005] diameter [PATO:0001334] of a plant.
Maximum root diameter.
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 1000
has broader root morphology trait
plant structure root
measured characteristic diameter
size
keyword root morphology trait
scope note
has exact match root width
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012111

Root system morphology

Property Value
URI https://w3id.org/APD/traits/trait_0012112
alternative label root_system_classification
description A root system morphology trait [TO:0000985] which are categorical [STATO:0000252] trait values that capture the branchiness [PATO:0002009] and texture [PATO:0000150] of the root system [PO:0025025] and the root system depth [TO:0001058].
Categorical root system descriptions that indicate the branchiness and texture of the root system and the depth of the root system.
comments The trait values listed are those specified in the root system classification scheme proposed by Cannon in 1949.
value type categorical variable
has narrower root_system_classification_type_i The primary root is fibrous or fleshy and is relatively long; mesophytic.
root_system_classification_type_ii The primary root is relatively short and is fibrous. Roots are well branched in a dry habitat but branch little in one that is wet.
root_system_classification_type_iii The primary root is usually long and fibrous. The first order lateral roots are well branched and are usually short; they are rather evenly distributed along the primary root; xerophytic.
root_system_classification_type_iv The primary root is usually long and slender like that of Type II. The first order lateral roots are well branched. Those closest to the surface of the ground, or not far below, are relatively long; xerophytic and mesophytic.
root_system_classification_type_v The primary root is divided-forked or branched-and fibrous, and with few first order lateral roots, or, in age, apparently with none; xerophytic.
root_system_classification_type_vi The primary root is usually as in Type V. The first order lateral roots are mostly on the upper portion of the primary root and are thus relatively superficial; xerophytic or in well aerated sandy soil where the rainfall is not light.
root_system_classification_type_vii Adventitious roots that are in one group and arise on a short horizontal, or vertical, axis of the shoot. They are fibrous or fleshly; mesophytic and hydrophytic.
root_system_classification_type_viii Adventitious roots that are in one group and are fibrous and fleshy, or fibrous and thick. The fibrous adventitious roots are absorbing-anchoring roots and are usually with laterals to the second order; meophytic.
root_system_classification_type_ix Adventitious roots that arise on the aerial stem and branches of some herbaceous taxa, shrubs, and trees; on creepers, vines, and runners; and on subterranean rhizomes and stolons.
has broader root morphology trait
plant structure root
measured characteristic volume
3-D extent
size
texture
spatial pattern, distribution
keyword root system
root system depth
root branchiness
root texture
scope note
has exact match
has close match
has related match
examples
references Cannon 1949
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note root_morphology
is in scheme https://w3id.org/APD/traits
identifier trait_0012112

Fine root volume to coarse root volume ratio

Property Value
URI https://w3id.org/APD/traits/trait_0012113
alternative label root_fine_root_coarse_root_ratio
description A root system morphology trait [TO:0000985] which is the ratio [PATO:0001470] of the volume [PATO:0000918] of fine roots [EnvThes:20581] (those <0.5mm in diameter) to the volume of coarse roots [APD:glossary/40026] (those >0.5 mm in diameter).
The ratio of the volume of fine roots (<0.5mm diameter) to the volume of coarse roots (>0.5mm diameter).
comments
value type continuous variable
unit mm3/mm3
cubic millimetre per cubic millimetre
minAllowedValue 0.05
maxAllowedValue 50
has broader root morphology trait
plant structure root
measured characteristic volume
ratio; proportion
keyword fine root
root branchiness
coarse root
scope note
has exact match
has close match
has related match
examples close match: Fine root dry mass fraction that is taproot (taproot dry mass per fine root dry mass) [TRY:2011] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012113

Root biomass depth distribution coefficient

Property Value
URI https://w3id.org/APD/traits/trait_0012114
alternative label root_distribution_coefficient
description A root system morphology trait [TO:0000985] which is a model parameter [STATO:0000034] indicating the distribution [PATO:0000060] of a plant’s roots [PO:0009005] with depth [PATO:0001595]. To calculate the model parameter, the required input is a plot of the cumulative root distribution from the surface, where [Y] is the cumulative fraction of roots from the surface to a depth [d]. An asymptotic nonlinear model is used to describe the vertical root distribution: Y = 1 - b^d where Y is the cumulative root fraction from the surface to soil depth d in centimeters and b the estimated parameter.
Root biomass depth distribution coefficient (‘B’ from Gale & Grigal 1987), where high values indicate that root biomass is allocated deeper in the soil. To calculate the model parameter, the required input is a plot of the cumulative root distribution from the surface, where [Y] is the cumulative fraction of roots from the surface to a depth [d]. An asymptotic nonlinear model is used to describe the vertical root distribution: Y = 1 - b^d where Y is the cumulative root fraction from the surface to soil depth d in centimeters and b the estimated parameter.
comments
value type continuous variable
unit {dimensionless}
minAllowedValue 0.5
maxAllowedValue 1
has broader root morphology trait
plant structure root
measured characteristic mass
model parameter
spatial pattern, distribution
keyword root branchiness
root system depth
root system
scope note
has exact match
has close match
has related match
examples close match: Root evenness of dry mass vertical distribution [TRY:3033] (https://www.try-db.org/de/de.php)
references Gale and Grigal 1987
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012114

Root system type (presence of taproot)

Property Value
URI https://w3id.org/APD/traits/trait_0012115
alternative label root_system_type
description A root system morphology trait [TO:0000985] which indicates if the root system [PO:0025025] is one where the primary root [PO:0020127] remains the dominant root, developing into a taproot, a long [PATO:0000573], thick [PATO:0000591] root with few branches, or whether the lateral roots [PO:0020121] are of similar length to the primary root.
Variable describing whether a plant has a tap root.
comments
value type categorical variable
has narrower root_system_type_tap_root_absent Tap root not present.
root_system_type_tap_root_present Tap root present.
root_system_type_tap_root_sometimes_present Tap root sometimes present.
has broader root morphology trait
plant structure root
measured characteristic structure
spatial pattern, distribution
keyword root system
primary root
lateral root
root branchiness
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note tap_root; root_tap_root_status
is in scheme https://w3id.org/APD/traits
identifier trait_0012115

Specific root length (SRL)

Property Value
URI https://w3id.org/APD/traits/trait_0012116
alternative label root_specific_root_length
description A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of root [PO:0009005] length [PATO:0000122] to root dry mass [TO:0000078].
The ratio of root length to root dry mass; SRL.
comments
value type continuous variable
unit m/g
metre per gram
minAllowedValue 0.5
maxAllowedValue 5000
has broader root morphology trait
plant structure root
measured characteristic length
ratio; proportion
mass
keyword biomass allocation
root system
primary root
lateral root
scope note
has exact match
has close match
has related match
examples exact match: Specific root length trait [TOP935] (https://top-thesaurus.org/index)
exact match: Root length per root dry mass (specific root length, SRL) [TRY:1080] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note specific_root_length
is in scheme https://w3id.org/APD/traits
identifier trait_0012116

Specific tap root length (STRL)

Property Value
URI https://w3id.org/APD/traits/trait_0012117
alternative label root_specific_taproot_length
description A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of primary root [PO:0020127] length [PATO:0000122] to primary root dry mass [TO:0000078]
The ratio of taproot length (primary root length) to taproot dry mass; STRL.
comments This trait is usually measured during the very early stage of growth, capturing how efficiently seedlings reach reliable water.
value type continuous variable
unit m/g
metre per gram
minAllowedValue 0.5
maxAllowedValue 5000
has broader root morphology trait
plant structure root
measured characteristic length
ratio; proportion
mass
keyword biomass allocation
root system
primary root
scope note
has exact match
has close match
has related match
examples close match: Coarse root length per coarse root dry mass (specific coarse root length, SRL) [TRY:1550] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note specific_taproot_length
is in scheme https://w3id.org/APD/traits
identifier trait_0012117

Root surface area per unit root dry mass (specific root area)

Property Value
URI https://w3id.org/APD/traits/trait_0012118
alternative label root_specific_root_area
description A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of root [PO:0009005] surface area [EnvThes:21564] to root dry mass [TO:0000078].
The ratio of root surface area to root dry mass; specific root area.
comments
value type continuous variable
unit mm2/mg
square millimetre per milligram
minAllowedValue 0.001
maxAllowedValue 1
has broader root morphology trait
plant structure root
measured characteristic area
size
ratio; proportion
area density
mass density, density
keyword biomass allocation
root system
scope note
has exact match
has close match
has related match
examples exact match: Root surface area per root dry mass (specific root area) [TRY:1089] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note specific_root_area
is in scheme https://w3id.org/APD/traits
identifier trait_0012118

Root wood density

Property Value
URI https://w3id.org/APD/traits/trait_0012119
alternative label root_wood_density
description A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of the dry mass [PATO:0000125] of a unit of wood [PO_0005848] from a root [PO:0009005]to its fresh fresh [EnvThes:21976] volume [PATO:0000918].
The ratio of woody root dry mass to woody root fresh volume.
comments
value type continuous variable
unit mg/mm3
milligram per cubic millimetre
minAllowedValue 0.05
maxAllowedValue 10
has broader root morphology trait
plant structure root
measured characteristic mass
volume density
ratio; proportion
mass density, density
keyword biomass allocation
root system
wood, secondary xylem
scope note
has exact match
has close match
has related match
examples exact match: Root tissue density (root dry mass per root volume) [TRY:82] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012119

Root to shoot ratio

Property Value
URI https://w3id.org/APD/traits/trait_0012120
alternative label root_shoot_ratio
description A whole plant morphology trait [TO:0000398] which is the ratio [PATO:0001470] of a plant’s root dry mass [TO:0000078] to its shoot dry mass [TO:0000552].
The ratio of total root system dry mass to total shoot system dry mass.
comments This trait is usually measured on experimentally grown seedlings where it is possible to collect the entire root system.
value type continuous variable
unit mg{root}/mg{shoot}
milligram per milligram
minAllowedValue 0.01
maxAllowedValue 15
has broader root morphology trait
stem morphology trait
plant structure plant structure plant structure root
whole plant
stem
measured characteristic mass
ratio; proportion
ratio; proportion
mass fraction
mass density, density
keyword biomass allocation
root system
scope note
has exact match root to shoot ratio
has close match
has related match
examples exact match: Root/shoot ratio [TRY:9] (https://www.try-db.org/de/de.php)
exact match: Shoot:root ratio [BROT:22] (http://doi.org/10.1038/sdata.2018.135)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012120

Root dry matter content (RDMC)

Property Value
URI https://w3id.org/APD/traits/trait_0012121
alternative label root_dry_matter_content
description A root mass trait [TO:0000279] which is the ratio [PATO:0001470] of root dry mass [TO:0000078] to root [PO:0009005] fresh mass [APD:glossary/40044].
The ratio of root dry mass to root fresh mass; root dry matter content, RDMC.
comments
value type continuous variable
unit mg/g
milligram per gram
minAllowedValue 10
maxAllowedValue 1000
has broader root morphology trait
plant structure root
measured characteristic mass
mass fraction
ratio; proportion
keyword biomass allocation
root system
fresh mass
scope note
has exact match
has close match
has related match
examples exact match: Root dry mass per root fresh mass (root dry matter content, RDMC) [TRY:785] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012121

Root mass fraction

Property Value
URI https://w3id.org/APD/traits/trait_0012122
alternative label root_mass_fraction
description A root mass trait [TO:0000279] which is the ratio [PATO:0001470] of the total root dry mass [TO:0000078] to whole plant [PO:0000003] dry mass [APD:glossary/40080].
The ratio of total root system dry mass to whole plant dry mass.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.01
maxAllowedValue 1
has broader root morphology trait
plant structure plant structure root
whole plant
measured characteristic mass
mass fraction
ratio; proportion
keyword biomass allocation
whole plant dry mass
root system
scope note
has exact match
has close match
has related match
examples exact match: Plant biomass and allometry: Root mass fraction (fraction of root dry mass per whole plant dry mass) [TRY:470] (https://www.try-db.org/de/de.php)
related match: root dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012122

Root vascular anatomy

Property Value
URI https://w3id.org/APD/traits/trait_0012123
alternative label root_vascular_anatomy
description A root system morphology trait [TO:0000985] which is the arrangement [NCIT:C61532] or type [NCIT:C25284] of the root [PO:0020127] vascular system [PO:0003011].
The arrangement or type of the root vascular system.
comments
value type categorical variable
has narrower root_vascular_anatomy_cord_roots A root anatomy found within three genera of Mirbelieae (Daviesia, Euchilopsis, Phyllota) wherein networks of interlinking vascular strands exist (Crisp 2003; Pate and Dixon 1996).
root_vascular_anatomy_unistelar_roots The commonly observed root anatomy, wherein there is a single vascular strand which displays secondary thickening.
has broader root morphology trait
plant structure root
measured characteristic volume
3-D extent
size
texture
spatial pattern, distribution
keyword root system
root vascular cambium
scope note
has exact match
has close match
has related match
examples
references Cannon 1949
date created 03/06/2025
date modified 03/06/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012123

Storage organ length

Property Value
URI https://w3id.org/APD/traits/trait_0012151
alternative label storage_organ_length
description A plant structure morphology trait [TO:0000839] which is the length [PATO:0000122] of a plant’s storage organ [EFO:0000997], where a storage organ is a plant part (organ [PO:0009008], tissue [PO:0009007], or other plant structure [PO:0009011]) whose function [NCIT:C25518] is carbohydrate [CHEBI:16646] (energy [PATO:0001021]) storage.
The length of an underground plant structure (tissues, organs, or other structures) where a plant stores excess carbohydrates (energy).
comments This can be the length of any of the structures documented in the trait storage_organ and should be documented as an entity context.
value type continuous variable
unit mm
millimetre
minAllowedValue 1
maxAllowedValue 1000
has broader stem morphology trait
root morphology trait
plant structure plant structure root
stem
measured characteristic length
size
keyword storage organ
energy
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012151

Storage organ diameter

Property Value
URI https://w3id.org/APD/traits/trait_0012152
alternative label storage_organ_diameter
description A plant structure morphology trait [TO:0000839] which is the diameter [PATO:0001334] of a plant’s storage organ [EFO:0000997], where a storage organ is a plant part (organ [PO:0009008], tissue [PO:0009007], or other plant structure [PO:0009011]) whose function [NCIT:C25518] is carbohydrate [CHEBI:16646] (energy [PATO:0001021]) storage.
The width of an underground plant structure (tissues, organs, or other structures) where a plant stores excess carbohydrates (energy).
comments This can be the length of any of the structures documented in the trait storage_organ and should be documented as an entity context.
value type continuous variable
unit mm
millimetre
minAllowedValue 1
maxAllowedValue 1000
has broader stem morphology trait
root morphology trait
plant structure plant structure root
stem
measured characteristic diameter
size
keyword storage organ
energy
scope note
has exact match
has close match tuberous root tuber diameter
has related match
examples exact match: Belowground plant organ diameter [TRY:2550] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012152

Seed accessory cost fraction

Property Value
URI https://w3id.org/APD/traits/trait_0012221
alternative label accessory_cost_fraction
description A reproductive shoot system [PO:0025082] biomass allocation [EnvThes:21360] trait which is one minus the ratio [PATO:0001470] of total plant seed [PO:0009010] dry [PATO:0001824] mass [PATO:0000125] to total plant reproductive shoot system tissue dry mass where reproductive tissues include all flower buds [PO:0000056], flowers [PO:0009046], fruits [PO:0009001], dispersal [EnvThes:21038] tissues, and seeds produced during the developmental process involved in reproduction [GO:0003006].
The fraction of total reproductive investment required to mature a seed that is invested in non-seed tissues. It is calculated as one minus, the ratio of total biomass investment in seeds to total biomass investment in all reproductive tissues, including flower buds, flowers, fruits, dispersal tissues, aborted seeds, and successfully matured seeds.
comments
value type continuous variable
unit mg/mg
milligram per milligram
minAllowedValue 0.01
maxAllowedValue 1
has broader reproductive shoot system morphology trait
plant structure plant structure plant structure plant structure reproductive shoot system
flower
fruit
seed
measured characteristic mass
keyword biomass allocation
reproduction
accessory cost
scope note
has exact match
has close match
has related match
examples close match: Plant biomass and allometry: Reproductive dry mass per plant dry mass (reproductive weight fraction) [TRY:1046] (https://www.try-db.org/de/de.php)
references Lord & Westoby 2006
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012221

Seed accessory cost mass

Property Value
URI https://w3id.org/APD/traits/trait_0012222
alternative label accessory_cost_mass
description A reproductive shoot system [PO:0025082] biomass allocation [EnvThes:21360] trait which is the sum of the dry [PATO:0001824] mass [PATO:0000125] of all flower buds [PO:0000055], flowers [PO:0009046], fruits [PO:0009001], aborted seeds, and dispersal [EnvThes:21038] tissues produced during the developmental process involved in reproduction [GO:0003006], but excludes mature [PATO:0001701] seed [PO:0009010] dry mass.
The mass of seed accessory costs, which is the total biomass investment in all reproductive tissues, excluding biomass invested in successfully matured seeds; it includes all biomass invested in flower buds, flowers, fruits, dispersal tissues and aborted seeds.
comments
value type continuous variable
unit mg
milligram
minAllowedValue 0.01
maxAllowedValue 10000
has broader reproductive shoot system morphology trait
plant structure plant structure plant structure plant structure reproductive shoot system
flower
fruit
seed
measured characteristic mass
keyword biomass allocation
reproduction
accessory cost
scope note
has exact match
has close match
has related match
examples
references Lord & Westoby 2006
date created 14/07/2021
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012222

Bud length

Property Value
URI https://w3id.org/APD/traits/trait_0012301
alternative label bud_length
description A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of the bud [PO:0000055] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the bud.
Linear dimension from the base to the apex of a bud, even if this is not the longest dimension.
comments
value type continuous variable
unit cm
minAllowedValue 0.01
maxAllowedValue 200
has broader floral organ morphology trait
plant structure plant structure reproductive shoot system
bud
measured characteristic length
size
scope note
has exact match reproduction
corolla
perianth
has close match
has related match
examples
references no linked references
date created 05/02/2024
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012301

Bud width

Property Value
URI https://w3id.org/APD/traits/trait_0012302
alternative label bud_width
description A floral organ morphology trait [TO:0000862] which is the width [PATO:0000921] of the bud [PO:0000055] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the bud. Width is the greatest linear dimension [PATO:0001708] perpendicular to [PATO:0002434] to the bud length [PATO:0000122]
Greatest linear width dimension of a bud, sometimes referred to as ‘diameter’ in buds where the width and height are identical; orthogonal to the bud length, the base-to-apex dimension.
comments
value type continuous variable
unit cm
minAllowedValue 0.01
maxAllowedValue 200
has broader floral organ morphology trait
plant structure plant structure reproductive shoot system
bud
measured characteristic width
size
scope note
has exact match reproduction
corolla
perianth
has close match
has related match
examples
references no linked references
date created 05/02/2024
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012302

Buds per inflorescence

Property Value
URI https://w3id.org/APD/traits/trait_0012351
alternative label buds_per_inflorescence
description A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of buds [PO:0000055] in an inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers or the entire reproductive shoot system [PO:0025082].
The count of buds in an inflorescence, where an inflorescence can be either a single cluster of flowers or the entire reproductive shoot system.
comments
value type continuous variable
unit {count}/{count}
minAllowedValue 1
maxAllowedValue 1000
has broader floral organ morphology trait
plant structure plant structure plant structure reproductive shoot system
bud
inflorescence
measured characteristic has number of
dimensionless ratio
ratio; proportion
count
tolerance
sensitivity
scope note
has exact match reproduction
reproductive shoot system
inflorescence
has close match
has related match
examples
references no linked references
date created 05/02/2024
date modified 05/02/2024
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012351

Inflorescence length

Property Value
URI https://w3id.org/APD/traits/trait_0012352
alternative label inflorescence_length
description A floral organ morphology trait [TO:0000862] which is the vertical [PATO:0001854] length [PATO:0000122] (height [PATO:0000119]) of the inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers [PO:0009046] or the entire reproductive shoot system [PO:0025082].
The vertical length (height) of an inflorescence.
comments
value type continuous variable
unit cm
minAllowedValue 0.01
maxAllowedValue 1500
has broader perianth morphology trait
plant structure plant structure plant structure reproductive shoot system
bud
inflorescence
measured characteristic length
size
keyword floral organ morphology trait
flower
reproductive shoot system
inflorescence morphology trait
scope note
has exact match inflorescence length
has close match
has related match
examples exact match: Inflorescence length (height) [TRY:2817] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012352

Inflorescence diameter

Property Value
URI https://w3id.org/APD/traits/trait_0012353
alternative label inflorescence_diameter
description A floral organ morphology trait [TO:0000862] which is the diameter [PATO:0001334] of the inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers [PO:0009046] or the entire reproductive shoot system [PO:0025082].
The diameter (of greatest width dimension) of an inflorescence.
comments
value type continuous variable
unit cm
minAllowedValue 0.01
maxAllowedValue 1500
has broader perianth morphology trait
plant structure plant structure plant structure reproductive shoot system
bud
inflorescence
measured characteristic diameter
size
keyword floral organ morphology trait
flower
reproductive shoot system
inflorescence morphology trait
scope note
has exact match inflorescence diameter
has close match
has related match
examples exact match: Inflorescence diameter [TRY:3587] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012353

Number of androecium parts in each whorl (Androecium structural merism)

Property Value
URI https://w3id.org/APD/traits/trait_0012410
alternative label flower_androecium_structural_merism
description An androecium morphology trait [TO:0000994] which is the total count [NCIT:C25463] of stamens [PO:0009029], staminodes [PO:0009077] or stamen bundles (fascicles [NCIT:C79692]) in each floral whorl [PO:0025023].
The number of stamens or stamen bundles (fascicles) in one whorl, considering both fertile stamens and staminodes.
comments This character is not applicable for spiral or irregular stamen arrangements, nor when there is a single structural stamen (i.e., one stamen, no staminodes; e.g., Chloranthus, Chloranthaceae). Definition derived from Schoenenberger et al. (2020) for their character ‘332. Androecium structural merism (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 100
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
androecium
stamen
staminode
fascicle
merism
whorl, floral whorl
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012410

Androecium structural phyllotaxis

Property Value
URI https://w3id.org/APD/traits/trait_0012411
alternative label flower_androecium_structural_phyllotaxis
description An androecium morphology trait [TO:0000994] which is the spatial pattern or distribution [PATO:0000060] (phyllotaxy [TO:0006014]) of stamen [PO:0009029], staminodes [PO:0009077] or stamen bundles (fascicles [NCIT:C79692]) in each floral whorl [PO:0025023].
Structural phyllotaxy of the androecium, considering both fertile stamens and staminodes. In cases of stamen fascicles, it is the phyllotaxis of fascicles, not individual stamens that are recorded.
comments This character is not applicable when there is a single structural stamen (i.e., one stamen, no staminodes; e.g., Chloranthus, Chloranthaceae). Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘330. Androecium structural phyllotaxy (D1)’
value type categorical variable
has narrower flower_androecium_structural_phyllotaxis_whorled Stamens and staminodes arranged in whorls.
flower_androecium_structural_phyllotaxis_spiral Stamens and staminodes have a spiral arrangement.
flower_androecium_structural_phyllotaxis_irregular Stamens and staminodes have an irregular arrangement.
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
keyword reproduction
androecium
phyllotaxy
stamen
staminode
fascicle
whorl, floral whorl
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012411

Number of androecium structural whorls

Property Value
URI https://w3id.org/APD/traits/trait_0012412
alternative label flower_androecium_structural_whorls_count
description An androecium morphology trait [TO:0000994] which is the total count [NCIT:C25463] of whorls [PO:0025023] comprising the androecium [PO:0009061], considering both stamen [PO:0009029] and staminodes [PO:0009077]. If stamens are grouped into fascicles [NCIT:C79692], the trait is the count of whorls of fascicles.
The number of whorls, considering both fertile stamens and staminodes. In cases of stamen fascicles, the count of whorls of fascicles is recorded.
comments This character is not applicable for spiral or irregular stamen arrangements, or when there is a single structural stamen (i.e., one stamen, no staminodes; e.g., Chloranthus, Chloranthaceae). Definition derived from Schoenenberger et al. (2020) for their character ‘331. Number of androecium structural whorls (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 100
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
androecium
stamen
staminode
fascicle
merism
whorl, floral whorl
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012412

Anther attachment

Property Value
URI https://w3id.org/APD/traits/trait_0012413
alternative label flower_anther_attachment
description An androecium morphology trait [TO:0000994] which indicates how the filament [PO:0009067] is attached to [PATO:0001667] the tissue [CARO:0000043] connecting the two thecae [PO:0009069] (pollen [PO:0025281] sacs) of the anthers [PO:0009066], affecting the spatial pattern [PATO:0000060] of the stamens [PO:0009029].
Anther attachment refers to the area of insertion of the filament on the anther connective (i.e. the tissue connecting the two thecae of an anther).
comments Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘312. Anther attachment (D1)’
value type categorical variable
has narrower flower_anther_attachment_basifixed Filament attached to the base of the connective. Laminar stamens are classified as basifixed.
flower_anther_attachment_dorsifixed Filament attached to the dorsal side of the anther.
flower_anther_attachment_dorsifixed_at_base Filament attached to the dorsal side of the anther, at the base.
flower_anther_attachment_ventrifixed Filament attached to the ventral side of the anther.
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
keyword reproduction
androecium
stamen
anther
pollen
connective
anther theca
filament
scope note
has exact match
has close match
has related match anther attachment quality
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012413

Connective extension (apical)

Property Value
URI https://w3id.org/APD/traits/trait_0012414
alternative label flower_anther_connective_extension
description An androecium morphology trait [TO:0000994] which is whether anthers [PO:0009066] have protrusions [NCIT:C50710] that extend above the two theca [PO:0009069] (pollen [PO:0025281] sacs), and, if present, the shape [PATO:0000052] and the relative length [PATO:0000122] of the protrusion relative to the length of the entire anther.
Apical (distal) connective extensions (also called ‘distal connective protrusions’) are sterile anther structures that distally extend beyond the level of the thecae (i.e., the two lateral pairs of pollen sacs of a tetrasporangiate anther).
comments This trait records both absence or presence of anther connective extensions and also the shape of these extensions in terms of length in relation to the length of the thecae. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘314. Connective extension (apical) (D1)’
value type categorical variable
has narrower flower_anther_connective_extension_absent Apical connective extensions are lacking.
flower_anther_connective_extension_short_extension Connective extension less than a third of the length of anther.
flower_anther_connective_extension_long_extension Connective extension more than a third but less than length of anther; e.g. Idiospermum australiense.
flower_anther_connective_extension_very_long_extension Connective extension more than length of anther; e.g. Galbulimima belgraveana.
flower_anther_connective_extension_present_general Apical connective extensions are present but no information on their length relative to the anthers.
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic structure
shape
length
keyword reproduction
androecium
stamen
anther
pollen
connective
anther theca
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012414

Anther dehiscence

Property Value
URI https://w3id.org/APD/traits/trait_0012415
alternative label flower_anther_dehiscence
description An androecium morphology trait [TO:0000994] which is the length [PATO:0000122] and shape [PATO:0000052] of the stomia [PO:0020101] that form the anther dehiscence zone [PO:0005011]; stomia are the usually slit-like [PATO:0002482] openings in the anther [PO:0009066] through which pollen [PO:0025281] is released.
Anther dehiscence refers to the type of opening of the anther when releasing its pollen through the stomia.
comments Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘313. Anther dehiscence (D1)’
value type categorical variable
has narrower flower_anther_dehiscence_longitudinal_slit Anther with longitudinal slits that extend along the entire length of each theca.
flower_anther_dehiscence_poricidal Anther where dehiscence of longitudinal slits is incomplete, with pollen exiting via a small pore at the tip.
flower_anther_dehiscence_pores_many Anther with pollen exiting from many pores along its length.
flower_anther_dehiscence_h_valvate Anther with two valves on longitudinal hinges, opening horizontally as salon doors; when the stomium bifurcates at its distal and/or proximal end and thus a valve is formed.
flower_anther_dehiscence_flap_valvate Anther with one or more flap-like valves on horizontal hinges, opening vertically.
flower_anther_dehiscence_transverse_slit Anther with horizontal slit.
flower_anther_dehiscence_short_basal_slits Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the base of the theca.
flower_anther_dehiscence_short_central_slits Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the center of the theca.
flower_anther_dehiscence_short_apical_slits Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the apical end of the theca.
flower_anther_dehiscence_t_valvate Anther with T-shaped stomium; reduced transverse slit at base.
flower_anther_dehiscence_common_stomium_confluent_thecae The two thecae of a single anther are fused at one end with a single pore through which pollen exits.
flower_anther_dehiscence_circular Anther with a transverse slit that goes entirely around the anther.
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic structure
shape
length
keyword reproduction
androecium
stamen
anther
pollen
stomium
anther dehiscence zone
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012415

Anther orientation

Property Value
URI https://w3id.org/APD/traits/trait_0012416
alternative label flower_anther_orientation
description An androecium morphology trait [TO:0000994] which is the position [PATO:0000140] and orientation [PATO:0000133] of the stomia [PO:0020101] that form the anther dehiscence zone [PO:0005011]; stomia are the slit-like [PATO:0002482] openings in the anther [PO:0009066] through which pollen [PO:0025281] is released.
Orientation of anther at anthesis.
comments Anthers of angiosperms are rather uniform in their basic structure. They normally have four microsporangia (pollen sacs) that are arranged pair-wise in two thecae. The two microsporangia of a theca usually release their pollen grains through a common opening (stomium). Often, it is difficult to establish anther orientation clearly in a flower as this is a gradual feature with many intermediate stages. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘311. Anther orientation (D1)’
value type categorical variable
has narrower flower_anther_orientation_apical When thecae are positioned in a transverse position at the tip of the connective and thus dehisce upward in the flower (e.g., Sinofranchetia, Endress and Hufford, 1989).
flower_anther_orientation_extrorse When the stomia of the thecae face the floral periphery.
flower_anther_orientation_introrse When the stomia of the thecae face the floral centre.
flower_anther_orientation_latrorse When pollen is released toward the side (i.e., toward neighbouring anthers).
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
orientation
position
keyword reproduction
androecium
stamen
anther
pollen
stomium
anther dehiscence zone
scope note
has exact match
has close match anther orientation
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012416

Flower colour

Property Value
URI https://w3id.org/APD/traits/trait_0012417
alternative label flower_colour
description A floral organ morphology trait [TO:0000862] which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the flower [PO:0009046. The petals [PO:0009032] (corolla [PO:0009059]), sepals [PO:0009031] (calyx [PO:0009060]), or the entire perianth [PO:0009058] may be responsible for the main colour observed.
The dominant flower colour, where the entire flower, the petals, or the sepals may be responsible for the observed colour.
comments Most data mapped to flower_colour has probably been scored based on perianth colour, but if a dataset does not differentiate between the two traits it is mapped to flower_colour.
value type categorical variable
has narrower flower_colour_blue_purple Flower colour is blue or purple.
flower_colour_green Flower colour is green.
flower_colour_pink Flower colour is pink.
flower_colour_red_brown Flower colour is red or brown.
flower_colour_white_cream Flower colour is white or cream.
flower_colour_yellow_orange Flower colour is yellow or orange.
flower_colour_grey Flower colour is grey.
flower_colour_black Flower colour is black.
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic colour / color
keyword reproduction
petal
corolla
sepal
calyx
perianth
scope note
has exact match flower color
has close match corolla color
has related match
examples exact match: Flower color [TOP242] (https://top-thesaurus.org/index)
exact match: Flower color [TRY:207] (https://www.try-db.org/de/de.php)
exact match: Flower_colour [GIFT:3.21.1] (https://gift.uni-goettingen.de)
exact match: flower color [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012417

Perianth colour

Property Value
URI https://w3id.org/APD/traits/trait_0012418
alternative label perianth_colour
description A floral organ morphology trait [TO:0000862] which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]).
The dominant colour of the perianth parts (sepals, petals and tepals).
comments This trait refers explicitly to the colour of the petals, sepals and tepals, while the trait ‘flower colour’ refers to the colour of the entire flower. See also data under flower_colour. Most data mapped to flower_colour has probably been scored based on perianth colour, but if a dataset does not differentiate between the two traits it is mapped to flower_colour.
value type categorical variable
has narrower perianth_colour_white Perianth colour is white.
perianth_colour_yellow Perianth colour is yellow.
perianth_colour_orange Perianth colour is orange.
perianth_colour_red Perianth colour is red.
perianth_colour_pink Perianth colour is pink.
perianth_colour_purple Perianth colour is purple.
perianth_colour_blue Perianth colour is blue.
perianth_colour_green Perianth colour is green.
perianth_colour_grey Perianth colour is grey.
perianth_colour_brown Perianth colour is brown.
perianth_colour_black Perianth colour is black.
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic colour / color
keyword reproduction
petal
corolla
sepal
calyx
perianth
scope note
has exact match
has close match corolla color
has related match
examples exact match: Flower color [TOP242] (https://top-thesaurus.org/index)
exact match: Flower color [TRY:207] (https://www.try-db.org/de/de.php)
exact match: Flower_colour [GIFT:3.21.1] (https://gift.uni-goettingen.de)
exact match: flower color [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references no linked references
date created 19/05/2023
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012418

Flower length

Property Value
URI https://w3id.org/APD/traits/trait_0012419
alternative label flower_length
description A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of the flower [PO:0009046] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the flower, measured at anthesis [PO:0009068]. It excludes the length of the pedicel [PO:0030112].
The length of the flower.
comments
value type continuous variable
unit cm
minAllowedValue 0.01
maxAllowedValue 200
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic length
keyword reproduction
petal
corolla
sepal
calyx
perianth
scope note
has exact match flower length
has close match
has related match
examples
references no linked references
date created 19/05/2023
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012419

Flower diameter

Property Value
URI https://w3id.org/APD/traits/trait_0012420
alternative label flower_diameter
description A floral organ morphology trait [TO:0000862] which is the maximum diameter [PATO:0001334] of the flower [PO:0009046] at anthesis [PO:0009068], considering all floral organs, but most commonly the diameter from petal [PO:0009032] tip to petal tip.
The maximum diameter of the flower at anthesis, considering all floral organs, but most commonly the diameter from petal tip to petal tip.
comments
value type continuous variable
unit cm
minAllowedValue 0.01
maxAllowedValue 200
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic diameter
keyword reproduction
petal
corolla
sepal
calyx
perianth
scope note
has exact match flower diameter
has close match
has related match
examples
references no linked references
date created 19/05/2023
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012420

Floral orientation

Property Value
URI https://w3id.org/APD/traits/trait_0012421
alternative label flower_orientation
description A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] and orientation [PATO:0000133] of the flower [PO:0009046].
The position and orientation of the flower.
comments
value type categorical variable
has narrower flower_orientation_up Flower orientation is vertical, with the floral entrance pointing up.
flower_orientation_lateral Flower orientation is horizontal, with the floral entrance perpendicular to main vertical axis.
flower_orientation_down Flower orientation is pendant, with the floral entrance pointing down.
flower_orientation_mixed Flower orientation is mixed, with different flowers exhibiting different orientations.
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
orientation
position
keyword reproduction
petal
corolla
sepal
calyx
perianth
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 19/05/2023
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012421

Maximum flower number

Property Value
URI https://w3id.org/APD/traits/trait_0012422
alternative label flower_count_maximum
description A floral organ morphology trait [TO:0000862] which is the maximum [NCIT:C25564] count [NCIT:C25463] of flowers [PO:0009046] on an individual [EFO:0000542].
Maximum flower number produced.
comments
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 1e+06
has broader floral organ morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
maximum
keyword reproduction
perianth
corolla
scope note
has exact match
has close match
has related match
examples exact match: Flower number per plant [TRY:2999] (https://www.try-db.org/de/de.php)
references no linked references
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012422

Number of fertile stamens

Property Value
URI https://w3id.org/APD/traits/trait_0012431
alternative label flower_fertile_stamens_count
description A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of fertile [PATO:0000955] stamens [PO:0009029] on a hermaphrodite [PATO:0001340] flower [PO:0009046] or male [PATO:0000384] flower.
Count of fertile stamens in bisexual or male flowers; does not score female flowers.
comments Definition derived from Schoenenberger et al. (2020) for their character ‘301. Number of fertile stamens (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 10000
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
androecium
stamen
scope note male or hermaphroditic flowers
has exact match stamen number
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012431

Filament presence and shape

Property Value
URI https://w3id.org/APD/traits/trait_0012432
alternative label flower_filament
description A floral organ morphology trait [TO:0000862] which is whether a filament [PO:0009067] is present [PATO:0000467] or absent [PATO:0000462], and if present, the shape [PATO:0000052] and the relative length [PATO:0000122], width [PATO:0000921], and thickness [PATO:0000915] of the filament.
Trait that considers the absence or presence of the filament, and in the latter case, the shape of the filament. Shape is considered in terms of length and width and is defined in relation to anther length/width.
comments The width of the filament may thus either be broad as in laminar (e.g., Eupomatiaceae) or bulky stamens (e.g., Chloranthaceae), or narrow (filamentous) as found in many core eudicots groups (e.g., Rosaceae). This trait is considered inapplicable when filaments are entirely fused with each other or to the perianth. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘305. Filament (D1)’
value type categorical variable
has narrower flower_filament_absent No filaments present
flower_filament_present_general Filaments present, with no specific defining characteristics.
flower_filament_long_and_narrow Filaments have a long and narrow shape. (Synonyms, filiform, slender)
flower_filament_short_and_narrow Filaments have a short and narrow shape; subsessile anthers.
flower_filament_long_and_wide Filaments have a long and wide laminar shape.
flower_filament_short_and_wide Filaments have a short and wide laminar shape.
flower_filament_long_and_very_wide Filaments have a long and very wide shape; for instance, the petal-like (outer) stamens of Nymphaea.
flower_filament_petaloid Filaments are long and wide and colorful, e.g. Canna.
flower_filament_fused_into_synandrium Filament structure formed when stamens fused together.
flower_filament_fused_into_fascicle_stalks Special case where the stamens are fused into the stalk of a fascicle of flowers.
flower_filament_long_and_narrow_divided_in_two Filaments have a long and narrow shape that is divided in two; e.g. Adoxa.
flower_filament_short_general Filament is short relative to the anther.
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic shape
structure
keyword reproduction
androecium
stamen
filament
anther
scope note inapplicable when filaments are entirely fused with each other or to the perianth
has exact match
has close match filament 2-D shape
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012432

Fusion of filaments

Property Value
URI https://w3id.org/APD/traits/trait_0012433
alternative label flower_filament_fusion
description A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of the stamen [PO:0009029] (or staminode [PO:0009077]) filaments [PO:0009067] that is fused [NCIT:C124537] together at anthesis [PO:0009068].
Fusion of stamen (and staminode) filaments among each other at anthesis (congenitally or postgenitally) is recorded on a continuous scale, from 0 (free filaments) to 1 (filaments fused along their entire length).
comments Partial fusion is recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the length of filaments). Definition derived from Schoenenberger et al. (2020) for their character ‘306. Fusion of filaments (C1)’
value type continuous variable
unit {dimensionless}
minAllowedValue 0
maxAllowedValue 1
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic dimensionless ratio
ratio; proportion
keyword reproduction
androecium
stamen
filament
connective
staminode
scope note
has exact match
has close match
has related match filament attached to
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012433

Fusion of filaments to inner perianth series

Property Value
URI https://w3id.org/APD/traits/trait_0012434
alternative label flower_filament_fusion_to_inner_perianth
description A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of the stamen [PO:0009029] (or staminode [PO:0009077]) filaments [PO:0009067] that are fused [NCIT:C124537] to the innermost perianth [PO:0009058] organ [PO:0009008] at anthesis [PO:0009068].
The fusion of filaments with the innermost perianth organs at anthesis on a continuous scale, from 0 (filaments completely free from perianth) to 1 (filaments fused along their entire length with the perianth; the ‘entire length’ of the filament is defined as the distance between the floral base and the joint between filament and anther).
comments Partial fusion is recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the length of perianth parts). If two (or more) stamen whorls (including staminodial whorls) differ in their extent of filament fusion with the perianth, this trait is recorded as a range of values. For example, if the filaments of an outermost stamen whorl are fused, up to 90% of their length, but an inner whorl only up to 50%, the trait is assigned a range of 0.5 to 0.9. The rationale for this is to provide a general trait that allows comparison of fusion among all angiosperms. Definition derived from Schoenenberger et al. (2020) for their character ‘308. Fusion of filaments to inner perianth series (C1)’
value type continuous variable
unit {dimensionless}
minAllowedValue 0
maxAllowedValue 1
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic dimensionless ratio
ratio; proportion
keyword reproduction
androecium
stamen
filament
connective
staminode
perianth
scope note
has exact match
has close match
has related match filament attached to
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012434

Gynoecium phyllotaxis

Property Value
URI https://w3id.org/APD/traits/trait_0012441
alternative label flower_gynoecium_phyllotaxis
description A gynoecium [PO:0009062] morphology trait which is either the count [NCIT:C25463] of whorls [PO:0025023] of carpels [PO:0009030] comprising the gynoecium or other descriptors of carpel phyllotaxy [TO:0006014].
Structural phyllotaxy of the gynoecium.
comments This trait is not applicable to unicarpellate flowers. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘400. Gynoecium phyllotaxy (D1)’
value type categorical variable
has narrower flower_gynoecium_phyllotaxis_one_whorl Flowers with a single whorl of carpels.
flower_gynoecium_phyllotaxis_two_or_more_whorls Flowers with two or more whorls of carpels.
flower_gynoecium_phyllotaxis_spiral Flowers with carpels in a spiral arrangement.
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
orientation
structure
has number of
count
keyword reproduction
gynoecium
carpel
whorl, floral whorl
phyllotaxy
scope note Not applicable to unicarpellate flowers
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012441

Placentation

Property Value
URI https://w3id.org/APD/traits/trait_0012442
alternative label flower_gynoecium_placentation
description A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] of the ovules [PO:0020003] within the ovary [PO:0009072], known as placentation [APD:glossary/40074].
The different types of placentation in apocarpous/unicarpellate and in syncarpous gynoecia.
comments In apocarpous and unicarpellate gynoecia, placentation is often described as marginal as the ovules are usually attached along the ventral slit (i.e., the zone where the carpel margins become postgenitally closed during carpel development). In syncarpous gynoecia there are three main types of placentation, 1) axile placentation refers to ovaries where the ovules are placed in the angle between carpel flanks in the center of the ovary; 2) in ovaries with parietal placentation the ovules attach to the ovary wall where two carpels meet; 3) in ovaries with free-central placentation the ovules are attached to a central column that emerges from the base of the ovary and protrudes into the non-septate ovary. It is not unusual that syncarpous gynoecia show a transition from proximally axile to a distally parietal placentation (e.g., Polemoniaceae). Basal and apical placentation may occur both in apocarpous/unicarpellate and in syncarpous gynoecia. Usually there is one longitudinal series of ovules attached to each carpel margin. However, both in apocarpous/unicarpellate and in syncarpous gynoecia there may be more than one series of ovules at the flanks of a carpel. These latter cases are referred to as laminar (or laminar-diffuse) placentation (e.g., in some Nymphaeaceae). A placenta that is protruding from its surroundings and has more than two series of ovules (either axile or parietal) is called protruding-diffuse. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘412. Placentation (D1)’
value type categorical variable
has narrower flower_gynoecium_placentation_axile Ovules are placed in the angle between carpel flanks in the center of the ovary; in syncarpous gynoecia.
flower_gynoecium_placentation_apical Ovules attach at the top of the ovary; pendulous; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
flower_gynoecium_placentation_basal Ovules attach at the base of the ovary; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
flower_gynoecium_placentation_parietal Ovules attach to the ovary wall where two carpels meet; in syncarpous gynoecia.
flower_gynoecium_placentation_marginal Ovules attach along the ventral slit; in apocarpous and unicarpellate gynoecia.
flower_gynoecium_placentation_free_central Ovules are attached to a central column that emerges from the base of the ovary and protrudes into the non-septate ovary; in syncarpous gynoecia.
flower_gynoecium_placentation_laminar When there is more than one series of ovules at the flanks of a carpel; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic structure
spatial pattern, distribution
position
keyword reproduction
gynoecium
carpel
plant ovary
placentation
plant ovule
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012442

Fusion of ovaries

Property Value
URI https://w3id.org/APD/traits/trait_0012443
alternative label flower_ovary_fusion
description A floral organ morphology trait [TO:0000862] which, for flowers with multiple carpels, is the proportion [PATO:0001470] of the length [PATO:0000122] of the ovaries [PO:0009072] that are fused [NCIT:C124537] together.
Degree of ovary fusion expressed as a fraction of the total length of the ovary (from the floral base to the apex of the ovary).
comments Fusion of styles and stigmas is not taken into account for this trait. Not applicable when there is a single carpel. Definition derived from Schoenenberger et al. (2020) for their character ‘403. Fusion of ovaries (C1)’
value type continuous variable
unit {dimensionless}
minAllowedValue 0
maxAllowedValue 1
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic dimensionless ratio
ratio; proportion
structure
keyword reproduction
gynoecium
carpel
plant ovary
scope note Not applicable when there is a single carpel.
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012443

Ovary position

Property Value
URI https://w3id.org/APD/traits/trait_0012444
alternative label flower_ovary_position
description A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] of the plant ovary [PO:0009072] relative to the other flower [PO:0009046] organs [PO:0009008], in particular whether the ovary is attached above or below the perianth [PO:0009058] organs (the petals [PO:0009032] and sepals [PO:0009031]).
The position of the flower ovary relative to the insertion level of the remaining floral organs.
comments The ovary is the part of the gynoecium where the ovules are produced. The ovary may be located on the receptacle and thus be positioned above the insertion level of the remaining floral organs (i.e., the ovary is superior and the flower is hypogynous). Alternatively, the ovary may be embedded in the receptacle and therefore be located below the insertion level of the remaining floral organs (i.e., the ovary is inferior and the flower is epigynous). Flowers with a hypanthium may either have a superior ovary (perigyny; e.g., many Rosaceae) or an inferior ovary (epiperigyny). It is also possible that the ovary is inferior to a certain degree only, such as half-inferior, if the receptacle is surrounding the ovary to its mid-level. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘102. Ovary position (D1)’
value type categorical variable
has narrower flower_ovary_position_inferior Flower with inferior ovary, where the ovary is embedded in the receptacle and therefore located below the insertion level of the remaining floral organs.
flower_ovary_position_superior Flower with superior ovary, where the ovary is positioned above the insertion level of the remaining floral organs.
flower_ovary_position_half_inferior Flower where the receptacle is surrounded by the ovary to its mid-level.
flower_ovary_position_one_quarter_inferior Flower where the receptacle surrounds up to one quarter of the ovary.
flower_ovary_position_three_quarters_inferior Flower where the receptacle surrounds more than three quarters of the ovary.
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
orientation
position
keyword reproduction
gynoecium
carpel
plant ovary
perianth
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012444

Number of ovules per functional carpel

Property Value
URI https://w3id.org/APD/traits/trait_0012445
alternative label flower_ovules_per_functional_carpel_count
description A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of ovules [PO:0020003] per fertile [PATO:0000955] carpel [PO:0009030].
Number of ovules per carpel recorded as a continuous character (with integer values of 1 and above).
comments Reduced (sterile) carpels are not taken into account here. Definition derived from Schoenenberger et al. (2020) for their character ‘411. Number of ovules per functional carpel (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 1e+05
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
gynoecium
carpel
plant ovary
plant ovule
scope note Only applicable to fertile carpels.
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012445

Perianth differentiation

Property Value
URI https://w3id.org/APD/traits/trait_0012461
alternative label flower_perianth_differentiation
description A floral organ morphology trait [TO:0000862] which indicates whether petals [PO:0009032] and sepals [PO:0009031] are differentiated or identical and the various ways they can be identical or different.
The ways in which perianth organs may look different from each other in a given flower.
comments Typically, outer perianth parts are sepaloid and protect the other floral organs during floral development, while inner organs are often petaloid and play a role in pollinator attraction (Endress, 1994). However, it is also possible that all parts are either sepaloid or petaloid but remain differentiated in shape, size, and/or texture. In case of spiral perianths, differentiation may be continuous (i.e., gradual), whereby two successively initiated organs are very similar or only slightly different, while the outermost and innermost organs at both ends of the spiral are very different from each other (e.g., Chimonanthus). In the special case of perianths consisting of a single whorl, they are scored as undifferentiated. This perianth differentiation trait may be seen as both functional (the parting vs. sharing of functions among perianth parts) and developmental (the expression of a genetic program for different forms of perianth parts vs. a single program for a single type of perianth part morphology). Within-whorl differentiation, whereby organs of the same whorl take different forms, is common in zygomorphic flowers (e.g., Balsaminaceae, Fabaceae, Orchidaceae) but is not taken into account with this trait. This trait is not applicable when the perianth is absent. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘234. Perianth differentiation (D1)’
value type categorical variable
has narrower flower_perianth_differentiation_undifferentiated Flower were all tepals are alike, including flowers with a single, undifferentiated whorl.
flower_perianth_differentiation_marked_differentiation_outer_sepaloid_inner_petaloid A typical flower with a distinct calyx and corolla.
flower_perianth_differentiation_marked_differentiation_among_petaloid_tepals A flower with tepals that all resemble petals, but with marked differentiation between whorls of tepals.
flower_perianth_differentiation_marked_differentiation_among_sepaloid_tepals A flower with tepals that all resemble sepals, but with marked differentiation between whorls of tepals.
flower_perianth_differentiation_marked_differentiation A flower with marked differentiation between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.
flower_perianth_differentiation_continuous_differentiation_outer_sepaloid_inner_petaloid A flower with multiple whorls, where there is continuous differentiation between the outer sepaloid parts and the inner petaloid parts.
flower_perianth_differentiation_continuous_differentiation_among_petaloid_tepals A flower with series of tepals that all resemble petals, which display continuous, gradual differentiation between whorls.
flower_perianth_differentiation_continuous_differentiation_among_sepaloid_tepals A flower with series of tepals that all resemble sepals, which display continuous, gradual differentiation between whorls.
flower_perianth_differentiation_weak_differentiation A flower with weak between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.
flower_perianth_differentiation_very_weak_differentiation A flower with very weak between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic structure
shape
keyword reproduction
perianth
sepal
petal
corolla
calyx
scope note Not applicable when the perianth is absent
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012461

Fusion of perianth

Property Value
URI https://w3id.org/APD/traits/trait_0012462
alternative label flower_perianth_fusion
description A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]) that are fused [NCIT:C124537] together at anthesis [PO:0009068].
Fusion of perianth organs (congenital or postgenital) at anthesis, recorded on a continuous scale, from 0 (free parts) to 1 (parts fused along their entire length).
comments Partial fusion was recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the length of perianth parts). In case of multiple whorls, this trait records within-whorl fusion. For example, if organs within each whorl are fused along their entire length, a trait value of 1 is recorded. In cases where organs of two whorls are fused into a common tubular structure, such as frequently observed in monocots (e.g., Polygonatum), the perianth is considered to be fused. If the two (or more) whorls differed in their extent of fusion, a range of trait values may be recorded. For example, if the calyx is only basally fused, up to 10% of its length, but the corolla is entirely fused, a range of 0.1 to 1 is recorded as the trait value for the species. Definition derived from Schoenenberger et al. (2020) for their character ‘204. Fusion of perianth (C1)’
value type continuous variable
unit {dimensionless}
minAllowedValue 0
maxAllowedValue 1
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic dimensionless ratio
ratio; proportion
structure
keyword reproduction
perianth
sepal
petal
corolla
calyx
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012462

Number of perianth parts in each whorl (Perianth merism)

Property Value
URI https://w3id.org/APD/traits/trait_0012463
alternative label flower_perianth_merism
description A perianth morphology trait [TO:0000275] which is the count [NCIT:C25463] of perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]) in each floral whorl [PO:0025023].
The number of perianth parts in each whorl, recorded as a continuous trait (with integer values of 1 and above).
comments Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent. Definition derived from Schoenenberger et al. (2020) for their character ‘232. Perianth merism (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 50
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
orientation
keyword reproduction
perianth
sepal
petal
corolla
calyx
merism
whorl, floral whorl
scope note Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent.
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012463

Number of perianth parts

Property Value
URI https://w3id.org/APD/traits/trait_0012464
alternative label flower_perianth_parts_count
description A perianth morphology trait [TO:0000275] which is the total count [NCIT:C25463] of perianth [PO:0009058] parts, including sepals [PO:0009031], petals [PO:0009032], and any form of tepal [PO:0009033].
The total number of perianth parts, including sepals, petals, or any form of tepal.
comments A value of zero was scored when the perianth is absent. In flowers with perianth whorls fused along their complete length (e.g., Convolvulus), perianth parts are counted based on merism (e.g., if a calyx has 5 distinct sepals and the corolla is entirely fused, then the corolla is often interpreted to consist of five fused petals), anatomy (e.g., number of vascular traces), development (e.g., number of primordia), or comparison with closely related taxa. Definition derived from Schoenenberger et al. (2020) for their character ‘201. Number of perianth parts (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 1000
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
perianth
sepal
petal
corolla
calyx
scope note
has exact match tepal number
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012464

Perianth phyllotaxis

Property Value
URI https://w3id.org/APD/traits/trait_0012465
alternative label flower_perianth_phyllotaxis
description A perianth morphology trait [TO:0000275] which is the phyllotaxy [TO:0006014] of the perianth [PO:0009058] parts (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]), at anthesis [PO:0009068]
Perianth phyllotaxis at anthesis.
comments Perianth parts may be organised in one or more whorls or along a continuous spiral, usually with wide divergence angles more or less equal to 137.5 degrees. Less frequently, perianth phyllotaxis may be irregular. Perianth phyllotaxis at anthesis may differ from phyllotaxis of perianth part primordia at their inception and it is not uncommon that spirally initiated perianths become whorled later through development. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘230. Perianth phyllotaxy (D1)’, with more information of differentiating between trait values in this manuscript.
value type categorical variable
has narrower flower_perianth_phyllotaxis_irregular Perianth parts lacking any symmetrical organisation.
flower_perianth_phyllotaxis_spiral Perianth parts organised along a continuous spiral, usually with wide divergence angles more or less equal to 137.5 degrees, the Fibonacci ‘golden angle’.
flower_perianth_phyllotaxis_whorled Perianth parts organised in one or more whorls.
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
orientation
keyword reproduction
perianth
sepal
petal
corolla
calyx
whorl, floral whorl
phyllotaxy
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012465

Symmetry of perianth

Property Value
URI https://w3id.org/APD/traits/trait_0012466
alternative label flower_perianth_symmetry
description A perianth morphology trait [TO:0000275] which is the symmetry [PATO:0000965] of the perianth [PO:0009058] (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]), whereby a perianth is only considered actinomorphic [PATO:0001328] if all perianth whorls [PO:0025023] are actinomorphic.
Trait capturing the symmetry of the perianth.
comments This trait is applied to the perianth as a whole. In case of flowers with two or more perianth whorls, species are considered actinomorphic if all whorls are actinomorphic and as zygomorphic if one or more whorls are zygomorphic. Trait is not applicable when the perianth is missing. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘207. Symmetry of perianth (D1)’
value type categorical variable
has narrower flower_perianth_symmetry_actinomorphic_general Perianth displaying radial symmetry.
flower_perianth_symmetry_actinomorphic_rotational Perianth that can be equally divided into three or more identical sections that, when rotated around the center of the flower by some number of degrees, exactly match each other in orientation and shape.
flower_perianth_symmetry_actinomorphic_strictly_polysymmetric Polysymmetry, with three or more planes of bilateral symmetry.
flower_perianth_symmetry_actinomorphic_spiral Perianth lacking a distinct plane of symmetry because their perianth parts are spirally arranged.
flower_perianth_symmetry_asymmetric Perianth lacking any planes of symmetry.
flower_perianth_symmetry_disymmetric Perianth with two orthogonal planes of bilateral symmetry.
flower_perianth_symmetry_zygomorphic Monosymmetric, with a single plane of bilateral symmetry.
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic spatial pattern, distribution
symmetry
orientation
keyword reproduction
perianth
sepal
petal
corolla
calyx
whorl, floral whorl
scope note Not applicable when the perianth is missing
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012466

Number of perianth whorls

Property Value
URI https://w3id.org/APD/traits/trait_0012467
alternative label flower_perianth_whorls_count
description A perianth morphology trait [TO:0000275] which is the count [NCIT:C25463] of floral whorls [PO:0025023] that comprise the perianth [PO:0009058] (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]).
The number of perianth whorls as a continuous trait (with integer values of 1 and above).
comments Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent. Definition derived from Schoenenberger et al. (2020) for their character ‘231. Number of perianth whorls (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 15
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
perianth
sepal
petal
corolla
calyx
whorl, floral whorl
scope note Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent.
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012467

Pollen grain aperture shape

Property Value
URI https://w3id.org/APD/traits/trait_0012471
alternative label flower_pollen_aperture_shape
description A floral organ morphology trait [TO:0000862] which is the shape [PATO:0000052], structure [PATO:0000141], and position [PATO:0000140] of the pollen grain [PO:0025281] apertures [GO:0062074].
Pollen grains are often described according to the shape, structure, and position of their apertures. This trait describes the main shape of the many described aperture types.
comments Aperture terminology is partly determined by the position of the aperture on the pollen grain (either at the pole or at the equator) and pollen grain polarity, in turn, is determined by the spatial orientation of the microspore in the meiotic tetrad. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘5002. Aperture shape (D1)’’
value type categorical variable
has narrower flower_pollen_aperture_shape_absent Pollen that lacks an aperture.
flower_pollen_aperture_shape_colporus Pollen aperture shape that combines the groove of colpate and the pore of porate apertures.
flower_pollen_aperture_shape_colpus Pollen aperture shape is elongate and grooved and may be positioned globally or equatorially.
flower_pollen_aperture_shape_pore Pollen aperture shape is round and pore-like and may be positioned globally or equatorially.
flower_pollen_aperture_shape_ring_like Aperture that completely encircles the pollen.
flower_pollen_aperture_shape_spiraperturate Pollen grain with one or more spiral apertures.
flower_pollen_aperture_shape_sulcus Pollen aperture shape is elongate and grooved and positioned at the pole.
flower_pollen_aperture_shape_syncolpus Pollen grain with two or more simple (or compound) colpi, the ends of which join at the pole.
flower_pollen_aperture_shape_ulcus Pollen aperture shape is round and pore-like and positioned at the pole.
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic shape
structure
position
spatial pattern, distribution
keyword reproduction
androecium
stamen
anther
pollen aperture
pollen
pollen tube
anther theca
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012471

Number of pollen grain apertures

Property Value
URI https://w3id.org/APD/traits/trait_0012472
alternative label flower_pollen_apertures_count
description A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of pollen apertures [GO:0062074] per pollen [PO:0025281] grain.
The number of apertures per pollen grain as a continuous character.
comments A pollen aperture is a structurally delimited region of the pollen grain wall through which the pollen tube emerges during pollen germination and which plays a role in harmomegathy. A value of zero is scored when pollen is inaperturate (i.e. when there is no distinct aperture). Aperturate pollen is scored with integer values of 1 and above. Definition derived from Schoenenberger et al. (2020) for their character ‘5000. Number of apertures (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 100
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
androecium
stamen
anther
pollen aperture
pollen
pollen tube
anther theca
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012472

Pollen grain length

Property Value
URI https://w3id.org/APD/traits/trait_0012473
alternative label flower_pollen_length
description A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of a pollen [PO:0025281] grain.
The largest dimension of the pollen grain.
comments Palynologists typically describe pollen dimensions (and shape) using two measures, the polar axis (P) and the total breadth or diameter of the equatorial axis. Here we use P or E or another measure, whichever is the longest.
value type continuous variable
unit um
micrometre
minAllowedValue 0
maxAllowedValue 1000
has broader androecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic length
keyword reproduction
pollen
scope note
has exact match
has close match
has related match
examples
references no linked references
date created 19/05/2023
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012473

Number of structural carpels

Property Value
URI https://w3id.org/APD/traits/trait_0012481
alternative label flower_structural_carpels_count
description A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of both fertile [PATO:0000955] and female sterile [PATO:0000892] carpels [PO:0009030] in a hermaphrodite [PATO:0001340] flower [PO:0009046] or female [PATO:0000383] flower.
Number of fertile or sterile carpels in bisexual or female flowers, recorded as a continuous character (with integer values of 1 and above).
comments Includes the number of co-occurring carpellodes (sterile carpels) because this number is often more easily obtained from the literature than the actual number of fertile carpels. However, the number of carpellodes in male flowers is ignored for this character. In multicarpellate, unilocular gynoecia with complete carpel fusion up to the stigma (e.g., Primula), it may be difficult to assess the number of carpels unequivocally. In such cases, the number of carpels is scored only if it is well established based on atomical or developmental investigations. Similarly, in gynoecia where one or more carpels are reduced (e.g., in the pseudomonomerous gynoecia of some Arecaceae, Stauffer et al., 2002), the total number of structural carpels is only scored when unequivocally determined in the literature. Definition derived from Schoenenberger et al. (2020) for their character ‘401. Number of structural carpels (C1)’
value type continuous variable
unit {count}
minAllowedValue 0
maxAllowedValue 10000
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic has number of
count
keyword reproduction
gynoecium
carpel
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012481

Floral structural sex

Property Value
URI https://w3id.org/APD/traits/trait_0012482
alternative label flower_structural_sex_type
description A floral organ morphology trait [TO:0000862] which is the phenotypic sex [PATO:0001894] of a flower [PO:0009046], indicating whether individual flowers have both male [PATO:0000384] and female [PATO:0000383] reproductive structures [UBERON:0005156] (bisexual) or only one type of reproductive organ (unisexual).
Trait that captures floral sex, distinguishing among three states, bisexual, incompletely unisexual, and unisexual.
comments Defined by Schoenenberger et al. (2020) for their character ‘100. Floral structural sex (D1)’. See the trait ‘sex_type’ for plant sex type.
value type categorical variable
has narrower flower_structural_sex_type_bisexual Hermaphrodite; flowers with both male and female reproductive organs.
flower_structural_sex_type_incompletely_unisexual Male flowers with pistillodes and/or female flowers with staminodes.
flower_structural_sex_type_unisexual Flowers with either male or female reproductive organs.
has broader floral organ morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic structure
organismal quality
keyword reproduction
phenotypic sex
perianth
corolla
hermaphrodite
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012482

Style differentiation

Property Value
URI https://w3id.org/APD/traits/trait_0012483
alternative label flower_style_differentiation
description A floral organ morphology trait [TO:0000862] which is whether a style [PO:0006469] is present [PATO:0000467] or absent [PATO:0000462], and if present, the shape [PATO:0000052] and the length [PATO:0000122], width [PATO:0000921], and thickness [PATO:0000915] of the style relative to the dimensions of the ovary [PO:0009072].
Presence or absence of a style, and, when present, the shape of the style in terms of length and width in relation to ovary length and width.
comments This trait does not distinguish between fused or free styles. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘404. Style differentiation (D1)’
value type categorical variable
has narrower flower_style_differentiation_absent Stigma sessile.
flower_style_differentiation_present_length_and_shape_unknown Style present but shape unknown.
flower_style_differentiation_present_short_and_thick Style present, and short and thick.
flower_style_differentiation_present_short_and_narrow Style present, and short and narrow, including short filiform styles.
flower_style_differentiation_present_long_and_wide Style present, and long and wide.
flower_style_differentiation_present_long_and_narrow Style present, and long and narrow, including long filiform styles.
flower_style_differentiation_present_petaloid Style present and petaloid.
flower_style_differentiation_continuous When the style is an apical extension of carpel with decurrent stigma.
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic shape
amount (presence or absence in organism
2-D extent
structure
keyword reproduction
gynoecium
style
plant ovary
carpel
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012483

Fusion of styles

Property Value
URI https://w3id.org/APD/traits/trait_0012484
alternative label flower_style_fusion
description A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of styles [PO:0006469] that are fused [NCIT:C124537] together.
The degree of fusion of styles at anthesis, recorded on a continuous scale, from 0 (free styles) to 1 (styles fused along their entire length, but excluding the stigmatic region).
comments Partial fusion is recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the total length of the styles). This character is not applicable in unicarpellate flowers. Definition derived from Schoenenberger et al. (2020) for their character ‘406. Fusion of styles (C1)’
value type continuous variable
unit {dimensionless}
minAllowedValue 0
maxAllowedValue 1
has broader gynoecium morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic dimensionless ratio
ratio; proportion
structure
keyword reproduction
gynoecium
style
carpel
scope note
has exact match
has close match
has related match
examples
references Schönenberger 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 24/05/2023
reviewed by Hervé Sauquet
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012484

Petal length

Property Value
URI https://w3id.org/APD/traits/trait_0012485
alternative label flower_petal_length
description A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of a petal [PO_0009032].
The length of an individual flower petal.
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 1
maxAllowedValue 100
has broader perianth morphology trait
plant structure plant structure reproductive shoot system
flower
measured characteristic length
size
keyword reproduction
petal
corolla
perianth
petal morphology trait
scope note applicable to flowering plants
has exact match petal length
has close match
has related match
examples exact match: Flower petal length [TRY:3576] (https://www.try-db.org/de/de.php)
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012485

Fruit dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0012511
alternative label fruit_dry_mass
description A fruit morphology trait [TO:0002629] which is the dry [PATO:0001824] mass [PATO:0000125] of a fruit [PO:0009001], including seeds [PO:0009010].
Fruit dry mass, including seeds. Fruits will generally be oven-dried.
comments
value type continuous variable
unit mg
milligram
minAllowedValue 1e-04
maxAllowedValue 1e+06
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic mass
keyword reproduction
scope note
has exact match fruit dry weight
has close match
has related match
examples exact match: Fruit dry mass [TOP112] (https://top-thesaurus.org/index)
exact match: Fruit dry mass [TRY:919] (https://www.try-db.org/de/de.php)
exact match: Fruit dry mass [BROT:25] (http://doi.org/10.1038/sdata.2018.135)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012511

Fruit length

Property Value
URI https://w3id.org/APD/traits/trait_0012512
alternative label fruit_length
description A fruit morphology trait [TO:0002629] which is the length [PATO:0000122] of a fresh [EnvThes:21976] fruit [PO:0009001] from the fruit proximal end [PO:0008002] (base) to the fruit distal end [PO:0008001] (apex).
Linear dimension from the base to the apex of a fresh fruit, even if this is not the longest dimension.
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 2000
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic length
size
keyword reproduction
scope note
has exact match fruit length
has close match
has related match
examples exact match: Fruit length [TOP92] (https://top-thesaurus.org/index)
exact match: Fruit length [TRY:918] (https://www.try-db.org/de/de.php)
exact match: Fruit length [GIFT:3.13] (https://gift.uni-goettingen.de)
close match: maximum fruit length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
close match: minimum fruit length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: AverageFruitLength_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
exact match: MinFruitLength_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
exact match: MaxFruitLength_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012512

Fruit width

Property Value
URI https://w3id.org/APD/traits/trait_0012513
alternative label fruit_width
description A fruit morphology trait [TO:0002629] which is the width [PATO:0000921] of a fresh [EnvThes:21976] fruit [PO:0009001], where width is the greatest linear dimension [PATO:0001708] perpendicular to [PATO:0002434] to the fruit length [PATO:0000122] and fruit height [PATO:0000119].
Greatest linear width dimension of a fresh fruit, sometimes referred to as ‘diameter’ in fruits where the width and height are identical; orthogonal to the fruit length, the base-to-apex dimension.
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 2000
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic width
size
keyword reproduction
scope note
has exact match fruit width
has close match
has related match
examples exact match: Fruit width [TOP96] (https://top-thesaurus.org/index)
exact match: Fruit width (diameter) [TRY:917] (https://www.try-db.org/de/de.php)
exact match: Fruit width [GIFT:3.14] (https://gift.uni-goettingen.de)
exact match: AverageFruitWidth_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
exact match: MinFruitWidth_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
exact match: MaxFruitWidth_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012513

Fruit breadth

Property Value
URI https://w3id.org/APD/traits/trait_0012514
alternative label fruit_height
description A fruit morphology trait [TO:0002629] which is the height [PATO:0000119] of a fresh [EnvThes:21976] fruit [PO:0009001], where height is the shorter linear width dimension [PATO:0001708] perpendicular to [PATO:0002434] both the fruit length [PATO:0000122] and fruit width [PATO:0000921].
Shorter linear width dimension of a fresh fruit and generally not reported in fruits where the width and height are identical; orthogonal to the length, the base-to-apex dimension.
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 2000
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic height
size
keyword reproduction
scope note
has exact match
has close match
has related match
examples exact match: Fruit thickness [TOP100] (https://top-thesaurus.org/index)
exact match: Fruit height [GIFT:3.15] (https://gift.uni-goettingen.de)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note fruit_breadth
is in scheme https://w3id.org/APD/traits
identifier trait_0012514

Fruit wall thickness

Property Value
URI https://w3id.org/APD/traits/trait_0012515
alternative label fruit_wall_thickness
description A fruit morphology trait [TO:0002629] which is the thickness [PATO:0000915] of the entire pericarp [PO:0009084] (the exocarp [PO:0009085], mesocarp [PO:0009087], and endocarp [PO:0009086] combined), although it often refers just to the fleshy [PATO:0002351] components of the pericarp (most frequently the exocarp and mesocarp).
Fruit wall thickness of a mature fresh fruit. This term sometimes applies explicitly to the entire pericarp (exocarp + mesocarp + endocarp), but more frequently will have a functional meaning, referring to the outer fleshy layer which might be just some components of the peripcarp (most frequently the exocarp and mesocarp).
comments
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 1000
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic thickness
keyword reproduction
pericarp
exocarp
mesocarp
endocarp
scope note
has exact match
has close match
has related match
examples
references Simpson 2019
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note fruit_wall_width
is in scheme https://w3id.org/APD/traits
identifier trait_0012515

Fruit type

Property Value
URI https://w3id.org/APD/traits/trait_0012516
alternative label fruit_type
description A fruit morphology trait [TO:0002629] which is the botanical classification [NCIT:C25161] or type [NCIT:C25284] of a fruit [PO:0009001].
Botanical fruit type classification.
comments Although this trait is explicitly for fruit types, it includes a single term strobilus to describe the ‘fruiting body’ of non-flowering plants.
value type categorical variable
has narrower fruit_type_achene A simple, dry, indehiscent fruit with a single seed that is attached to the pericarp at one point only. (Synonyms, cypsela)
fruit_type_anthocarp A ripened ovary surrounded by attached accessory floral tissues; the floral tissues may be fleshy or dry. (Synonyms, accessory_fruit, pseudocarp, diclesium)
fruit_type_berry A simple, indehiscent fruit with one to many seeds embedded in a fleshy pericarp.
fruit_type_capsule A dry, dehiscent fruit composed of two or more united carpels; different types of capsules distinguished by type of dehiscence (e.g. valves, slits, pores).
fruit_type_caryopsis A simple, dry, indehiscent fruit with a single seed which is fused to the pericarp at one point only; the fruit type of all Poaceae (Synonyms, grain)
fruit_type_compound_fruit_other A fruit derived from more than one flower. (Synonyms, aggregate_fruit)
fruit_type_drupe A simple indehiscent fruit with a hard, stony endocarp and a fleshy mesocarp.
fruit_type_follicle A dry, dehiscent fruit derived from a single carpel that splits along one suture.
fruit_type_legume A dry, dehiscent fruit derived from a single carpel that splits along two sutures; applies to Fabaceae. (Synonyms, pod)
fruit_type_legume_indehiscent A dry, indehiscent fruit derived from a single carpel; applies to Fabaceae.
fruit_type_mericarp A fruitlet (seed-containing portion of a fruit) that is a half-carpel; the individual fruitlets separate from one another at maturity; a specific type of schizocarp. (In some datasets, term likely refers to any fruitlet derived from a portion of a schizocarp that separates at maturity, not explicitly just a half-carpel.)
fruit_type_multiple_fruit A fruit composed of several fruitlets developing from a single flower (e.g. raspberries, magnolias); fruits developing from more than one flower are included under compound (aggregate) fruits. (Synonym, apocarp)
fruit_type_nut A simple, dry, indehiscent fruit with a single seed and a hard pericarp.
fruit_type_nutlet A diminutive of nut, referring to a simple, dry, indehiscent structure with a single seed that might be an entire fruit (nut; monocarp) or a portion of a fruit, such as the individual components derived from an apocarp (fruit with multiple unfused ovaries) or schizocarp (fruit with multiple ovaries that separate at maturity).
fruit_type_pepo A fleshy fruit derived from a single ovary that is divided into several carpels, has multiple seeds, and has a tough rind when mature.
fruit_type_pome A fleshy fruit with the bulk of the fleshy tissue derived from flora tissue that surrounds the inferior ovary.
fruit_type_pyrene A fleshy fruit in which the seed(s) are covered by a hard layer of endocarp.
fruit_type_samara A winged, dry fruit.
fruit_type_schizocarp Category that includes all fruits derived from a compound pistil that separate into individual carpellary constituents at maturity; broad terms that includes dry and fleshy fruits, some with accessory tissues, others without. The individual seed-containing units that a schizocarp breaks apart into include mericarps, nutlets, and fruitlets.
fruit_type_silique A type of capsule (dry, dehiscent fruit) where the fruit is divided into two parts by a thin partition; more than three times as long as wide; applies to Brassicaceae (Synonyms, silicle, siliqua)
fruit_type_strobilus A modified reproductive shoot system found in many nonflowering plants, especially gymnosperms, comprised of overlapping bracts (scales). (Synonyms, cone)
fruit_type_syncarp A compound fruit produced by the adhesion of the fruits from several flowers.
fruit_type_syconium A fleshy compound fruit, in which the fruitlets are enclosed in accessory material, such as the hollow-centered fruits of Moraceae. (Synonyms, synconium, fig)
fruit_type_utricle A bladdery or inflated dry fruit with a single seed; essential an achene in which the pericarp is significantly larger than the mature seed.
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic structure
composition
shape
keyword reproduction
scope note
has exact match fruit type
has close match
has related match
examples exact match: Fruit type [TRY:99] (https://www.try-db.org/de/de.php)
exact match: Fruit_type_1 [GIFT:3.16.1] (https://gift.uni-goettingen.de)
exact match: fruit type [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Fruit type [BROT:26] (http://doi.org/10.1038/sdata.2018.135)
references Kew Seed Information Database 2022
Beentje 2010
Spjut 2003
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012516

Fruit fleshiness

Property Value
URI https://w3id.org/APD/traits/trait_0012517
alternative label fruit_fleshiness
description A fruit morphology trait [TO:0002629] which indicates whether a mature [PATO:0001701] fresh [EnvThes:21976] fruit [PO:0009001] is fleshy [PATO:0002351].
Binary variable, dividing entire, true fruits into ‘dry’ versus ‘fleshy’ based on whether their botanical fruit type is fleshy when mature and fresh.
comments This trait explicitly refers to fruit fleshiness, not diaspore or seed fleshiness; diaspore fleshiness is a separate trait.
value type categorical variable
has narrower fruit_fleshiness_dry Fruits are dry when mature and fresh.
fruit_fleshiness_fleshy Fruits are fleshy when mature and fresh.
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic composition
structure
texture
keyword reproduction
fleshy
scope note
has exact match
has close match
has related match
examples close match: Fruit type [TRY:99] (https://www.try-db.org/de/de.php)
exact match: Fruit_dryness_1 [GIFT:3.18.1] (https://gift.uni-goettingen.de)
references Kew Seed Information Database 2022
Beentje 2010
Spjut 2003
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012517

Fruit dehiscence

Property Value
URI https://w3id.org/APD/traits/trait_0012518
alternative label fruit_dehiscence
description A fruit morphology trait [TO:0002629] which indicates whether mature [PATO:0001701] fruits [PO:0009001] are dehiscent [GO:0009900].
Binary variable, dividing fruits into ‘dehiscent’ versus ‘indehiscent’ based on whether the individual fruit units are dehiscent when mature.
comments
value type categorical variable
has narrower fruit_dehiscence_dehiscent Fruits are dehiscent when mature.
fruit_dehiscence_indehiscent Fruits are indehiscent when mature.
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic structure
keyword reproduction
dehiscence
scope note
has exact match
has close match
has related match
examples close match: Fruit type [TRY:99] (https://www.try-db.org/de/de.php)
exact match: Dehiscence_1 [GIFT:3.17.1] (https://gift.uni-goettingen.de)
references Kew Seed Information Database 2022
Beentje 2010
Spjut 2003
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012518

Fruit colour

Property Value
URI https://w3id.org/APD/traits/trait_0012519
alternative label fruit_colour
description A fruit morphology trait [TO:0002629] which is the primary [NCIT:C25251] colour [PATO:0000014] of the exocarp [PO:0009085] of a mature [PATO:0001701] fruit [PO:0009001].
Human visible dominant colour of a fresh fruit’s exocarp (outermost layer) at maturity.
comments If a fruit is multiple colours, they are listed as a space-delimited string.
value type categorical variable
has narrower fruit_colour_pink Fruit’s exocarp is pink.
fruit_colour_red Fruit’s exocarp is red.
fruit_colour_orange Fruit’s exocarp is orange.
fruit_colour_yellow Fruit’s exocarp is yellow.
fruit_colour_green Fruit’s exocarp is green.
fruit_colour_blue Fruit’s exocarp is blue.
fruit_colour_purple Fruit’s exocarp is purple.
fruit_colour_white Fruit’s exocarp is white.
fruit_colour_cream Fruit’s exocarp is cream.
fruit_colour_brown Fruit’s exocarp is brown.
fruit_colour_black Fruit’s exocarp is black.
fruit_colour_grey Fruit’s exocarp is grey.
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic colour / color
keyword reproduction
exocarp
pericarp
scope note
has exact match fruit color
exocarp color
has close match
has related match
examples exact match: Fruit/seed color [TRY:585] (https://www.try-db.org/de/de.php)
close match: Dispersal unit color [TRY:392] (https://www.try-db.org/de/de.php)
exact match: Fruit_colour [GIFT:3.22.1] (https://gift.uni-goettingen.de)
exact match: MainFruitColors [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
close match: FruitColorDescription [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references Kissling 2019
Delmas 2020
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012519

Fruit surface hairiness

Property Value
URI https://w3id.org/APD/traits/trait_0012521
alternative label fruit_surface_hairs
description A fruit morphology trait [TO:0002629] which is the presence or absence of trichomes [PO:0006504] (hairs) on the surface (exocarp [PO:0009085]) of a mature [PATO:0001701] fruit [PO:0009001].
Surface hairiness of a mature fruit, including vestiture (trichome cover) and bristles.
comments
value type categorical variable
has narrower fruit_surface_hairs_glabrous Mature fruit do not have hairs (trichomes).
fruit_surface_hairs_hairy Mature fruit have hairs (trichomes).
has broader fruit morphology trait
plant structure plant structure reproductive shoot system
fruit
measured characteristic texture
pilosity
keyword trichome
fruit morphology trait
scope note
has exact match fruit hairiness
has close match
has related match
examples
references no linked references
date created 01/04/2025
date modified 01/04/2025
date reviewed
reviewed by no reviewers
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012521

Seed dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0012610
alternative label seed_dry_mass
description A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of a mature [PATO:0001701] seed [PO:0009010].
Dry mass of a mature seed, including both oven dried and air-dried samples.
comments Standard methods people will have used to dry seeds include, ‘fresh’ (at dispersal, mature); ‘air dried’ (at local ambient conditions); ‘seed bank air dried’ (to 15% relative humidity); and ‘oven dried’ (>100 deg C for a set number of hours; e.g. seed bank standard is 103 deg C for 17 hours). It is expected that some observations in AusTraits mapped onto ‘seed_dry_mass’ will actually include both the seed and some dispersal tissue, if the two cannot easily be separated; these should be mapped to ‘diaspore_dry_mass’.
value type continuous variable
unit mg
milligram
minAllowedValue 1e-05
maxAllowedValue 1e+06
has broader seed morphology trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
keyword reproduction
scope note
has exact match seed weight, seed mass
seed mass
has close match
has related match
examples exact match: Seed dry mass [TOP111] (https://top-thesaurus.org/index)
exact match: Seed dry mass [TRY:26] (https://www.try-db.org/de/de.php)
close match: seed weight & seed shape [LEDA:5.2] (https://uol.de/en/landeco/research/leda/standards)
exact match: Seed mass [GIFT:3.2] (https://gift.uni-goettingen.de)
exact match: seed mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
exact match: Seed dry mass [BROT:27] (http://doi.org/10.1038/sdata.2018.135)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
Saatkamp 2018
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note seed_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0012610

Diaspore dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0012611
alternative label diaspore_dry_mass
description A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of a seed [PO:0009010] and the seed dispersal [EnvThes:21042] structures [PATO:0000141].
Dry mass of a plant’s dispersal unit, including the seed and any dispersal appendages; synonymous with the term dispersule and often synonymous with the term disseminule.
comments Data mapped onto this trait have been very explicitly defined as a ‘diaspore’. There will be additional values mapped as either ‘seed_dry_mass’ or ‘fruit_dry_mass’ that are also diaspores, but have identified only as being a ‘seed’ or ‘fruit’. In addition, it is expected that some observations in AusTraits mapped onto ‘seed_dry_mass’ will actually include both the seed and some dispersal tissue, if the two cannot easily be separated.
value type continuous variable
unit mg
milligram
minAllowedValue 1e-05
maxAllowedValue 1e+06
has broader seed morphology trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
keyword dispersal
reproduction
seed dispersal
scope note
has exact match
has close match
has related match
examples exact match: Dispersule dry mass trait [TOP110] (https://top-thesaurus.org/index)
exact match: Dispersal unit dry mass [TRY:233] (https://www.try-db.org/de/de.php)
close match: seed weight & seed shape [LEDA:5.2] (https://uol.de/en/landeco/research/leda/standards)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note diaspore_mass
is in scheme https://w3id.org/APD/traits
identifier trait_0012611

Seed embryo and endosperm dry mass

Property Value
URI https://w3id.org/APD/traits/trait_0012612
alternative label seed_dry_mass_reserve
description A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of the components of a seed [PO:0009010] that provide energy [PATO:0001021] to a seedling [EnvThes:20698], generally the embryo [PO:0009009] and, if present, the endosperm [PO:0009089].
Dry mass of a seed’s embryo and endosperm (if present), the components of a seed that provide energy to a seedling. These tissues are distinct from a seed’s dispersal structures or protective tissues.
comments
value type continuous variable
unit mg
milligram
minAllowedValue 1e-05
maxAllowedValue 1e+05
has broader seed morphology trait
plant structure plant structure reproductive shoot system
seed
measured characteristic mass
keyword energy
plant embryo
endosperm
reproduction
seedling
storage organ
scope note
has exact match
has close match
has related match
examples
references Westoby 1990
Lamont & Groom 2013
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note seed_mass_reserve
is in scheme https://w3id.org/APD/traits
identifier trait_0012612

Seed length

Property Value
URI https://w3id.org/APD/traits/trait_0012613
alternative label seed_length
description A seed morphology trait [TO:0000184] which is the greatest linear dimension [PATO:0001708] (axis) of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed width [PATO:0000921] and seed height [PATO:0000119] axes [NCIT:C25154].
Longest linear seed dimension.
comments Observations of this trait in AusTraits are almost certainly an undocumented mix of dimensions of true seeds and diaspores. When all or part of the fruit is not easily separated from the true seed, many researchers will have recorded and submitted a diaspore length, but labeled the data ‘seed length’. The intent of this measurement is that it excludes appendages, although it is likely some submitted measurements include small appendages that are not easily separated from the seed.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 1000
has broader seed morphology trait
plant structure plant structure reproductive shoot system
seed
measured characteristic length
size
keyword reproduction
scope note
has exact match seed length
has close match
has related match
examples exact match: Seed length [TOP91] (https://top-thesaurus.org/index)
exact match: Seed length [TRY:27] (https://www.try-db.org/de/de.php)
close match: seed weight & seed shape [LEDA:5.2] (https://uol.de/en/landeco/research/leda/standards)
exact match: Seed length [GIFT:3.10] (https://gift.uni-goettingen.de)
exact match: seed length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references Pérez-Harguindeguy 2013
Kew Seed Information Database 2022
Saatkamp 2018
date created 14/07/2021
date modified 05/02/2024
date reviewed 30/11/2021
reviewed by Elizabeth Wenk
Hervé Sauquet
Russell Barrett
Carl Gosper
Lydia Guja
Gregory J. Jordan
Mark Ooi
Karen D. Sommerville
Lily Dun
change note
is in scheme https://w3id.org/APD/traits
identifier trait_0012613

Seed width

Property Value
URI https://w3id.org/APD/traits/trait_0012614
alternative label seed_width
description A seed morphology trait [TO:0000184] which is the longest [NCIT:C25558] width [PATO:0000921] dimension of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed length [PATO:0000122] and seed height [PATO:0000119] axes [NCIT:C25154].
Longest width dimension of a seed; orthogonal to the length and height axes.
comments Observations of this trait in AusTraits are almost certainly an undocumented mix of dimensions of true seeds and diaspores. When all or part of the fruit is not easily separated from the true seed, many researchers will have recorded and submitted a diaspore width, but labeled the data ‘seed width’. The intent of this measurement is that it excludes appendages, although it is likely some submitted measurements include small appendages that are not easily separated from the seed.
value type continuous variable
unit mm
millimetre
minAllowedValue 0.01
maxAllowedValue 1000