The AusTraits Plant Dictionary (APD) Ontology

Release 2.1.0

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https://w3id.org/APD/index.html

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https://w3id.org/APD/release/2.1.0/index.html

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https://w3id.org/APD/release/2.0.0/index.html

Authors:

Elizabeth Wenk, Daniel Falster, Rachael Gallagher, Hervé Sauquet, Rowan Brownlee, Carl Boettiger

Contributors:

Tony Auld, Russell Barrett, Timothy Brodribb, Brendan Choat, David Coleman, Félix de Tombeur, Lily Dun, David Ellsworth, Carl Gosper, Lydia Guja, Gregory J. Jordan, Tom Le Breton, Andy Leigh, Patricia Lu-Irving, Belinda Medlyn, Rachael Nolan, Mark Ooi, Karen D. Sommerville, Ruby Stephens, Peter Vesk, Matthew White, Ian J. Wright, Sophie Yang

Human readable tables:

Serializations:

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Vocabulary service:

Source:

Contributing:

Cite as:

Wenk EH, Sauquet H, Gallagher RV, Brownlee R, Boettiger C, Coleman D, Yang S, Auld T, Barrett RL, Brodribb T, Choat B, Dun L, Ellsworth D, Gosper C, Guja L, Jordan GJ, Breton T, Leigh A, Irving PMedlyn B, Nolan R, Ooi M, Sommerville KD, Vesk P, White M, Wright IJ & Falster DS (2024) The AusTraits Plant Dictionary. Scientific Data 11, 537. DOI: 10.1038/s41597-024-03368-z

Abstract

This document describes the schema available from the APD namespace. The APD offers detailed descriptions for more than 500 plant trait concepts. It includes trait focused on plant morphology, plant nutrient concentrations, plant physiology, plant life history, and plant fire response. The definitions will be useful to researchers from diverse disciplines, including plant functional ecology, plant taxonomy, and conservation biology. All trait concepts are supported by comprehensive metadata including trait descriptions, allowable trait values, allowable ranges, preferred units, keywords, references, and links to matches in a selection of trait databases. The traits describe here also fully support the AusTraits plant trait database, <doi.org/10.5281/zenodo.3568417>.

Table of Contents

1. Introduction
2. Trait groups: Hierarchical trait groupings, included to enhance trait findability.
3. Trait concepts: Traits defined within the vocabulary.
4. Values for categorical traits: Allowed categorical trait values to support the defined trait concepts.
5. Glossary: Terms used as keywords that were not available in other published vocabularies.
6. Acknowledgements

1. Introduction

The APD is a compilation of the traits concepts used by AusTraits, a database of Australian plant traits. APD includes definitions for nearly 500 traits pertaining to plant functional ecology and plant morphology. The metadata for each trait concept has been reviewed by multiple people. Metadata includes a trait definition, accepted units and ranges (for numeric traits), allowable values (for categorical traits), keywords and trait groupings to enhance trait findability, references, and links to identical/similar traits in other trait databases. APD will be published in a machine-readable format (rdf representation), allowing the traits to be readily re-used by other databases. This is a human-friendly output, with embedded links to the references, other trait databases and keywords/concepts borrowed from other ontologies.

The APD is a community resource and we welcome any suggestions for edits, including additional key references for each trait or other trait databases for which we should add links.

2. Trait groups

The APD Trait groups express broad trait groupings, with each trait concept mapped into one or more trait groups as narrower concepts.

plant trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000000
description	A measurable or observable characteristic of a cellular component [GO:0005575], biological process [GO:0008150] or molecular function [GO:0003674] that is part of, or has participant a plant anatomical entity [PO:0025131] and/or a plant structure development stage [PO:0009012]. This includes physiological, molecular and biochemical traits as well.
has narrower	biochemical trait plant morphology trait biological process trait, physiological process trait life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000000

biochemical trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000002
description	A plant trait [TO:0000387] which is a quality of a chemical entity [CHEBI:24431], a plant metabolite [CHEBI:76924], or a molecular function [GO:0003674], which occurs in a cellular component [GO:0005575], plant anatomical entity [PO:0025131] or during a plant structure development stage [PO:0009012].
has narrower	mineral and ion content trait metabolite content trait stable isotope ratio determination
has broader	plant trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000002

mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000004
description	A biochemical trait [TO:0000277] that is the amount of a particular mineral or ion constituent occurring in a plant anatomical entity [PO:0025131].
has narrower	leaf mineral and ion content trait stem mineral and ion content trait bark mineral and ion content trait root mineral and ion content trait reproductive shoot system mineral and ion content trait cell or tissue mineral and ion content trait
has broader	biochemical trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000004

leaf mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000006
description	A mineral and ion content trait which is measured on a leaf [PO:0025034].
has narrower	live leaf mineral and ion content trait senesced leaf mineral and ion content trait
has broader	mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000006

live leaf mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000008
description	A mineral and ion content trait which is measured on a live [PATO:0001421] leaf [PO:0025034].
has narrower	Leaf aluminium (Al) content per unit leaf dry mass Leaf boron (B) content per unit leaf dry mass Leaf carbon (C) content per unit leaf dry mass Leaf calcium (Ca) content per unit leaf dry mass Leaf chlorine (Cl) content per unit leaf dry mass Leaf chromium (Cr) content per unit leaf dry mass Leaf cobalt (Co) content per unit leaf dry mass Leaf copper (Cu) content per unit leaf dry mass Leaf iron (Fe) content per unit leaf dry mass Leaf potassium (K) content per unit leaf area Leaf potassium (K) content per unit leaf dry mass Leaf magnesium (Mg) content per unit leaf dry mass Leaf manganese (Mn) content per unit leaf dry mass Leaf molybdenum (Mo) content per unit leaf dry mass Leaf nitrogen (N) content per unit leaf area Leaf nitrogen (N) content per unit leaf dry mass Leaf sodium (Na) content per unit leaf dry mass Leaf nickel (Ni) content per unit leaf dry mass Leaf phosphorus (P) content per unit leaf area Leaf phosphorus (P) content per unit leaf dry mass Leaf sulphur (S) content per unit leaf dry mass Leaf selenium (Se) content per unit leaf dry mass Leaf silicon (Si) content per unit leaf dry mass Leaf zinc (Zn) content per unit leaf dry mass Leaf carbon to nitrogen ratio (C/N Leaf nitrogen to phosphorus ratio (N/P) per unit leaf dry mass Leaf cell wall nitrogen (N) per unit cell wall dry mass Leaf cell wall nitrogen (N) per unit leaf N content Lipid fraction of leaf P content per dry mass Nucleic acid fraction of leaf P content per dry mass Residual P fraction of leaf P content per dry mass Inorganic P fraction of leaf P content per dry mass Metabolite fraction of leaf P content per dry mass Leaf ash content per unit leaf dry mass Leaf photosynthetic nitrogen use efficiency during Amax measurement (PNUE Leaf photosynthetic nitrogen use efficiency during Asat measurement (PNUE Leaf photosynthetic phosphorus use efficiency during Amax measurement PPUE Leaf photosynthetic phosphorus use efficiency during Asat measurement (PPUE
has broader	leaf mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000008

senesced leaf mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000108
description	A mineral and ion content trait which is measured on a senescent [PATO:0001487] leaf [PO:0025034].
has narrower	Senesced leaf aluminium (Al) content per unit leaf dry mass Senesced leaf boron (B) content per unit leaf dry mass Senesced leaf carbon (C) content per unit leaf dry mass Senesced leaf calcium (Ca) content per unit leaf dry mass Senesced leaf copper (Cu) content per unit leaf dry mass Senesced leaf iron (Fe) content per unit leaf dry mass Senesced leaf potassium (K) content per unit leaf dry mass Senesced leaf magnesium (Mg) content per unit leaf dry mass Senesced leaf manganese (Mn) content per unit leaf dry mass Senesced leaf molybdenum (Mo) content per unit leaf dry mass Senesced leaf nitrogen (N) content per unit leaf dry mass Senesced leaf sodium (Na) content per unit leaf dry mass Senesced leaf nickel (Ni) content per unit leaf dry mass Senesced leaf phosphorus (P) content per unit leaf dry mass Senesced leaf sulphur (S) content per unit leaf dry mass Senesced leaf zinc (Zn) content per unit leaf dry mass
has broader	leaf mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000108

stem mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000206
description	A mineral and ion content trait which is measured on a stem [PO:0009047].
has narrower	wood mineral and ion content trait senesced wood mineral and ion content trait Stem carbon (C) content per unit stem dry mass Stem nitrogen (N) content per unit stem dry mass Bark aluminium (Al) content per unit bark dry mass Bark boron (B) content per unit bark dry mass Bark carbon (C) content per unit bark dry mass Bark calcium (Ca) content per unit bark dry mass Bark copper (Cu) content per unit bark dry mass Bark iron (Fe) content per unit bark dry mass Bark potassium (K) content per unit bark dry mass Bark magnesium (Mg) content per unit bark dry mass Bark manganese (Mn) content per unit bark dry mass Bark nitrogen (N) content per unit bark dry mass Bark sodium (Na) content per unit bark dry mass Bark phosphorus (P) content per unit bark dry mass Bark sulphur (S) content per unit bark dry mass Bark zinc (Zn) content per unit bark dry mass Bark ash content per unit bark dry mass
has broader	mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000206

wood mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000408
description	A mineral and ion content trait which is measured on wood [NCIT:C93062].
has narrower	Wood carbon (C) content per unit wood dry mass Wood calcium (Ca) content per unit wood dry mass Wood potassium (K) content per unit wood dry mass Wood magnesium (Mg) content per unit wood dry mass Wood nitrogen (N) content per unit wood dry mass Wood sodium (Na) content per unit wood dry mass Wood phosphorus (P) content per unit wood dry mass
has broader	stem mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000408

senesced wood mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000508
description	A mineral and ion content trait which is measured on senescent [PATO:0001487] wood [NCIT:C93062].
has narrower	Dead wood calcium (Ca) content per unit dead wood dry mass Dead wood potassium (K) content per unit dead wood dry mass Dead wood magnesium (Mg) content per unit dead wood dry mass Dead wood nitrogen (N) content per unit dead wood dry mass Dead wood sodium (Na) content per unit dead wood dry mass Dead wood phosphorus (P) content per unit dead wood dry mass
has broader	stem mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000508

bark mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000608
description	A mineral and ion content trait which is measured on bark [PO:0004518].
has broader	mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000608

root mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0000808
description	A mineral and ion content trait which is measured on a root [PO:0009005].
has narrower	Root carbon (C) content per unit root dry mass Root nitrogen (N) content per unit root dry mass Root phosphorus (P) content per unit root dry mass
has broader	mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0000808

reproductive shoot system mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0001006
description	A mineral and ion content trait which is measured on the reproductive shoot system [PO:0025082].
has narrower	Flower nitrogen (N) content per unit flower dry mass Fruit calcium (Ca) content per unit fruit dry mass Fruit potassium (K) content per unit fruit dry mass Fruit magnesium (Mg) content per unit fruit dry mass Fruit nitrogen (N) content per unit fruit dry mass Fruit phosphorus (P) content per unit fruit dry mass Fruit sulphur (S) content per unit fruit dry mass Seed calcium (Ca) content per unit seed dry mass Seed potassium (K) content per unit seed dry mass Seed magnesium (Mg) content per unit seed dry mass Seed nitrogen (N) content per unit seed dry mass Seed phosphorus (P) content per unit seed dry mass Seed sulphur (S) content per unit seed dry mass
has broader	mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0001006

cell or tissue mineral and ion content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0001502
description	A mineral and ion content trait which is measured on an individual tissue [CARO:0000043] type or cell [CL:0000000] type.
has narrower	Leaf rubisco nitrogen (N) content per unit leaf N content Leaf thylakoid protein nitrogen (N) content per unit leaf N content Leaf epidermis calcium (Ca) content per unit leaf fresh mass Leaf hypodermis calcium (Ca) content per unit leaf fresh mass Leaf internal parenchyma cell calcium (Ca) content per unit leaf fresh mass Leaf palisade mesophyll cell calcium (Ca) content per unit leaf fresh mass Leaf sclerenchyma cell calcium (Ca) content per unit leaf fresh mass Leaf spongy mesophyll cell calcium (Ca) content per unit leaf fresh mass Leaf epidermis phosphorus (P) content per unit leaf fresh mass Leaf hypodermis phosphorus (P) content per unit leaf fresh mass Leaf internal parenchyma cell phosphorus (P) content per unit leaf fresh mass Leaf palisade mesophyll cell phosphorus (P) content per unit leaf fresh mass Leaf sclerenchyma cell phosphorus (P) content per unit leaf fresh mass Leaf spongy mesophyll cell phosphorus (P) content per unit leaf fresh mass
has broader	mineral and ion content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0001502

metabolite content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0002004
description	The trait associated with an assay to determine the affected metabolites profile.
has narrower	carbohydrate content trait lipid content trait phenolic compound content trait pigment content trait protein content trait
has broader	biochemical trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0002004

carbohydrate content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0002005
description	A metabolite content trait [TO:0000281] which is the amount of carbohydrate [CHEBI:16646] in a plant structure [PO:0009011].
has narrower	Leaf total non-structural carbohydrate content per unit leaf area Leaf total non-structural carbohydrate content per unit leaf dry mass Leaf cellulose content per unit leaf dry mass Leaf starch content per unit leaf area Leaf soluble starch content per unit leaf area Leaf soluble starch content per unit leaf dry mass Leaf soluble sugar content per unit leaf area Leaf soluble sugar content per unit leaf dry mass Stem soluble starch content per unit stem dry mass Stem soluble sugar content per unit stem dry mass Bark cellulose content per unit bark dry mass Root soluble starch content per unit root dry mass Root soluble sugar content per unit root dry mass
has broader	metabolite content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0002005

lipid content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0002006
description	The total lipid [CHEBI:18059] content measured in a plant or a plant part.
has narrower	Seed oil content per unit seed dry mass
has broader	metabolite content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0002006

phenolic compound content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0002007
description	Amount of phenolics present in the plant part, whole plant or product.
has narrower	Leaf lignin content per unit leaf dry mass Total leaf phenolic content per unit leaf dry mass Leaf tannin content per unit leaf dry mass Bark lignin content per unit bark dry mass Bark tannin content per unit bark dry mass
has broader	metabolite content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0002007

pigment content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0002008
description	A metabolite content trait [TO:0000281] which is the amount of pigment [NCIT:C747] in a plant structure [PO:0009011].
has narrower	Leaf carotenoid content per unit leaf area Leaf carotenoid content per unit leaf dry mass Leaf total chlorophyll content (chlorophyll A + B) per unit leaf area Leaf total chlorophyll content (chlorophyll A + B) per unit leaf dry mass Leaf chlorophyll A content per unit leaf area Leaf chlorophyll A content per unit leaf dry mass Leaf chlorophyll B content per unit leaf area Leaf chlorophyll B content per unit leaf dry mass Ratio of leaf chlorophyll A content to leaf chlorophyll B content Leaf chlorophyll content in SPAD units
has broader	metabolite content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0002008

protein content trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0002009
description	The total protein content measured in a plant or a plant part.
has narrower	Leaf soluble protein content per unit leaf area Leaf insoluble protein content per unit leaf area Leaf rubisco content per unit leaf dry mass Seed protein content per unit seed dry mass
has broader	metabolite content trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0002009

stable isotope ratio determination

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0003004
description	A mineral and ion content trait [TO:0000465] which is a ratio [PATO:0001470] of two naturally occurring stable isotopes [EnvThes:20310], generally expressed in delta notation.
has narrower	Bark stable carbon isotope composition (delta13C Leaf stable carbon isotope composition (delta13C Stem stable carbon isotope composition (delta13C Root stable carbon isotope composition (delta13C Wood stable carbon isotope composition (delta13C Bark stable nitrogen isotope composition (delta15N Leaf stable nitrogen isotope composition (delta15N Stem stable nitrogen isotope composition (delta15N Root stable nitrogen isotope composition (delta15N Wood stable nitrogen isotope composition (delta15N Leaf xylem stable nitrogen isotope composition (delta15N Root xylem stable nitrogen isotope composition (delta15N Leaf stable oxygen isotope composition (delta18O Stem water stable oxygen isotope composition (delta18O
has broader	biochemical trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0003004

plant morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0010001
description	A plant trait [TO:0000387] which is a morphological quality of a plant anatomical entity [PO:0025131] or a constituent cellular component [GO:0005575] contained therein. Includes traits of plant structures [PO:0009011] such as plant cell [PO:0009002], plant organ [PO:0009008], portion of plant tissue [PO:0009007], vascular system [PO:0000034], whole plant [PO:0000003] and seed [PO:0009010], as well as those of plant anatomical spaces [PO:0025117] and portions of plant substance [PO:0025161].
has narrower	whole plant morphology trait plant structure morphology trait vascular tissue morphology trait plant structure strength trait
has broader	plant trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0010001

whole plant morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0010004
description	A plant structure morphology trait [TO:0000839] which is a quality of a whole plant [PO:0000003].
has narrower	plant embryo morphology trait Plant canopy width Plant canopy breadth Plant vegetative height Height of a climbing plant Plant reproductive height Plant growth form Life form
has broader	plant morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0010004

plant embryo morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0010104
description	A whole plant morphology trait [TO:0000398] which is a quality of a plant embryo [PO:0009009].
has narrower	Embryo colour Cotyledon function Cotyledon position at germination Cotyledon hairiness Hypocotyl hairiness Seedling first true leaf type Seedling first node leaf count Seedling germination location
has broader	whole plant morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0010104

plant structure morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011001
description	A plant morphology trait [TO:0000017] which is a quality of a plant structure [PO:0009011].
has narrower	leaf morphology trait portion of plant tissue morphology trait plant cell morphology trait stem morphology trait bark morphology trait root system morphology trait reproductive shoot system morphology trait Plant spinescence Physical defence structures Plant sex type Clonal spread mechanism Storage organ Plant fire tolerance strategy
has broader	plant morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011001

leaf morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011202
description	A phyllome morphology trait [TO:0000747] which is quality of a leaf [PO:0025034].
has narrower	leaf size trait leaf mass trait leaf shape trait leaf position trait leaf stomatal complex morphology trait leaf optical properties trait Cotyledon function Number of leaflets per leaf Leaf glaucousness Mature leaf hairiness Juvenile phase leaf hairiness Immature leaf hairiness Leaf surface colour Leaf surface reflectivity Average leaf epidermal cell density Lower leaf side epidermal cell density Upper leaf side epidermal cell density Palisade cell length Palisade cell width Number of layers of palisade cells Spongy mesophyll cell thickness Xylem to leaf area ratio (XLA Leaf secondary vein angle Major leaf vein density Length of all minor and major leaf lamina veins per unit area Leaf vein frequency Leaf xylem vessel density Leaf xylem vessel diameter Leaf xylem vessel wall thickness Leaf xylem vessel length Leaf work to punch Leaf specific work to punch Leaf work to shear Leaf specific work to shear (fracture toughness Leaf work to tear Leaf specific work to tear Plant photosynthetic organ Physical defence structures Plant climbing mechanisms Plant succulence
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011202

leaf size trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011204
description	Refer to size [PATO:0000117]: A morphology quality inhering in a bearer by virtue of the bearers physical magnitude.
has narrower	Leaf area Leaflet area Leaf length Leaf width Leaf thickness Petiole length Petiole width Leaflet length Leaflet width Leaf mass per area Leaf lamina mass per area Leaf area ratio (LAR Leaf fresh mass per leaf area
has broader	leaf morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011204

leaf mass trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011206
description	A leaf morphology trait [TO:0000505] which is the weight of a leaf [PO:0025034].
has narrower	Leaf dry mass Leaflet dry mass Leaf fresh mass Leaf mass per area Leaf lamina mass per area Leaf tissue density Leaf mass fraction Leaf dry matter content (LDMC Leaf fresh mass per leaf area Leaf water content per unit leaf area (leaf succulence Leaf water content per unit leaf dry mass Leaf water content per unit leaf fresh mass Leaf water content per unit saturated leaf mass Leaf cell wall fraction Leaf dry mass to stem dry mass ratio Stem dry mass to vegetative shoot dry mass ratio (support fraction
has broader	leaf morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011206

leaf shape trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011304
description	A leaf morphology trait [TO:0000748] which is the variation in shapes and forms of a leaf [PO:0025034]. Refer to shape [PATO:0000052]: A morphological quality inhering in a bearer by virtue of the bearers ratios of distances between its features [points, edges, surfaces and also holes etc].
has narrower	Plant spinescence Leaf type Leaf shape Leaf base shape Leaf margin Leaf margin posture Leaf lobation Leaf compoundness Leaf divisions Leaf lamina posture (leaf 3-dimensionality Leaf lamina posture (leaf 3-dimensional shape
has broader	leaf morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011304

leaf position trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011402
description	A leaf morphology trait [TO:0000748] which refers to the position [PATO:0000140] of leaves [PO:0025034] relative to the stem [PO:0009047].
has narrower	Leaf phyllotaxis Leaf arrangement Leaf axil angle Leaf inclination angle Leaf pendulousness
has broader	leaf morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011402

portion of plant tissue morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011502
description	A plant structure morphology trait [TO:0000839] which is a quality of a portion of plant tissue [PO:0009007].
has narrower	Cuticle thickness on the lower leaf surface Cuticle thickness on the upper leaf surface Leaf epidermis thickness Lower leaf side epidermis thickness Upper leaf side epidermis thickness Lower leaf side hypodermis thickness Upper leaf side hypodermis thickness Lower palisade mesophyll thickness Upper palisade mesophyll thickness Number of layers of palisade cells Spongy mesophyll cell thickness
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011502

plant cell morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011503
description	A plant structure morphology trait [TO:0000839] which is a quality of a plant cell [PO:0009002].
has narrower	Average leaf epidermal cell density Lower leaf side epidermal cell density Upper leaf side epidermal cell density Palisade cell length Palisade cell width Cell cross-sectional area
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011503

leaf stomatal complex morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011602
description	A vascular leaf morphology trait [TO:0000419] which is a quality of the stomatal complexes [PO:0002000] present on the leaf surface.
has narrower	Stomatal density on the lower leaf surface Stomatal density on the upper leaf surface Stomatal density averaged across both leaf surfaces Stomatal distribution Stomatal hairiness Guard cell length
has broader	leaf morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011602

leaf optical properties trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011702
description	Leaf morphology trait [TO:0000748] associated with the leafs optical [NCIT:C16940] properties, including the absorption [SWEET:soprocw:Absorption], transmission [SWEET:soprocw:Transmission], and reflection [SWEET:soprocw:Reflection] of light [OMIT:0009112].
has narrower	Leaf visible light transmission Leaf visible light absorption Leaf visible light reflection Leaf infra-red light reflection
has broader	leaf morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011702

stem morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011802
description	A shoot axis morphology trait [TO:0000739] which is the quality of a stem [PO:0009047].
has narrower	stem mass trait Stem diameter at breast height Stem count Stem length Stem cross-sectional area Wood cross-sectional area Terminal twig cross-sectional area Terminal twig length Wood density Herbaceous stem density Stem hairiness Huber value Root to shoot ratio Storage organ length Storage organ diameter Stem xylem vessel density Stem xylem vessel diameter Stem xylem vessel hydraulic mean diameter Stem xylem vessel lumen fraction Stem xylem vessel multiple fraction Stem non-lumen fraction xylem vessel wall fraction Xylem vulnerability index Stem xylem vessel wall thickness Stem xylem vessel length Wood axial parenchyma fraction Wood conduit fraction Wood fibre fraction Wood ray parenchyma fraction Wood tracheid fraction Stem modulus of elasticity Xylem modulus of elasticity Modulus of rupture Plant growth form Plant photosynthetic organ Woodiness Detailed woodiness categories Physical defence structures Plant climbing mechanisms Plant succulence Stem growth habit
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011802

stem mass trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0011902
description	A stem morphology trait [TO:0000361] which is the mass [PATO:0000125] of a stem [PO:0009047].
has narrower	Leaf dry mass to stem dry mass ratio Stem dry mass to vegetative shoot dry mass ratio (support fraction Side branch dry mass to whole plant dry mass ratio Stem dry matter content (SDMC Stem water content per unit saturated stem mass Stem mass fraction Stem saturated water content per unit stem dry mass
has broader	stem morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0011902

bark morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012002
description	A plant structure morphology trait [TO:0000736] which is a quality of bark [PO:0004518].
has narrower	bark mass trait Bark morphology, Eucalyptus Bark thickness Scaled bark thickness Bark density Bark dry mass per unit bark surface area Bark water content per unit bark dry mass Bark water content per unit saturated bark mass Bark modulus of elasticity Physical defence structures
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012002

bark mass trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012006
description	A bark morphology trait which is the mass of bark.
has broader	bark morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012006

root system morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012102
description	A collective plant organ structure morphology trait [TO:0000842] which is a quality of a root system [PO:0025025].
has narrower	root morphology trait root mass trait root branchiness trait
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012102

root morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012104
description	A plant axis morphology trait [TO:0000738] which is a quality of a root [PO:0025025].
has narrower	Root diameter Root system morphology Fine root volume to coarse root volume ratio Root biomass depth distribution coefficient Root system type (presence of taproot Specific root length (SRL Specific tap root length (STRL Root surface area per unit root dry mass (specific root area Root wood density Root to shoot ratio Root dry matter content (RDMC Root mass fraction Root vascular anatomy Storage organ length Storage organ diameter Plant nitrogen fixation capacity Plant root structures
has broader	root system morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012104

root mass trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012106
description	A root morphology trait [TO:0000043] which is the weight of a root [PO:0009005].
has broader	root system morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012106

root branchiness trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012108
description	A root system morphology trait [TO:0000985] which is the branchiness [PATO:0002009] of the root system [PO:0025025].
has broader	root system morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012108

reproductive shoot system morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012202
description	A shoot system morphology trait [TO:0000077] that is a quality of a reproductive shoot system [PO:0025082].
has narrower	floral organ morphology trait fruit morphology trait seed morphology trait Seed accessory cost fraction Seed accessory cost mass Life form
has broader	plant structure morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012202

floral organ morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012404
description	A plant organ morphology trait [TO:0000736] which is a quality of a floral organ [PO:0025395].
has narrower	perianth morphology trait androecium morphology trait gynoecium morphology trait Bud length Bud width Buds per inflorescence Maximum flower number Floral structural sex Pollination syndrome Pollination system
has broader	reproductive shoot system morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012404

perianth morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012406
description	A collective phyllome structure morphology trait [TO:0000845] which is a quality of a perianth [PO:0009058].
has narrower	Inflorescence length Inflorescence diameter Flower colour Perianth colour Flower length Flower diameter Floral orientation Perianth differentiation Fusion of perianth Number of perianth parts in each whorl (Perianth merism Number of perianth parts Perianth phyllotaxis Symmetry of perianth Number of perianth whorls Petal length
has broader	floral organ morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012406

androecium morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012408
description	A collective phyllome structure morphology trait [TO:0000845] that is a quality of the androecium [PO:0009061].
has narrower	Number of androecium parts in each whorl (Androecium structural merism Androecium structural phyllotaxis Number of androecium structural whorls Anther attachment Connective extension (apical Anther dehiscence Anther orientation Number of fertile stamens Filament presence and shape Fusion of filaments Fusion of filaments to inner perianth series Pollen grain aperture shape Number of pollen grain apertures Pollen grain length
has broader	floral organ morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012408

gynoecium morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012409
description	A collective phyllome structure morphology trait [TO:0000845] that is a quality of the gynoecium [PO:0009062].
has narrower	Gynoecium phyllotaxis Placentation Fusion of ovaries Ovary position Number of ovules per functional carpel Number of structural carpels Style differentiation Fusion of styles
has broader	floral organ morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012409

fruit morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012504
description	A multi-tissue plant structure morphology trait [TO:0000836] which is a quality of a fruit [PO:0009001].
has narrower	Fruit dry mass Fruit length Fruit width Fruit breadth Fruit wall thickness Fruit type Fruit fleshiness Fruit dehiscence Fruit colour Fruit surface hairiness Diaspore fleshiness Dispersal appendage Dispersal unit Diaspore dispersal syndrome
has broader	reproductive shoot system morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012504

seed morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0012604
description	A multi-tissue plant structure morphology trait [TO:0000836] which is a quality of a seed [PO:0009010].
has narrower	Seed dry mass Diaspore dry mass Seed embryo and endosperm dry mass Seed length Seed width Seed height Seed volume Seed count Seed shape Seed surface hairs Seed surface texture Seed surface reflectivity Diaspore fleshiness Dispersal appendage Dispersal unit Seed colour Diaspore dispersal syndrome
has broader	reproductive shoot system morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0012604

vascular tissue morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0013001
description	A portion of plant tissue morphology trait [TO:0000843] which is a quality of a vascular tissue and its parts.
has narrower	leaf vein morphology trait xylem vessel morphology trait Wood axial parenchyma fraction Wood conduit fraction Wood fibre fraction Wood ray parenchyma fraction Wood tracheid fraction
has broader	plant morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0013001

leaf vein morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0013006
description	A vascular leaf morphology trait [TO:0000419] which is a quality of a leaf lamina vein [PO:0020138].
has narrower	Leaf secondary vein angle Major leaf vein density Length of all minor and major leaf lamina veins per unit area Leaf vein frequency
has broader	vascular tissue morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0013006

xylem vessel morphology trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0013102
description	A vascular tissue morphology trait [TO:0000470] which is a quality of a xylem vessel [PO:0025417].
has narrower	Stem xylem vessel density Leaf xylem vessel density Stem xylem vessel diameter Stem xylem vessel hydraulic mean diameter Leaf xylem vessel diameter Stem xylem vessel lumen fraction Stem xylem vessel multiple fraction Stem non-lumen fraction xylem vessel wall fraction Xylem vulnerability index Leaf xylem vessel wall thickness Stem xylem vessel wall thickness Leaf xylem vessel length Stem xylem vessel length
has broader	vascular tissue morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0013102

plant structure strength trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0014001
description	A plant structure morphology trait [TO:0000736] pertaining to the strength [PATO:0001230] of a plant structure [PO:0009011] and often indicating the amount of work [PATO:0001026] required to break or deform the plant structure.
has narrower	Leaf work to punch Leaf specific work to punch Leaf work to shear Leaf specific work to shear (fracture toughness Leaf work to tear Leaf specific work to tear Bark modulus of elasticity Stem modulus of elasticity Xylem modulus of elasticity Modulus of rupture
has broader	plant morphology trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0014001

biological process trait, physiological process trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020001
description	Refers to process quality [PATO:0001236]: A quality which inheres in an process. A plant trait [TO:0000387] which is a process quality [PATO:0001236] of a biological process [GO:0008150] which occurs in a plant anatomical entity [PO:0025131], a cellular component [GO:0005575], or during a plant structure development stage [PO:0009012].
has narrower	photosynthetic trait water transport trait, hydraulic trait nutrient recycling trait
has broader	plant trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020001

photosynthetic trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020202
description	A plant physiological process trait [TO:0000283] associated with photosynthesis [GO:0015979].
has narrower	gas exchange trait photosystem performance trait Plant photosynthetic pathway Plant photosynthetic organ
has broader	biological process trait, physiological process trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020202

gas exchange trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020204
description	A plant photosynthesis trait pertaining to the movement [NCIT:C37994] of carbon dioxide [CHEBI:16526], water [CHEBI:15377], and oxygen [CHEBI:15379] into and out of a plant cellular metabolic processes [GO:0044237] where these molecules are the reactant [SIO:000879] or product [SIO:010363].
has narrower	photosynthetic rate trait respiration rate trait transpiration rate trait carbon dioxide concentration trait Leaf stomatal conductance to water vapour per unit leaf area under ambient conditions (gsw Leaf stomatal conductance to water vapour per unit leaf area during Amax measurement (gsw Leaf stomatal conductance to water vapour per unit leaf area during Asat measurement (gsw Leaf stomatal water vapour resistance under ambient conditions Leaf mesophyll conductance to carbon dioxide per unit leaf area (gm Leaf mesophyll conductance to carbon dioxide per unit leaf mass (gm Leaf cuticular conductance Leaf conductance to surface water (ksurf Leaf foliar water uptake Leaf photosynthetic nitrogen use efficiency during Amax measurement (PNUE Leaf photosynthetic nitrogen use efficiency during Asat measurement (PNUE Leaf photosynthetic phosphorus use efficiency during Amax measurement PPUE Leaf photosynthetic phosphorus use efficiency during Asat measurement (PPUE Integrated water use efficiency Intrinsic water use efficiency (WUEi Instantaneous water use efficiency (WUE
has broader	photosynthetic trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020204

photosynthetic rate trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020206
description	The total amount of carbon fixed by the plant per unit time and is measured by determining the changes in CO2 concentration over a plant leaf in a glass enclosure.
has narrower	Bark photosynthesis Leaf photosynthesis rate per unit leaf area under ambient light and CO2 (A Leaf photosynthesis rate per unit leaf area under saturating light and CO2 (Amax Leaf photosynthesis rate per unit leaf area under saturating light and ambient CO2 (Asat Leaf photosynthesis rate per unit leaf dry mass under ambient light and CO2 (A Leaf photosynthesis rate per unit leaf dry mass under saturating light and CO2 (Amax Leaf photosynthesis rate per unit leaf dry mass under saturating light and ambient CO2 (Asat
has broader	gas exchange trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020206

respiration rate trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020208
description	A plant gas exchange trait which is the net rate of carbon dioxide [CHEBI:16526] released from a whole plant [PO:0000003] or plant organ [PO:0009008] due to cellular respiration [GO:0045333] and photorespiration [GO:0009853].
has narrower	Leaf respiration rate per unit leaf area, in the dark (Rdark Leaf respiration rate per unit leaf dry mass, in the dark (Rdark Leaf respiration rate per unit leaf area, in the light (Rday Stem respiration rate per unit stem area, in the dark
has broader	gas exchange trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020208

transpiration rate trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020210
description	A plant gas exchange trait which is the rate of transpiration [ENVO:02500035].
has narrower	Leaf transpiration per unit leaf area under ambient conditions (E Leaf transpiration per unit leaf area during Amax measurement (E Leaf transpiration per unit leaf area during Asat measurement (E Leaf transpiration rate per unit leaf area, in the dark Integrated plant transpiration Whole plant sapflow
has broader	gas exchange trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020210

carbon dioxide concentration trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020212
description	A plant gas exchange trait which is the concentration [PATO:0000033] of carbon dioxide [CHEBI:16526].
has narrower	Leaf internal CO2 concentration during Amax measurement (ci Internal CO2 concentration during Asat measurement (ci Internal CO2 concentration under ambient conditions (ci Ratio of internal to external CO2 concentrations (ci/ca CO2 concentration inside chloroplasts (cc Ambient CO2 concentration (ca
has broader	gas exchange trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020212

photosystem performance trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0020812
description	A plant photosynthesis trait pertaining to the performance [NCIT:C60778] of the photosystem [GO:0009521] complexes, especially with regard to how effectively they process photosynthetically active radiation [ENVO:01001864] during the photosynthesis light reactions [GO:0019684].
has narrower	Leaf Jmax per unit leaf area (Jmax Leaf Jmax per unit leaf area at 25 deg C (Jmax25 Leaf Jmax per unit leaf mass (Jmax Leaf Vcmax per unit leaf area (Vcmax Leaf Vcmax per unit leaf area at 25 deg C (Vcmax25 Leaf Vcmax per unit leaf mass (Vcmax Leaf Jmax to leaf Vcmax ratio at 25 deg C Leaf maximum quantum yield (Fv/Fm Leaf ambient quantum yield Leaf quantum yield, gas exchange measurement Leaf heat tolerance maximum critical temperature (Tcrit-cold or Tcrit-min Leaf heat tolerance minimum critical temperature (Tcrit-hot or Tcrit-max Leaf heat tolerance maximum temperature (Tmax-hot Photochemical reflectance index (PRI
has broader	photosynthetic trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0020812

water transport trait, hydraulic trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0021002
description	A plant physiological process trait [TO:0000283] pertaining to the movement [NCIT:C37994] of water [CHEBI:15377] through a plants vascular system [PO:0000034], including water stress traits [TO:0000237].
has narrower	Leaf secondary vein angle Major leaf vein density Length of all minor and major leaf lamina veins per unit area Leaf vein frequency Stem xylem vessel density Leaf xylem vessel density Stem xylem vessel diameter Stem xylem vessel hydraulic mean diameter Leaf xylem vessel diameter Stem xylem vessel lumen fraction Stem xylem vessel multiple fraction Stem non-lumen fraction xylem vessel wall fraction Xylem vulnerability index Leaf xylem vessel wall thickness Stem xylem vessel wall thickness Leaf xylem vessel length Stem xylem vessel length Wood axial parenchyma fraction Wood conduit fraction Wood fibre fraction Wood ray parenchyma fraction Wood tracheid fraction Water band index Stem hydraulic conductivity (Kh Sapwood specific hydraulic conductivity (Ks Theoretical sapwood specific hydraulic conductivity (Ks Stem specific hydraulic conductivity (Ks Leaf specific hydraulic conductance (kleaf Leaf specific hydraulic conductivity (Kl Root hydraulic conductivity (Kh Root specific hydraulic conductivity Pre-dawn water potential Midday water potential Stem sapwood capacitance (C Leaf capacitance (Cleaf Root sapwood capacitance (C Leaf xylem pressure, 50% lost conductance (P50) (leaf hydraulic vulnerability Stem xylem pressure, 12% lost conductivity (P12 Stem xylem pressure, 50% lost conductivity (P50 Stem xylem pressure, 88% lost conductivity (P88 Hydraulic safety margin, 50 Hydraulic safety margin, 88 Leaf turgor loss point Osmotic potential Osmotic potential at full tugor Bulk modulus of elasticity (e Leaf relative water content predawn Leaf relative water content at turgor loss point Root xylem pressure, 50% lost conductivity (P50 Root xylem pressure, 12% lost conductivity (P12 Root xylem pressure, 88% lost conductivity (P88 Leaf xylem pressure, 12% lost conductance (P12 Leaf xylem pressure, 88% lost conductance (P88
has broader	biological process trait, physiological process trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0021002

nutrient recycling trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0022002
description	A plant physiological process trait [TO:0000283] associated with the resorption [NCIT:C93204] and recycling [NCIT:C126395] of macronutrients [CHEBI:33937] and micronutrients [CHEBI:27027] as plant structures [PO:0009011] senesce [PATO:0001487].
has narrower	Leaf nitrogen resorption efficiency Leaf phosphorus resorption efficiency
has broader	biological process trait, physiological process trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0022002

life history trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030001
description	A plant trait [TO:0000387] pertaining to an individuals life history [ECOCORE:00000018].
has narrower	whole plant phenotype trait plant phenological trait interspecific interactions trait genetic structure trait reproductive structure life history trait fire response trait chemical stress sensitivity trait environmental tolerance trait Ephemeral life history class Plant growth substrate Clonal spread mechanism Storage organ Plant vegetative response to disturbances other than fire
has broader	plant trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030001

whole plant phenotype trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030006
description	A plant life history trait which is a whole plant phenotype [FLOPO:0000089].
has narrower	Plant spinescence Plant growth form Life form Life history Lifespan Plant alternative energy and nutrient acquisition strategies Woodiness Detailed woodiness categories Plant climbing mechanisms Plant succulence Stem growth habit Competitive stratum Vegetative reproduction ability Bud bank location Time from fire to fruiting Time from fire until 50% of individuals are fruiting
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030006

plant phenological trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030008
description	A growth and development trait [TO:0000357] which is the timing of the occurrence of plant structure development stages [PO:0009012] and the relevant biological processes [GO:0008150].
has narrower	Leaf phenology Leaf lifespan Foliage time Age of reproductive maturity Environmental flowering cues Flowering time, by month Fruiting time, by month Seedling recruitment time, by month Seedling establishment conditions Canopy light environment required for reproduction Canopy light environment required for seedling establishment Seed storage location Post-fire flowering Time from fire to first flowering Time from fire until 50% of individuals are flowering Time from fire to peak flowering Time from fire until flowering declines
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030008

interspecific interactions trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030009
description	A plant life history trait that captures an interspecific interaction between organisms [GO:0044419].
has narrower	Dispersal appendage Plant nitrogen fixation capacity Plant root structures Plant parasitism status Pollination syndrome Pollination system Diaspore dispersal syndrome Diaspore dispersal agents
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030009

genetic structure trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030059
description	A plant life history trait that captures information about genetic structures.
has narrower	Plant genome size Chromosome ploidy
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030059

reproductive structure life history trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030504
description	A plant life history trait pertaining to a plants reproductive structures [UBERON:0005156].
has narrower	Seed storage location Serotiny Seedbank longevity class Seedbank longevity Dormancy type Seed germination treatment Seed germination proportion Seed viability Seed germination time Time from fire to first flowering Time from fire until 50% of individuals are flowering Time from fire to peak flowering Time from fire until flowering declines Time from fire to fruiting Time from fire until 50% of individuals are fruiting
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030504

fire response trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030602
description	A plant life history trait indicating adaptations [NCIT:C158868] to increase survival rate [EnvThes:21065] or seed germination [GO_0009845] rate following fire [ENVO:01000786] or indicating a plants response [NCIT:C25755] to fire.
has narrower	Leaf ash content per unit leaf dry mass Bark ash content per unit bark dry mass Ephemeral life history class Seedling recruitment time, by month Seedling establishment conditions Seed storage location Serotiny Seedbank longevity class Seedbank longevity Sprout depth Post-fire resprouting capacity Post-fire proportion resprouting individuals Post-fire resprouting capacity of juvenile plants Time from seedling germination until individuals survive a fire Post-fire to pre-fire stem ratio Fire exposure level Post-fire recruitment Post-fire flowering Time from fire to first flowering Time from fire until 50% of individuals are flowering Time from fire to peak flowering Time from fire until flowering declines Time from fire to fruiting Time from fire until 50% of individuals are fruiting Fuel bed bulk density Fuel consumption by fire Fire rate of spread Leaf smoulder duration Leaf flame duration Leaf flame and smoulder duration Fire time to ignition Plant fire tolerance strategy
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030602

chemical stress sensitivity trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030702
description	Response by the plant in terms of resistivity or senility to chemical stress.
has narrower	Plant soil salinity tolerance Plant salt tolerance strategy Plant calcium sensitivity
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030702

environmental tolerance trait

Property	Value
URI	https://w3id.org/APD/traits/trait_group_0030704
description	A plant life history trait indicating adaptations [NCIT:C158868] to specific environmental conditions [ENVO:01000203] or tolerance [NCIT:C43414] to specific environmental stresses [EnvThes:21245].
has narrower	Leaf heat tolerance maximum critical temperature (Tcrit-cold or Tcrit-min Leaf heat tolerance minimum critical temperature (Tcrit-hot or Tcrit-max Leaf heat tolerance maximum temperature (Tmax-hot Leaf ice nucleation temperature Photochemical reflectance index (PRI Water band index Modified normalized difference vegetation index (modified NDVI Modified chlorophyll absorption ratio index 705 Leaf chlorophyll content in SPAD units Plant succulence Foliage time Environmental flowering cues Seedling recruitment time, by month Seedling establishment conditions Plant resource requirements and tolerance Plant flood regime response Plant water-logging tolerance Plant inundation tolerance Plant snow tolerance Plant soil salinity tolerance Plant salt tolerance strategy Plant calcium sensitivity
has broader	life history trait
is in scheme	https://w3id.org/APD/traits
identifier	trait_group_0030704

3. Trait concepts

The APD Trait concepts express the traits defined by the APD.

Leaf aluminium (Al) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000012
alternative label	leaf_Al_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of aluminium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0.01
maxAllowedValue	12000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	aluminium atom
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf aluminium content per leaf dry mass [TOP449] (https://top-thesaurus.org/index) exact match: Leaf aluminium (Al) content per leaf dry mass [TRY:249] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000012

Leaf boron (B) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000014
alternative label	leaf_B_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of boron in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0.01
maxAllowedValue	1000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	boron atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf boron content per leaf dry mass [TOP450] (https://top-thesaurus.org/index) exact match: Leaf boron (B) content per leaf dry mass [TRY:250] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000014

Leaf carbon (C) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000016
alternative label	leaf_C_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of carbon in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	50
maxAllowedValue	750
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon atom
scope note	none
has exact match	leaf carbon content Leaf carbon content
has close match
has related match
examples	exact match: Leaf carbon content per leaf dry mass [TOP452] (https://top-thesaurus.org/index) exact match: Leaf carbon (C) content per leaf dry mass [TRY:13] (https://www.try-db.org/de/de.php) exact match: leaf carbon content per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000016

Leaf calcium (Ca) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000018
alternative label	leaf_Ca_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of calcium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	1000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf calcium content per leaf dry mass [TOP451] (https://top-thesaurus.org/index) exact match: Leaf calcium (Ca) content per leaf dry mass [TRY:252] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000018

Leaf chlorine (Cl) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000020
alternative label	leaf_Cl_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorine [CHEBI:23116] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of chlorine in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	10
maxAllowedValue	10000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	chlorine atom
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf chlorine content per leaf dry mass [TOP453] (https://top-thesaurus.org/index) exact match: Leaf chlorine (Cl) content per leaf dry mass [TRY:539] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000020

Leaf chromium (Cr) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000022
alternative label	leaf_Cr_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chromium [CHEBI:28073] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of chromium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	ppm parts per million
minAllowedValue	0.01
maxAllowedValue	150
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	chromium atom
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf chromium (Cr) content per leaf dry mass [TRY:254] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000022

Leaf cobalt (Co) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000024
alternative label	leaf_Co_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of cobalt [CHEBI:27638] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of cobalt in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	ppm parts per million
minAllowedValue	0.01
maxAllowedValue	150
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	cobalt atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf cobalt content per leaf dry mass [TOP454] (https://top-thesaurus.org/index) exact match: Leaf cobalt (Co) content per leaf dry mass [TRY:253] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000024

Leaf copper (Cu) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000026
alternative label	leaf_Cu_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of copper in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0.01
maxAllowedValue	150
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	copper atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf copper content per leaf dry mass [TOP455] (https://top-thesaurus.org/index) exact match: Leaf copper (Cu) content per leaf dry mass [TRY:255] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000026

Leaf iron (Fe) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000028
alternative label	leaf_Fe_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of iron in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	1
maxAllowedValue	10000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	iron atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf iron content per leaf dry mass [TOP456] (https://top-thesaurus.org/index) exact match: Leaf iron (Fe) content per leaf dry mass [TRY:256] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000028

Leaf potassium (K) content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0000029
alternative label	leaf_K_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of potassium in a leaf to leaf area.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	10
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	potassium atom macronutrient
scope note	none
has exact match	leaf potassium content
has close match
has related match
examples	exact match: Leaf potassium content per leaf area [TOP483] (https://top-thesaurus.org/index) exact match: Leaf potassium (K) content per leaf area [TRY:52] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000029

Leaf potassium (K) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000030
alternative label	leaf_K_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of potassium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom macronutrient
scope note	none
has exact match	leaf potassium content
has close match
has related match
examples	exact match: Leaf potassium content per leaf dry mass [TOP464] (https://top-thesaurus.org/index) exact match: Leaf potassium (K) content per leaf dry mass [TRY:44] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000030

Leaf magnesium (Mg) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000032
alternative label	leaf_Mg_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of magnesium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf magnesium content per leaf dry mass [TOP457] (https://top-thesaurus.org/index) exact match: Leaf magnesium (Mg) content per leaf dry mass [TRY:257] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000032

Leaf manganese (Mn) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000034
alternative label	leaf_Mn_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of manganese in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	1
maxAllowedValue	20000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	manganese atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf manganese content per leaf dry mass [TOP458] (https://top-thesaurus.org/index) exact match: Leaf manganese (Mn) content per leaf dry mass [TRY:258] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000034

Leaf molybdenum (Mo) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000036
alternative label	leaf_Mo_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of molybdenum [CHEBI:28685] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of molybdenum in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0
maxAllowedValue	100
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	molybdenum atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf molybdenum content per leaf dry mass [TOP459] (https://top-thesaurus.org/index) exact match: Leaf molybdenum (Mo) content per leaf dry mass [TRY:259] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000036

Leaf nitrogen (N) content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0000037
alternative label	leaf_N_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of nitrogen in a leaf to leaf area.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.08
maxAllowedValue	50
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	nitrogen atom macronutrient
scope note	none
has exact match	Leaf nitrogen content leaf nitrogen content leaf nitrogen concentration
has close match
has related match
examples	exact match: Leaf nitrogen content per leaf area [TOP481] (https://top-thesaurus.org/index) exact match: Leaf nitrogen (N) content per leaf area [TRY:50] (https://www.try-db.org/de/de.php) exact match: leaf nitrogen content per leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000037

Leaf nitrogen (N) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000038
alternative label	leaf_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of nitrogen in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.4
maxAllowedValue	100
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient
scope note	none
has exact match	Leaf nitrogen content leaf nitrogen content leaf nitrogen concentration
has close match
has related match
examples	exact match: Leaf nitrogen content per leaf dry mass [TOP462] (https://top-thesaurus.org/index) exact match: Leaf nitrogen (N) content per leaf dry mass [TRY:14] (https://www.try-db.org/de/de.php) exact match: leaf nitrogen content per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Mass-based leaf nitrogen content [BROT:7] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000038

Leaf sodium (Na) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000040
alternative label	leaf_Na_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of sodium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	1
maxAllowedValue	20000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	sodium atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf sodium content per leaf dry mass [TOP460] (https://top-thesaurus.org/index) exact match: Leaf sodium (Na) content per leaf dry mass [TRY:260] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000040

Leaf nickel (Ni) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000042
alternative label	leaf_Ni_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nickel [CHEBI:28112] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of nickel in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0.1
maxAllowedValue	100
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nickel atom
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf nickel content per leaf dry mass [TOP461] (https://top-thesaurus.org/index) exact match: Leaf nickel (Ni) content per leaf dry mass [TRY:261] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000042

Leaf phosphorus (P) content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0000043
alternative label	leaf_P_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of phosphorus in a leaf to leaf area.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.005
maxAllowedValue	1.2
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	phosphorus atom macronutrient
scope note	none
has exact match	leaf phosphorus content
has close match
has related match
examples	exact match: Leaf phosphorus content per leaf area [TOP482] (https://top-thesaurus.org/index) exact match: Leaf phosphorus (P) content per leaf area [TRY:51] (https://www.try-db.org/de/de.php) exact match: leaf phosphorus content per leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000043

Leaf phosphorus (P) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000044
alternative label	leaf_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of phosphorus in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.03
maxAllowedValue	12
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom macronutrient
scope note	none
has exact match	leaf phosphorus content
has close match
has related match
examples	exact match: Leaf phosphorus content per leaf dry mass [TOP463] (https://top-thesaurus.org/index) exact match: Leaf phosphorus (P) content per leaf dry mass [TRY:15] (https://www.try-db.org/de/de.php) exact match: leaf phosphorus content per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000044

Leaf sulphur (S) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000046
alternative label	leaf_S_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of sulphur in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	50
maxAllowedValue	50000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	sulfur atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf sulfur content per leaf dry mass [TOP466] (https://top-thesaurus.org/index) exact match: Leaf sulfur (S) content per leaf dry mass [TRY:262] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000046

Leaf selenium (Se) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000048
alternative label	leaf_Se_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of selenium [CHEBI:27568] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of selenium in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	ppm parts per million
minAllowedValue	0.01
maxAllowedValue	100
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	selenium atom
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf selenium (Se) content per leaf dry mass [TRY:263] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000048

Leaf silicon (Si) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000050
alternative label	leaf_Si_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of silicon [CHEBI:27573] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of silicon in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades. Leaf silicon (Si) must not be confused with leaf silica (SiO2). In plants, Si is mostly found as hydrated amorphous silica (SiO2 , nH2O).
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	200
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	silicon atom
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf silicon content per leaf dry mass [TOP465] (https://top-thesaurus.org/index) exact match: Leaf silicon (Si) content per leaf dry mass [TRY:264] (https://www.try-db.org/de/de.php)
references	Cooke & Leishman 2011 de Tombeur 2023
date created	14/07/2021
date modified	05/02/2024
date reviewed	13/02/2023
reviewed by	Félix de Tombeur
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000050

Leaf zinc (Zn) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000052
alternative label	leaf_Zn_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of zinc in a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0.1
maxAllowedValue	1000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	zinc atom micronutrient
scope note	none
has exact match
has close match
has related match
examples	exact match: Leaf zinc content per leaf dry mass [TOP467] (https://top-thesaurus.org/index) exact match: Leaf zinc (Zn) content per leaf dry mass [TRY:268] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000052

Leaf carbon to nitrogen ratio (C/N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0000090
alternative label	leaf_CN_ratio
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a leaf [PO:0025034] to the mass of nitrogen [CHEBI:25555] in the leaf. The ratio of the mass of carbon in a leaf to the mass of nitrogen in the leaf.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	10
maxAllowedValue	1000
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass dimensionless ratio ratio; proportion
keyword	carbon atom macronutrient nitrogen atom
scope note	none
has exact match	Ratio leaf nitrogen to leaf carbon C/N
has close match
has related match
examples	exact match: Leaf carbon/nitrogen (C/N) ratio [TRY:146] (https://www.try-db.org/de/de.php) exact match: leaf carbon content per leaf nitrogen content [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000090

Leaf nitrogen to phosphorus ratio (N/P) per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000091
alternative label	leaf_NP_ratio
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a leaf [PO:0025034] to the mass of phosphorus [CHEBI:28659] in the leaf. The ratio of the mass of nitrogen in a leaf to the mass of phosphorus in the leaf.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	1
maxAllowedValue	100
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass dimensionless ratio ratio; proportion
keyword	nitrogen atom macronutrient phosphorus atom
scope note
has exact match
has close match
has related match
examples	exact match: Leaf nitrogen/phosphorus (N/P) ratio [TRY:56] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	N_to_P_ratio
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000091

Senesced leaf aluminium (Al) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000112
alternative label	leaf_senesced_Al_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of aluminium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	aluminium atom senescent plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter aluminium content per litter dry mass [TOP583] (https://top-thesaurus.org/index) exact match: Litter aluminium (Al ) content per litter dry mass [TRY:987] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Al_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000112

Senesced leaf boron (B) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000114
alternative label	leaf_senesced_B_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of boron in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	boron atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter boron content per litter dry mass [TOP584] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_B_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000114

Senesced leaf carbon (C) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000116
alternative label	leaf_senesced_C_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of carbon in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.1
maxAllowedValue	1
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon atom senescent plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter carbon content per litter dry mass [TOP586] (https://top-thesaurus.org/index) exact match: Litter carbon (C) content per litter dry mass [TRY:72] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_C_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000116

Senesced leaf calcium (Ca) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000118
alternative label	leaf_senesced_Ca_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of calcium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	1000
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter calcium content per litter dry mass [TOP585] (https://top-thesaurus.org/index) exact match: Litter (leaf) calcium (Ca) content per leaf litter dry mass [TRY:943] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Ca_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000118

Senesced leaf copper (Cu) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000126
alternative label	leaf_senesced_Cu_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of copper in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0.01
maxAllowedValue	100
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	copper atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter copper content per litter dry mass [TOP589] (https://top-thesaurus.org/index) exact match: Litter (leaf) copper (Cu) content per leaf litter dry mass [TRY:947] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Cu_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000126

Senesced leaf iron (Fe) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000128
alternative label	leaf_senesced_Fe_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of iron in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	1
maxAllowedValue	10000
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	iron atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter iron content per litter dry mass [TOP590] (https://top-thesaurus.org/index) exact match: Litter (leaf) iron (Fe) content per leaf litter dry mass [TRY:950] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Fe_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000128

Senesced leaf potassium (K) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000130
alternative label	leaf_senesced_K_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of potassium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom senescent macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter potassium content per litter dry mass [TOP598] (https://top-thesaurus.org/index) exact match: Litter (leaf) potassium (K) content per leaf litter dry mass [TRY:958] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_K_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000130

Senesced leaf magnesium (Mg) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000132
alternative label	leaf_senesced_Mg_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of magnesium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	100
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter magnesium content per litter dry mass [TOP591] (https://top-thesaurus.org/index) exact match: Litter (leaf) magnesium (Mg) content per leaf litter dry mass [TRY:952] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Mg_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000132

Senesced leaf manganese (Mn) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000134
alternative label	leaf_senesced_Mn_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of manganese in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	1
maxAllowedValue	10000
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	manganese atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter manganese content per litter dry mass [TOP592] (https://top-thesaurus.org/index) exact match: Litter (leaf) manganese (Mn) content per leaf litter dry mass [TRY:953] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Mn_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000134

Senesced leaf molybdenum (Mo) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000136
alternative label	leaf_senesced_Mo_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of molybdenum [CHEBI:28685] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of molybdenum in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0
maxAllowedValue	30
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	molybdenum atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter molybdenum content per litter dry mass [TOP593] (https://top-thesaurus.org/index) exact match: Litter molybdenium (Mo) content per litter dry mass [TRY:999] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Mo_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000136

Senesced leaf nitrogen (N) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000138
alternative label	leaf_senesced_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of nitrogen in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	100
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom senescent macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter nitrogen content per litter dry mass [TOP596] (https://top-thesaurus.org/index) exact match: Litter (leaf) nitrogen (N) content per leaf litter dry mass [TRY:954] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_N_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000138

Senesced leaf sodium (Na) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000140
alternative label	leaf_senesced_Na_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of sodium in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	10
maxAllowedValue	20000
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	sodium atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter sodium content per litter dry mass [TOP594] (https://top-thesaurus.org/index) exact match: Litter sodium (Na) content per litter dry mass [TRY:1004] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Na_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000140

Senesced leaf nickel (Ni) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000142
alternative label	leaf_senesced_Ni_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nickel [CHEBI:28112] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of nickel in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	10
maxAllowedValue	1e+05
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nickel atom senescent plant litter
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Ni_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000142

Senesced leaf phosphorus (P) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000144
alternative label	leaf_senesced_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of phosphorus in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.005
maxAllowedValue	15
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom senescent macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter phosphorus content per litter dry mass [TOP597] (https://top-thesaurus.org/index) exact match: Litter (leaf) phosphorus (P) content per leaf litter dry mass [TRY:957] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_P_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000144

Senesced leaf sulphur (S) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000146
alternative label	leaf_senesced_S_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of sulphur in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	100
maxAllowedValue	50000
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	sulfur atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter sulfur content per litter dry mass [TOP600] (https://top-thesaurus.org/index) exact match: Litter (leaf) sulfur (S) content per leaf litter dry mass [TRY:961] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_S_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000146

Senesced leaf zinc (Zn) content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000152
alternative label	leaf_senesced_Zn_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in a senescent [PATO:0001487] leaf [PO:0025034] (leaf litter [ENVO:01000628]) to the senescent leaf dry mass [TO:0001014]. The ratio of the mass of zinc in a senescent leaf (leaf litter) to the senescent leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/kg milligram per kilogram
minAllowedValue	0
maxAllowedValue	100
has broader	senesced leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	zinc atom senescent micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Litter zinc content per litter dry mass [TOP601] (https://top-thesaurus.org/index) exact match: Litter (leaf) zinc (Zn) content per leaf litter dry mass [TRY:962] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	senesced_leaf_Zn_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000152

Leaf nitrogen resorption efficiency

Property	Value
URI	https://w3id.org/APD/traits/trait_0022012
alternative label	leaf_N_resorption
description	A leaf [PO:0025034] nutrient recycling trait which is one minus the ratio [PATO:0001470] of leaf nitrogen [CHEBI:25555] content in a senescent [PATO:0001487] leaf (leaf litter [ENVO:01000628]) to leaf nitrogen content in a live [PATO:0001421] leaf, indicating what proportion of nitrogen is resorbed [NCIT:C93204] from the senescent leaf before it is shed. A leaf nutrient recycling trait which is one minus the ratio of leaf nitrogen content in a senescent leaf (leaf litter) to leaf nitrogen content in a live leaf, indicating what proportion of nitrogen is resorbed from the senescent leaf before it is shed.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	-1.1
maxAllowedValue	1
has broader	nutrient recycling trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion dimensionless ratio
keyword	nitrogen atom senescent resorption plant litter macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	resorption_leaf_N
is in scheme	https://w3id.org/APD/traits
identifier	trait_0022012

Leaf phosphorus resorption efficiency

Property	Value
URI	https://w3id.org/APD/traits/trait_0022013
alternative label	leaf_P_resorption
description	A leaf [PO:0025034] nutrient recycling trait which is one minus the ratio [PATO:0001470] of leaf phosphorus [CHEBI:28659] content in a senescent [PATO:0001487] leaf (leaf litter [ENVO:01000628]) to leaf phosphorus content in a live [PATO:0001421] leaf, indicating what proportion of phosphorus is resorbed [NCIT:C93204] from the senescent leaf before it is shed. A leaf nutrient recycling trait which is one minus the ratio of leaf phosphorus content in a senescent leaf (leaf litter) to leaf phosphorus content in a live leaf, indicating what proportion of phosphorus is resorbed from the senescent leaf before it is shed.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	-1.1
maxAllowedValue	1
has broader	nutrient recycling trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion dimensionless ratio
keyword	phosphorus atom senescent resorption plant litter macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	resorption_leaf_P
is in scheme	https://w3id.org/APD/traits
identifier	trait_0022013

Stem carbon (C) content per unit stem dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000216
alternative label	stem_C_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in a stem [PO:0009047] to stem dry mass. The ratio of the mass of carbon in a stem to stem dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	100
maxAllowedValue	1000
has broader	stem mineral and ion content trait
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon atom
scope note
has exact match	stem carbon content
has close match
has related match
examples	exact match: Stem carbon content per stem dry mass [TOP491] (https://top-thesaurus.org/index) exact match: Stem carbon (C) content per stem dry mass [TRY:407] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000216

Stem nitrogen (N) content per unit stem dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000238
alternative label	stem_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in a stem [PO:0009047] to stem dry mass. The ratio of the mass of nitrogen in a stem to stem dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	1000
has broader	stem mineral and ion content trait
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Stem nitrogen content per stem dry mass [TOP501] (https://top-thesaurus.org/index) exact match: Stem nitrogen (N) content per stem dry mass [TRY:406] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000238

Wood carbon (C) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000416
alternative label	wood_C_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in wood [PO:0005848] to wood dry mass. The ratio of the mass of carbon in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	50
maxAllowedValue	750
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon atom wood plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood carbon content per wood dry mass trait [TOP732] (https://top-thesaurus.org/index) close match: Stem carbon (C) content per stem dry mass [TRY:407] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000416

Wood calcium (Ca) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000418
alternative label	wood_Ca_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in wood [PO:0005848] to wood dry mass. The ratio of the mass of calcium in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom wood micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood calcium content per wood dry mass trait [TOP731] (https://top-thesaurus.org/index) close match: Stem calcium (Ca) content per stem dry mass [TRY:623] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000418

Wood potassium (K) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000430
alternative label	wood_K_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in wood [PO:0005848] to wood dry mass. The ratio of the mass of potassium in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	100
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom wood macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood potassium content per wood dry mass trait [TOP744] (https://top-thesaurus.org/index) close match: Stem potassium (K) content per stem dry mass [TRY:628] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000430

Wood magnesium (Mg) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000432
alternative label	wood_Mg_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in wood [PO:0005848] to wood dry mass. The ratio of the mass of magnesium in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom wood micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood magnesium content per wood dry mass trait [TOP737] (https://top-thesaurus.org/index) close match: Stem magnesium (Mg) content per stem dry mass [TRY:624] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000432

Wood nitrogen (N) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000438
alternative label	wood_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in wood [PO:0005848] to wood dry mass. The ratio of the mass of nitrogen in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom wood macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood nitrogen content per wood dry mass trait [TOP742] (https://top-thesaurus.org/index) exact match: Wood nitrogen (N) content per wood dry mass [TRY:1229] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000438

Wood sodium (Na) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000440
alternative label	wood_Na_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in wood [PO:0005848] to wood dry mass. The ratio of the mass of sodium in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	sodium atom wood micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood sodium content per wood dry mass trait [TOP740] (https://top-thesaurus.org/index) close match: Stem sodium (Na) content per stem dry mass [TRY:627] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000440

Wood phosphorus (P) content per unit wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000444
alternative label	wood_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in wood [PO:0005848] to wood dry mass. The ratio of the mass of phosphorus in wood to wood dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom wood macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Wood phosphorus content per wood dry mass trait [TOP743] (https://top-thesaurus.org/index) close match: Stem phosphorus (P) content per stem dry mass [TRY:616] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000444

Dead wood calcium (Ca) content per unit dead wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000518
alternative label	wood_dead_Ca_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood. The ratio of the mass of calcium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	senesced wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom senescent wood micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Coarse woody debris calcium content per coarse woody debris dry mass trait [TOP750] (https://top-thesaurus.org/index) close match: Litter (other than leaf: twigs, bark, flowers…) calcium (Ca) content per litter dry mass [TRY:2892] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	dead_wood_Ca_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000518

Dead wood potassium (K) content per unit dead wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000530
alternative label	wood_dead_K_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood. The ratio of the mass of potassium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	senesced wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom senescent wood macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Coarse woody debris potassium content per coarse woody debris dry mass trait [TOP763] (https://top-thesaurus.org/index) close match: Litter (other than leaf: twigs, bark, flowers…) potassium (K) content per litter dry mass [TRY:2891] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	dead_wood_K_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000530

Dead wood magnesium (Mg) content per unit dead wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000532
alternative label	wood_dead_Mg_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood. The ratio of the mass of magnesium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	senesced wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom senescent wood micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Coarse woody debris magnesium content per coarse woody debris dry mass trait [TOP756] (https://top-thesaurus.org/index) close match: Litter (other than leaf: twigs, bark, flowers…) magnesium (Mg) content per litter dry mass [TRY:2893] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	dead_wood_Mg_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000532

Dead wood nitrogen (N) content per unit dead wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000538
alternative label	wood_dead_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood. The ratio of the mass of nitrogen in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	senesced wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom senescent wood macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Coarse woody debris nitrogen content per coarse woody debris dry mass trait [TOP761] (https://top-thesaurus.org/index) close match: Litter twig nitrogen (N) content per twig dry mass [TRY:1199] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	dead_wood_N_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000538

Dead wood sodium (Na) content per unit dead wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000540
alternative label	wood_dead_Na_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood. The ratio of the mass of sodium in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	1
has broader	senesced wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	sodium atom senescent wood micronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Coarse woody debris sodium content per coarse woody debris dry mass trait [TOP759] (https://top-thesaurus.org/index) close match: Litter (other than leaf: twigs, bark, flowers…) sodium (Na) content per litter dry mass [TRY:2898] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	dead_wood_Na_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000540

Dead wood phosphorus (P) content per unit dead wood dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000544
alternative label	wood_dead_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in senescent [PATO:0001487] wood [PO:0005848] (litter [ENVO:01000628]) to the dry mass of the senesced wood. The ratio of the mass of phosphorus in senescent wood (litter) to the dry mass of the senesced wood.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	senesced wood mineral and ion content trait
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom senescent wood macronutrient plant litter
scope note
has exact match
has close match
has related match
examples	exact match: Coarse woody debris phosphorus content per coarse woody debris dry mass trait [TOP762] (https://top-thesaurus.org/index) close match: Litter other than leaf phosphorus (P) content per litter dry mass [TRY:2890] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	dead_wood_P_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000544

Bark aluminium (Al) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000612
alternative label	bark_Al_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of aluminium [CHEBI:28984] in bark [PO:0004518] to bark dry mass. The ratio of the mass of aluminium in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	aluminium atom
scope note
has exact match
has close match
has related match
examples	exact match: Bark aluminium content per bark dry mass [TOP373] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000612

Bark boron (B) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000614
alternative label	bark_B_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of boron [CHEBI:27560] in bark [PO:0004518] to bark dry mass. The ratio of the mass of boron in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	1
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	boron atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark boron content per bark dry mass [TOP374] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000614

Bark carbon (C) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000616
alternative label	bark_C_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in bark [PO:0004518] to bark dry mass. The ratio of the mass of carbon in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	50
maxAllowedValue	750
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon atom
scope note
has exact match
has close match
has related match
examples	exact match: Bark carbon content per bark dry mass [TOP376] (https://top-thesaurus.org/index) exact match: Bark carbon (C) content per bark dry mass [TRY:617] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000616

Bark calcium (Ca) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000618
alternative label	bark_Ca_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in bark [PO:0004518] to bark dry mass. The ratio of the mass of calcium in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark calcium content per bark dry mass [TOP375] (https://top-thesaurus.org/index) exact match: Bark calcium (Ca) content per bark dry mass [TRY:2957] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000618

Bark copper (Cu) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000626
alternative label	bark_Cu_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of copper [CHEBI:28694] in bark [PO:0004518] to bark dry mass. The ratio of the mass of copper in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1e-04
maxAllowedValue	1
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	copper atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark copper content per bark dry mass [TOP379] (https://top-thesaurus.org/index) exact match: Bark copper (Cu) content per bark dry mass [TRY:2958] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000626

Bark iron (Fe) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000628
alternative label	bark_Fe_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of iron [CHEBI:18248] in bark [PO:0004518] to bark dry mass. The ratio of the mass of iron in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	iron atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark iron content per bark dry mass [TOP380] (https://top-thesaurus.org/index) exact match: Bark iron (Fe) content per bark dry mass [TRY:2959] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000628

Bark potassium (K) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000630
alternative label	bark_K_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in bark [PO:0004518] to bark dry mass. The ratio of the mass of potassium in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	100
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom macronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark potassium content per bark dry mass [TOP388] (https://top-thesaurus.org/index) exact match: Bark potassium (K) content per bark dry mass [TRY:2960] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000630

Bark magnesium (Mg) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000632
alternative label	bark_Mg_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in bark [PO:0004518] to bark dry mass. The ratio of the mass of magnesium in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark magnesium content per bark dry mass [TOP381] (https://top-thesaurus.org/index) exact match: Bark magnesium (Mg) content per bark dry mass [TRY:2961] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000632

Bark manganese (Mn) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000634
alternative label	bark_Mn_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of manganese [CHEBI:18291] in bark [PO:0004518] to bark dry mass. The ratio of the mass of manganese in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	1
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	manganese atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark manganese content per bark dry mass [TOP382] (https://top-thesaurus.org/index) exact match: Bark manganese (Mn) content per bark dry mass [TRY:2962] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000634

Bark nitrogen (N) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000638
alternative label	bark_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in bark [PO:0004518] to bark dry mass. The ratio of the mass of nitrogen in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark nitrogen content per bark dry mass [TOP386] (https://top-thesaurus.org/index) exact match: Bark nitrogen (N) content per bark dry mass [TRY:618] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000638

Bark sodium (Na) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000640
alternative label	bark_Na_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sodium [CHEBI:26708] in bark [PO:0004518] to bark dry mass. The ratio of the mass of sodium in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	100
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	sodium atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark sodium content per bark dry mass [TOP384] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000640

Bark phosphorus (P) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000644
alternative label	bark_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in bark [PO:0004518] to bark dry mass. The ratio of the mass of phosphorus in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom macronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark phosphorus content per bark dry mass [TOP387] (https://top-thesaurus.org/index) exact match: Bark phosphorus (P) content per bark dry mass [TRY:619] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000644

Bark sulphur (S) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000646
alternative label	bark_S_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in bark [PO:0004518] to bark dry mass. The ratio of the mass of sulphur in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	sulfur atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark sulfur content per bark dry mass [TOP390] (https://top-thesaurus.org/index) exact match: Bark sulfur (S) content per bark dry mass [TRY:2963] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000646

Bark zinc (Zn) content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000652
alternative label	bark_Zn_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of zinc [CHEBI:27363] in bark [PO:0004518] to bark dry mass. The ratio of the mass of zinc in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	stem mineral and ion content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	zinc atom micronutrient
scope note
has exact match
has close match
has related match
examples	exact match: Bark zinc content per bark dry mass [TOP391] (https://top-thesaurus.org/index) exact match: Bark zinc (Zn) content per bark dry mass [TRY:2964] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000652

Root carbon (C) content per unit root dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000816
alternative label	root_C_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carbon [CHEBI:27594] in roots [PO:0009005] to root dry mass. The ratio of the mass of carbon in roots to root dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	50
maxAllowedValue	750
has broader	root mineral and ion content trait
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon atom
scope note
has exact match	root carbon content
has close match
has related match
examples	exact match: Root carbon content per root dry mass trait [TOP713] (https://top-thesaurus.org/index) exact match: Root carbon (C) content per root dry mass [TRY:84] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000816

Root nitrogen (N) content per unit root dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000838
alternative label	root_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in roots [PO:0009005] to root dry mass. The ratio of the mass of nitrogen in roots to root dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	root mineral and ion content trait
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient
scope note
has exact match	root nitrogen content
has close match
has related match
examples	exact match: Root nitrogen content per root dry mass trait [TOP723] (https://top-thesaurus.org/index) exact match: Root nitrogen (N) content per root dry mass [TRY:80] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000838

Root phosphorus (P) content per unit root dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0000844
alternative label	root_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in roots [PO:0009005] to root dry mass. The ratio of the mass of phosphorus in roots to root dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	100
has broader	root mineral and ion content trait
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom macronutrient
scope note
has exact match	root system phosphorus content
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0000844

Flower nitrogen (N) content per unit flower dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001038
alternative label	flower_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in flowers [PO:0009046] to flower dry mass. The ratio of the mass of nitrogen in flowers to flower dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.5
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	mass mass fraction ratio; proportion
keyword	reproduction nitrogen atom macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001038

Fruit calcium (Ca) content per unit fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001118
alternative label	fruit_Ca_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in fruit [PO:0009001] to fruit dry mass. The ratio of the mass of calcium in fruit to fruit dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.5
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom micronutrient reproduction
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001118

Fruit potassium (K) content per unit fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001130
alternative label	fruit_K_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in fruit [PO:0009001] to fruit dry mass. The ratio of the mass of potassium in fruit to fruit dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.5
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom macronutrient reproduction
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001130

Fruit magnesium (Mg) content per unit fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001132
alternative label	fruit_Mg_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in fruit [PO:0009001] to fruit dry mass. The ratio of the mass of magnesium in fruit to fruit dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.05
maxAllowedValue	10
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom micronutrient reproduction
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001132

Fruit nitrogen (N) content per unit fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001138
alternative label	fruit_N_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in fruit [PO:0009001] to fruit dry mass. The ratio of the mass of nitrogen in fruit to fruit dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.5
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient reproduction
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001138

Fruit phosphorus (P) content per unit fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001144
alternative label	fruit_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in fruit [PO:0009001] to fruit dry mass. The ratio of the mass of phosphorus in fruit to fruit dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom macronutrient reproduction
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001144

Fruit sulphur (S) content per unit fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001146
alternative label	fruit_S_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in fruit [PO:0009001] to fruit dry mass. The ratio of the mass of sulphur in fruit to fruit dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.05
maxAllowedValue	10
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass mass fraction ratio; proportion
keyword	sulfur atom micronutrient reproduction
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001146

Seed calcium (Ca) content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001218
alternative label	seed_Ca_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in seeds [PO:0009010] to seed dry mass. The ratio of the mass of calcium in seeds to seed dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom micronutrient reproduction
scope note
has exact match
has close match
has related match
examples	exact match: Seed calcium content per seed dry mass [TOP566] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_Ca_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001218

Seed potassium (K) content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001230
alternative label	seed_K_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of potassium [CHEBI:26216] in seeds [PO:0009010] to seed dry mass. The ratio of the mass of potassium in seeds to seed dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	potassium atom macronutrient reproduction
scope note
has exact match
has close match
has related match
examples	exact match: Seed potassium content per seed dry mass [TOP579] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_K_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001230

Seed magnesium (Mg) content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001232
alternative label	seed_Mg_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of magnesium [CHEBI:25107] in seeds [PO:0009010] to seed dry mass. The ratio of the mass of magnesium in seeds to seed dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	magnesium atom micronutrient reproduction
scope note
has exact match
has close match
has related match
examples	exact match: Seed magnesium content per seed dry mass [TOP572] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_Mg_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001232

Seed nitrogen (N) content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001238
alternative label	seed_N_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in seeds [PO:0009010] to seed dry mass. The ratio of the mass of nitrogen in seeds to seed dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	10
maxAllowedValue	1000
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient reproduction
scope note
has exact match	concentration of nitrogen in seeds
has close match
has related match
examples	exact match: Seed nitrogen content per seed dry mass [TOP577] (https://top-thesaurus.org/index) exact match: Seed nitrogen (N) content per dry mass (or diaspore) [TRY:2945] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_N_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001238

Seed phosphorus (P) content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001244
alternative label	seed_P_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in seeds [PO:0009010] to seed dry mass. The ratio of the mass of phosphorus in seeds to seed dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom macronutrient reproduction
scope note
has exact match	seed phosphorus content
has close match
has related match
examples	exact match: Seed phosphorus content per seed dry mass [TOP578] (https://top-thesaurus.org/index) exact match: Seed phosphorus (P) content per dry mass (or diaspore) [TRY:3000] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_P_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001244

Seed sulphur (S) content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001246
alternative label	seed_S_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sulphur [CHEBI:26833] in seeds [PO:0009010] to seed dry mass. The ratio of the mass of sulphur in seeds to seed dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	reproductive shoot system mineral and ion content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	sulfur atom micronutrient reproduction
scope note
has exact match
has close match
has related match
examples	exact match: Seed sulfur content per seed dry mass [TOP581] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_S_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001246

Leaf cell wall nitrogen (N) per unit cell wall dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001511
alternative label	leaf_cell_wall_N_per_cell_wall_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in the plant cell wall [GO:0009505] in a leaf [PO:0025034] to the cell wall dry mass in the leaf. The ratio of the mass of nitrogen in the plant cell wall in a leaf to the cell wall dry mass in the leaf.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mmol{N}/g micromole per gram
minAllowedValue	0.1
maxAllowedValue	10
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient cell plant-type cell wall
scope note
has exact match
has close match
has related match
examples	exact match: Leaf cell wall nitrogen (N) content per cell wall dry mass [TRY:3376] (https://www.try-db.org/de/de.php) close match: Leaf cell wall nitrogen (N) content per leaf area [TRY:3377] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	leaf_cell_wall_N
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001511

Leaf cell wall nitrogen (N) per unit leaf N content

Property	Value
URI	https://w3id.org/APD/traits/trait_0001512
alternative label	leaf_cell_wall_N_per_leaf_N
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in the plant cell wall [GO:0009505] in a leaf [PO:0025034] to the mass of nitrogen in the leaf. The ratio of the mass of nitrogen in the plant cell wall in a leaf to the mass of nitrogen in the leaf.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.01
maxAllowedValue	1
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient cell plant-type cell wall
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	leaf_cell_wall_N_fraction
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001512

Leaf rubisco nitrogen (N) content per unit leaf N content

Property	Value
URI	https://w3id.org/APD/traits/trait_0001513
alternative label	leaf_rubisco_N_per_total_leaf_N
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in rubisco [MESH:D012273] in a leaf [PO:0025034] to the mass of nitrogen in a leaf. The ratio of the mass of nitrogen in rubisco in a leaf to the mass of nitrogen in a leaf.
comments	This trait is most commonly measured on leaf tissue. If the Rubisco concentration of a different tissue is measured, this should be explicitly noted in the methods or as a method context.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.05
maxAllowedValue	0.5
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient cell protein Ribulose-Bisphosphate Carboxylase chloroplast
scope note
has exact match
has close match
has related match	rubisco to nitrogen content ratio
examples	exact match: Leaf nitrogen (N) fraction in Rubisco [TRY:187] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	cell_rubisco_N_per_total_N
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001513

Leaf thylakoid protein nitrogen (N) content per unit leaf N content

Property	Value
URI	https://w3id.org/APD/traits/trait_0001514
alternative label	leaf_thylakoid_N_per_total_leaf_N
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of nitrogen [CHEBI:25555] in thylakoid [GO:0009579] proteins [CHEBI:36080] in a leaf [PO:0025034] to the mass of nitrogen in a leaf. The ratio of the mass of nitrogen in thylakoid proteins in a leaf to the mass of nitrogen in a leaf.
comments	This trait is most commonly measured on leaf tissue. If the Rubisco concentration of a different tissue is measured, this should be explicitly noted in the methods or as a method context.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.05
maxAllowedValue	0.3
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	nitrogen atom macronutrient cell protein thylakoid chloroplast
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	cell_thylakoid_N_per_total_N
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001514

Leaf epidermis calcium (Ca) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001611
alternative label	leaf_epidermis_Ca_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in epidermal cells [PO:0004013] to epidermal cell fresh mass [APD:glossary/40044]. The ratio of the mass of calcium in epidermal cells to epidermal cell fresh mass.
comments
value type	continuous variable
unit	umol{Ca}/g micromole per gram
minAllowedValue	1
maxAllowedValue	500
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom epidermal cell cell micronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_epidermis_Ca_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001611

Leaf hypodermis calcium (Ca) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001612
alternative label	leaf_hypodermis_Ca_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in the hypodermis [PO:0005051] to hypodermis fresh mass [APD:glossary/40044]. The ratio of the mass of calcium in the hypodermis to hypodermis fresh mass.
comments
value type	continuous variable
unit	umol{Ca}/g micromole per gram
minAllowedValue	1
maxAllowedValue	500
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom hypodermis cell micronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_hypodermis_Ca_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001612

Leaf internal parenchyma cell calcium (Ca) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001613
alternative label	leaf_internal_parenchyma_Ca_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in parenchyma cells [PO:0000074] to parenchyma cell fresh mass [APD:glossary/40044]. The ratio of the mass of calcium in parenchyma cells to parenchyma cell fresh mass.
comments
value type	continuous variable
unit	umol{Ca}/g micromole per gram
minAllowedValue	1
maxAllowedValue	500
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom parenchyma cell cell micronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_internal_parenchyma_Ca_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001613

Leaf palisade mesophyll cell calcium (Ca) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001614
alternative label	leaf_palisade_mesophyll_Ca_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in palisade mesophyll cells [PO:0006206] to palisade mesophyll cell fresh mass [APD:glossary/40044]. The ratio of the mass of calcium in palisade mesophyll cells to palisade mesophyll cell fresh mass.
comments
value type	continuous variable
unit	umol{Ca}/g micromole per gram
minAllowedValue	1
maxAllowedValue	100
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom palisade mesophyll cell cell micronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_palisade_mesophyll_Ca_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001614

Leaf sclerenchyma cell calcium (Ca) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001615
alternative label	leaf_sclerenchyma_Ca_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in sclerenchyma cells [PO:0000077] to sclerenchyma cell fresh mass [APD:glossary/40044]. The ratio of the mass of calcium in sclerenchyma cells to sclerenchyma cell fresh mass.
comments
value type	continuous variable
unit	umol{Ca}/g micromole per gram
minAllowedValue	1
maxAllowedValue	100
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom sclerenchyma cell cell micronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_sclerenchyma_Ca_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001615

Leaf spongy mesophyll cell calcium (Ca) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001616
alternative label	leaf_spongy_mesophyll_Ca_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of calcium [CHEBI:22984] in spongy mesophyll cells [PO:0006205] to spongy mesophyll cell fresh mass [APD:glossary/40044]. The ratio of the mass of calcium in spongy mesophyll cells to spongy mesophyll cell fresh mass.
comments
value type	continuous variable
unit	umol{Ca}/g micromole per gram
minAllowedValue	1
maxAllowedValue	1000
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	calcium atom spongy mesophyll cell cell micronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_spongy_mesophyll_Ca_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001616

Leaf epidermis phosphorus (P) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001661
alternative label	leaf_epidermis_P_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in epidermal cells [PO:0004013] to epidermal cell fresh mass [APD:glossary/40044]. The ratio of the mass of phosphorus in epidermal cells to epidermal cell fresh mass.
comments
value type	continuous variable
unit	umol{P}/g micromole per gram
minAllowedValue	0
maxAllowedValue	10
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom epidermal cell cell macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_epidermis_P_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001661

Leaf hypodermis phosphorus (P) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001662
alternative label	leaf_hypodermis_P_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in the hypodermis [PO:0005051] to hypodermis fresh mass [APD:glossary/40044]. The ratio of the mass of phosphorus in the hypodermis to hypodermis fresh mass.
comments
value type	continuous variable
unit	umol{P}/g micromole per gram
minAllowedValue	0
maxAllowedValue	10
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom hypodermis cell macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_hypodermis_P_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001662

Leaf internal parenchyma cell phosphorus (P) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001663
alternative label	leaf_internal_parenchyma_P_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in parenchyma cells [PO:0000074] to parenchyma cell fresh mass [APD:glossary/40044]. The ratio of the mass of phosphorus in parenchyma cells to parenchyma cell fresh mass.
comments
value type	continuous variable
unit	umol{P}/g micromole per gram
minAllowedValue	0
maxAllowedValue	10
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom parenchyma cell cell macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_internal_parenchyma_P_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001663

Leaf palisade mesophyll cell phosphorus (P) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001664
alternative label	leaf_palisade_mesophyll_P_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in palisade mesophyll cells [PO:0006206] to palisade mesophyll cell fresh mass [APD:glossary/40044]. The ratio of the mass of phosphorus in palisade mesophyll cells to palisade mesophyll cell fresh mass.
comments
value type	continuous variable
unit	umol{P}/g micromole per gram
minAllowedValue	0
maxAllowedValue	30
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom palisade mesophyll cell cell macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_palisade_mesophyll_P_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001664

Leaf sclerenchyma cell phosphorus (P) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001665
alternative label	leaf_sclerenchyma_P_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in sclerenchyma cells [PO:0000077] to sclerenchyma cell fresh mass [APD:glossary/40044]. The ratio of the mass of phosphorus in sclerenchyma cells to sclerenchyma cell fresh mass.
comments
value type	continuous variable
unit	umol{P}/g micromole per gram
minAllowedValue	0
maxAllowedValue	10
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom sclerenchyma cell cell macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_sclerenchyma_P_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001665

Leaf spongy mesophyll cell phosphorus (P) content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001666
alternative label	leaf_spongy_mesophyll_P_per_fresh_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in spongy mesophyll cells [PO:0006205] to spongy mesophyll cell fresh mass [APD:glossary/40044]. The ratio of the mass of phosphorus in spongy mesophyll cells to spongy mesophyll cell fresh mass.
comments
value type	continuous variable
unit	umol{P}/g micromole per gram
minAllowedValue	0
maxAllowedValue	30
has broader	cell or tissue mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom spongy mesophyll cell cell macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	cell_spongy_mesophyll_P_per_fresh_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001666

Lipid fraction of leaf P content per dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001667
alternative label	leaf_lipid_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to lipids [CHEBI:18059] to leaf total dry mass [TO:0001014]. The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom lipid macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001667

Nucleic acid fraction of leaf P content per dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001668
alternative label	leaf_nucleic_acid_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of the fraction of total phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to nucleic acids [CHEBI:33696] to leaf total dry mass [TO:0001014]. The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.001
maxAllowedValue	10
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom nucleic acid macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001668

Residual P fraction of leaf P content per dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001669
alternative label	leaf_residual_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to residual P to leaf total dry mass [TO:0001014]. The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0
maxAllowedValue	1
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001669

Inorganic P fraction of leaf P content per dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001670
alternative label	leaf_inorganic_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] that is inorganic [NCIT:C66829] P to leaf total dry mass [TO:0001014]. The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.01
maxAllowedValue	20
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom inorganic macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001670

Metabolite fraction of leaf P content per dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0001671
alternative label	leaf_metabolite_P_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phosphorus [CHEBI:28659] in a leaf [PO:0025034] allocated to metabolites [NCIT:C61154] to leaf total dry mass [TO:0001014]. The ratio of the mass of phosphorus in the lipid fraction of a leaf to leaf dry mass.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1e-04
maxAllowedValue	10
has broader	live leaf mineral and ion content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phosphorus atom metabolite macronutrient
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0001671

Leaf total non-structural carbohydrate content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002021
alternative label	leaf_total_non-structural_carbohydrates_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of non-structural carbohydrates [CHEBI:16646] (sugars [APD:glossary/40106]) in a leaf [PO:0025034] to leaf area [PATO:0001323], where non-structural carbohydrates are carbohydrates that do not comprise the cell wall [GO:0009505]. The ratio of the mass of non-structural carbohydrates (sugars) in a leaf to leaf area, where non-structural carbohydrates are carbohydrates that do not comprise the cell wall.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	50
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	carbohydrate starch sugar non-structural carbohydrates
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002021

Leaf total non-structural carbohydrate content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002022
alternative label	leaf_total_non-structural_carbohydrates_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of non-structural carbohydrates [CHEBI:16646] (sugars [APD:glossary/40106]) in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where non-structural carbohydrates are carbohydrates that do not comprise the cell wall [GO:0009505]. The ratio of the mass of non-structural carbohydrates (sugars) in a leaf to leaf dry mass, where non-structural carbohydrates are carbohydrates that do not comprise the cell wall.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	10
maxAllowedValue	200
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	carbohydrate starch sugar non-structural carbohydrates
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002022

Leaf cellulose content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002024
alternative label	leaf_cellulose_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of cellulose [CHEBI:62967] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of cellulose in a leaf to leaf dry mass.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.01
maxAllowedValue	1
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	amorphous cellulose carbohydrate
scope note
has exact match
has close match
has related match	cellulose content
examples	exact match: Leaf cellulose content per leaf dry mass [TOP819] (https://top-thesaurus.org/index) exact match: Leaf cellulose content per leaf dry mass [TRY:92] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002024

Leaf starch content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002025
alternative label	leaf_starch_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of total starch [CHEBI:28017] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of total starch in a leaf to leaf area.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	50
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	starch carbohydrate
scope note
has exact match
has close match
has related match	starch content
examples	exact match: Leaf starch content per leaf dry mass [TRY:505] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	starch_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002025

Leaf soluble starch content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002027
alternative label	leaf_soluble_starch_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of water soluble starch in a leaf to leaf area.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	100
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	starch carbohydrate
scope note
has exact match
has close match
has related match
examples	exact match: Leaf starch content per leaf dry mass [TRY:505] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	soluble_starch_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002027

Leaf soluble starch content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002028
alternative label	leaf_soluble_starch_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of water soluble starch in a leaf to leaf dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	200
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	starch carbohydrate
scope note
has exact match
has close match
has related match
examples	exact match: Leaf starch content per leaf dry mass [TRY:505] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002028

Leaf soluble sugar content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002031
alternative label	leaf_soluble_sugars_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a leaf [PO:0025034] to leaf area [PATO:0001323], where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233]. The ratio of the mass of sugar in a leaf to leaf area, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	100
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	carbohydrate monosaccharide disaccharide sugar
scope note
has exact match
has close match
has related match	total soluble sugar content
examples	exact match: Leaf sugar content per leaf area [TOP920] (https://top-thesaurus.org/index) exact match: Leaf sugar content per leaf dry mass [TRY:506] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	soluble_sugars_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002031

Leaf soluble sugar content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002032
alternative label	leaf_soluble_sugars_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233]. The ratio of the mass of sugar in a leaf to leaf dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	200
has broader	carbohydrate content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	carbohydrate monosaccharide disaccharide sugar
scope note
has exact match
has close match
has related match	total soluble sugar content
examples	exact match: Leaf sugar content per leaf dry mass [TOP919] (https://top-thesaurus.org/index) exact match: Leaf sugar content per leaf dry mass [TRY:506] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002032

Leaf soluble protein content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002035
alternative label	leaf_soluble_protein_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] protein [CHEBI:36080] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of water soluble protein in a leaf to leaf area.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	20
has broader	protein content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	protein
scope note
has exact match	Leaf soluble protein content
has close match
has related match	soluble protein content
examples	close match: Leaf protein content per leaf dry mass [TRY:503] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	soluble_protein_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002035

Leaf insoluble protein content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002037
alternative label	leaf_insoluble_protein_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] insoluble [PATO:0001538] protein [CHEBI:36080] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of water insoluble protein in a leaf to leaf area.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	20
has broader	protein content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	protein
scope note
has exact match
has close match
has related match
examples	close match: Leaf protein content per leaf dry mass [TRY:503] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	insoluble_protein_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002037

Leaf lignin content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002050
alternative label	leaf_lignin_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of lignin [CHEBI:6457] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of lignin in a leaf to leaf dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	10
maxAllowedValue	1000
has broader	phenolic compound content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	lignin
scope note
has exact match
has close match
has related match	lignin content
examples	exact match: Leaf lignin content per leaf dry mass [TOP844] (https://top-thesaurus.org/index) exact match: Leaf lignin content per leaf dry mass [TRY:87] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed	13/02/2023
reviewed by	Félix de Tombeur
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002050

Total leaf phenolic content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002052
alternative label	leaf_phenol_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of phenol [CHEBI:33853] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where phenol content is measured as gallic acid [CHEBI:30778] equivalents. The ratio of the mass of phenol in a leaf to leaf dry mass, where phenol content is measured as gallic acid equivalents.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	1000
has broader	phenolic compound content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phenol gallic acid
scope note
has exact match
has close match
has related match
examples	exact match: Leaf phenols content per leaf dry mass [TRY:147] (https://www.try-db.org/de/de.php)
references	Salminen & Karonen 2011
date created	14/07/2021
date modified	05/02/2024
date reviewed	13/02/2023
reviewed by	Félix de Tombeur
change note	leaf_phenol_content_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002052

Leaf tannin content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002054
alternative label	leaf_tannin_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of tannins [CHEBI:26848] (a class of polyphenols [CHEBI:26195]) in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of tannins (a class of polyphenols) in a leaf to leaf dry mass.
comments	Tannins are a specific class of polyphenolic compounds found in plants.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	1000
has broader	phenolic compound content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	phenol tannin polyphenol
scope note
has exact match
has close match
has related match
examples	exact match: Leaf tannin content per leaf dry mass [TRY:148] (https://www.try-db.org/de/de.php)
references	Salminen & Karonen 2011
date created	14/07/2021
date modified	06/06/2025
date reviewed	13/02/2023
reviewed by	Félix de Tombeur
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002054

Leaf carotenoid content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002055
alternative label	leaf_carotenoid_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carotenoids [CHEBI:23044] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of carotenoids in a leaf to leaf area.
comments
value type	continuous variable
unit	umol{carotenoid}/m2 micromole per square metre
minAllowedValue	10
maxAllowedValue	1000
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	carotenoid
scope note
has exact match
has close match
has related match	carotenoid content
examples	exact match: Leaf carotenoid content per leaf area [TRY:491] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	carotenoid_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002055

Leaf carotenoid content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002056
alternative label	leaf_carotenoid_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of carotenoids [CHEBI:23044] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of carotenoids in a leaf to leaf dry mass.
comments
value type	continuous variable
unit	mmol{carotenoid}/kg millimole per kilogram
minAllowedValue	0.1
maxAllowedValue	10
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	carotenoid
scope note
has exact match
has close match
has related match	carotenoid content
examples	exact match: Leaf carotenoid content per leaf dry mass [TRY:809] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	carotenoid_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002056

Leaf total chlorophyll content (chlorophyll A + B) per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002081
alternative label	leaf_chlorophyll_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll [CHEBI:28966] in a leaf [PO:0025034] to leaf area [PATO:0001323], where chlorophyll content is the sum of chlorophyll a [CHEBI:18230] and chlorophyll b [CHEBI:27888] content. The ratio of the mass of chlorophyll in a leaf to leaf area, where chlorophyll content is the sum of chlorophyll a and chlorophyll b content.
comments
value type	continuous variable
unit	umol{chlorophyll}/m2 micromole per square metre
minAllowedValue	10
maxAllowedValue	2000
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	chlorophyll chlorophyll a chlorophyll b chloroplast
scope note
has exact match	Leaf chlorophyll content
has close match	leaf chlorophyll content
has related match	chlorophyll content
examples	exact match: Leaf chlorophyll content per leaf area [TRY:413] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002081

Leaf total chlorophyll content (chlorophyll A + B) per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002082
alternative label	leaf_chlorophyll_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll [CHEBI:28966] in a leaf [PO:0025034] to leaf dry mass [TO:0001014], where chlorophyll content is the sum of chlorophyll a [CHEBI:18230] and chlorophyll b [CHEBI:27888] content. The ratio of the mass of chlorophyll in a leaf to leaf dry mass, where chlorophyll content is the sum of chlorophyll a and chlorophyll b content.
comments
value type	continuous variable
unit	mmol{chlorophyll}/kg millimole per kilogram
minAllowedValue	0.1
maxAllowedValue	15
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	chlorophyll chlorophyll a chlorophyll b chloroplast
scope note
has exact match	Leaf chlorophyll content
has close match	leaf chlorophyll content
has related match	chlorophyll content
examples	exact match: Leaf chlorophyll content per leaf dry mass [TRY:164] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002082

Leaf chlorophyll A content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002083
alternative label	leaf_chlorophyll_A_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of chlorophyll a in a leaf to leaf area.
comments
value type	continuous variable
unit	umol{chlorophyllA}/m2 micromole per square metre
minAllowedValue	50
maxAllowedValue	2000
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	chlorophyll a chlorophyll chloroplast
scope note
has exact match
has close match	Chlorophyll a chlorophyll a content
has related match
examples	exact match: Leaf chlorophyll content per leaf area (Chl a) [TRY:3476] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_A_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002083

Leaf chlorophyll A content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002084
alternative label	leaf_chlorophyll_A_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of chlorophyll a in a leaf to leaf dry mass.
comments
value type	continuous variable
unit	mmol{chlorophyllA}/kg millimole per kilogram
minAllowedValue	0.1
maxAllowedValue	10
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	chlorophyll a chlorophyll chloroplast
scope note
has exact match	Chlorophyll a
has close match	chlorophyll a content
has related match
examples	exact match: Leaf chlorophyll content per leaf dry mass (Chl a) [TRY:3474] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_A_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002084

Leaf chlorophyll B content per unit leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0002085
alternative label	leaf_chlorophyll_B_per_area
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll b [CHEBI:27888] in a leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of chlorophyll b in a leaf to leaf area.
comments
value type	continuous variable
unit	umol{chlorophyllB}/m2 micromole per square metre
minAllowedValue	10
maxAllowedValue	1000
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	chlorophyll b chlorophyll chloroplast
scope note
has exact match
has close match	Chlorophyll b chlorophyll b content
has related match
examples	exact match: Leaf chlorophyll content per leaf area (Chl b) [TRY:3477] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_B_per_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002085

Leaf chlorophyll B content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002086
alternative label	leaf_chlorophyll_B_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll b [CHEBI:27888] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of chlorophyll b in a leaf to leaf dry mass.
comments
value type	continuous variable
unit	mmol{chlorophyllB}/kg millimole per kilogram
minAllowedValue	0.01
maxAllowedValue	10
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	chlorophyll b chlorophyll chloroplast
scope note
has exact match	Chlorophyll b
has close match	chlorophyll b content
has related match
examples	exact match: Leaf chlorophyll content per leaf dry mass (Chl b) [TRY:3475] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_B_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002086

Ratio of leaf chlorophyll A content to leaf chlorophyll B content

Property	Value
URI	https://w3id.org/APD/traits/trait_0002087
alternative label	leaf_chlorophyll_A_B_ratio
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of chlorophyll a [CHEBI:18230] in a leaf [PO:0025034] to the mass of chlorophyll b [CHEBI:27888] in the leaf. The ratio of the mass of chlorophyll a in a leaf to the mass of chlorophyll b in the leaf.
comments
value type	continuous variable
unit	umol{chlorophyllA}/umol{chlorophyllB} micromole per micromole
minAllowedValue	0.1
maxAllowedValue	6
has broader	pigment content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion dimensionless ratio
keyword	chlorophyll a chlorophyll b chlorophyll chloroplast
scope note
has exact match	Chlorophyll a to b ratio chlorophyll ratio
has close match
has related match
examples	exact match: Leaf chlorophyll a/b ratio [TRY:416] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note	chlorophyll_A_B_ratio
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002087

Leaf rubisco content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002090
alternative label	leaf_rubisco_per_leaf_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of rubisco [MESH:D012273] in a leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of rubisco in a leaf to leaf dry mass.
comments	This trait is most commonly measured on leaf tissue. If the Rubisco concentration of a different tissue is measured, this should be explicitly noted in the methods or as a method context.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	5
maxAllowedValue	50
has broader	protein content trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	Ribulose-Bisphosphate Carboxylase chloroplast protein
scope note
has exact match	Leaf rubisco content
has close match	rubisco content
has related match
examples	close match: Leaf Rubisco content per leaf area [TRY:504] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	cell_rubisco_concentration
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002090

Stem soluble starch content per unit stem dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002127
alternative label	stem_soluble_starch_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in a stem [PO:0009047] to stem dry mass. The ratio of the mass of water soluble starch in a stem to stem dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	carbohydrate content trait
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	starch carbohydrate
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002127

Stem soluble sugar content per unit stem dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002131
alternative label	stem_soluble_sugars_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in a stem [PO:0009047] to stem dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233]. The ratio of the mass of sugar in a stem to stem dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	200
has broader	carbohydrate content trait
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	carbohydrate monosaccharide disaccharide sugar
scope note
has exact match
has close match
has related match	sugar content
examples	exact match: Stem sugar content per stem dry mass [TOP914] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002131

Bark cellulose content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002224
alternative label	bark_cellulose_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of cellulose [CHEBI:62967] in bark [PO:0004518] to bark dry mass. The ratio of the mass of cellulose in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.1
maxAllowedValue	1
has broader	carbohydrate content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	amorphous cellulose
scope note
has exact match
has close match
has related match
examples	exact match: Bark cellulose content per bark dry mass [TOP813] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002224

Bark lignin content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002250
alternative label	bark_lignin_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of lignin [CHEBI:6457] in bark [PO:0004518] to bark dry mass. The ratio of the mass of lignin in bark to bark dry mass.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.01
maxAllowedValue	1
has broader	phenolic compound content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	lignin
scope note
has exact match
has close match
has related match
examples	exact match: Bark lignin content per bark dry mass [TOP838] (https://top-thesaurus.org/index)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002250

Bark tannin content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002255
alternative label	bark_tannin_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of tannins [CHEBI:26848] (polyphenols [CHEBI:26195]) in bark [PO:0004518] to bark dry mass. The ratio of the mass of tannins (polyphenols) in bark to bark dry mass.
comments	Tannins are a specific class of polyphenolic compounds found in plants.
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	1e-05
maxAllowedValue	1
has broader	phenolic compound content trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	tannin polyphenol
scope note
has exact match
has close match
has related match
examples
references	Salminen & Karonen 2011
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002255

Root soluble starch content per unit root dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002327
alternative label	root_soluble_starch_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] soluble [PATO:0001537] starch [CHEBI:28017] in roots [PO:0009005] to root dry mass. The ratio of the mass of water soluble starch in roots to root dry mass.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	0.1
maxAllowedValue	100
has broader	carbohydrate content trait
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	starch carbohydrate
scope note
has exact match
has close match
has related match
examples	exact match: Root starch content per dry mass [TRY:3147] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002327

Root soluble sugar content per unit root dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002331
alternative label	root_soluble_sugars_per_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of sugar [APD:glossary/40106] in roots [PO:0009005] to root dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates [CHEBI:16646] that are soluble in [PATO:0001537] water [CHEBI:15377], including both monosaccharides [CHEBI:35381] and disaccharides [CHEBI:36233]. The ratio of the mass of sugar in roots to root dry mass, where ‘sugar’ refers to an informal grouping of carbohydrates that are soluble in water, including both monosaccharides and disaccharides.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	1
maxAllowedValue	200
has broader	carbohydrate content trait
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	carbohydrate monosaccharide disaccharide sugar
scope note
has exact match
has close match
has related match
examples	exact match: Root sugar content per root dry mass trait [TOP921] (https://top-thesaurus.org/index) exact match: Root sugar content per root dry mass [TRY:662] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002331

Seed protein content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002534
alternative label	seed_protein_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of protein [CHEBI:36080] in a seed [PO:0009010] to seed dry mass [TO:0000181]. The ratio of the mass of protein in a seed to seed dry mass.
comments	It is best practice for the denominator to be seed dry mass. If data are reported on a per seed fresh mass basis, this should be explicitly noted in the methods or as a method context.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	1e-05
maxAllowedValue	1
has broader	protein content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	protein seed weight, seed mass
scope note
has exact match	seed protein content
has close match
has related match
examples	exact match: Seed protein content per seed mass [TRY:97] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_protein_content
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002534

Seed oil content per unit seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002544
alternative label	seed_oil_per_seed_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of lipids [CHEBI:18059] in a seed [PO:0009010] to seed dry mass [TO:0000181]. The ratio of the mass of lipids in a seed to seed dry mass.
comments	It is best practice for the denominator to be seed dry mass. If data are reported on a per seed fresh mass basis, this should be explicitly noted in the methods or as a method context.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	1e-04
maxAllowedValue	1
has broader	lipid content trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass mass fraction ratio; proportion
keyword	lipid seed weight, seed mass
scope note
has exact match
has close match
has related match
examples	exact match: Seed lipid content per seed dry mass [TOP880] (https://top-thesaurus.org/index) exact match: Seed oil content per seed mass [TRY:96] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seed_oil_content
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002544

Leaf ash content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002822
alternative label	leaf_ash_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of leaf [PO:0025034] ash [ENVO:02000090] remaining after a combustion process [ENVO:01000839] to the leaf dry mass before the combustion process. The ratio of the mass of leaf ash remaining after a combustion process to the leaf dry mass before the combustion process.
comments	Includes measurements on leaf analogs, including cladodes and phylloclades.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	1e-04
maxAllowedValue	1
has broader	live leaf mineral and ion content trait fire response trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	ash combustion process fire
scope note
has exact match
has close match
has related match
examples	exact match: Leaf ash content per leaf dry mass [TRY:333] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	leaf_ash_content_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002822

Bark ash content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0002824
alternative label	bark_ash_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of bark [PO:0004518] ash [ENVO:02000090] remaining after a combustion process [ENVO:01000839] to the bark dry mass before the combustion process. The ratio of the mass of bark ash remaining after a combustion process to the bark dry mass before the combustion process.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	1e-04
maxAllowedValue	1
has broader	stem mineral and ion content trait fire response trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	ash combustion process fire
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	bark_ash_content_per_dry_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0002824

Bark stable carbon isotope composition (delta13C)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003011
alternative label	bark_delta13C
description	Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of bark [PO:0004518]. Carbon stable isotope ratio, d13C, of bark.
comments	delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-50
maxAllowedValue	0
has broader	stable isotope ratio determination
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon stable isotope ratio d13C carbon-13 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Bark carbon (C) isotope signature (delta 13C) [TRY:829] (https://www.try-db.org/de/de.php) related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003011

Leaf stable carbon isotope composition (delta13C)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003012
alternative label	leaf_delta13C
description	Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of leaves [PO:0025034]. Carbon stable isotope ratio, d13C, of leaves.
comments	delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-50
maxAllowedValue	0
has broader	stable isotope ratio determination
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon stable isotope ratio d13C carbon-13 atom isotope
scope note
has exact match	Delta C13
has close match
has related match
examples	exact match: Leaf carbon isotope signature (delta 13C) [TRY:89] (https://www.try-db.org/de/de.php) related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003012

Stem stable carbon isotope composition (delta13C)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003013
alternative label	stem_delta13C
description	Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of stems [PO:0009047]. Carbon stable isotope ratio, d13C, of stems.
comments	delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-50
maxAllowedValue	0
has broader	stable isotope ratio determination
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon stable isotope ratio d13C carbon-13 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Stem carbon isotope ratio (delta 13C) [TRY:4049] (https://www.try-db.org/de/de.php) related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	29/03/2023
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003013

Root stable carbon isotope composition (delta13C)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003014
alternative label	root_delta13C
description	Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of roots [PO:0009005]. Carbon stable isotope ratio, d13C, of roots.
comments	delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-50
maxAllowedValue	0
has broader	stable isotope ratio determination
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon stable isotope ratio d13C carbon-13 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Root carbon isotope signature (delta 13C) [TRY:90] (https://www.try-db.org/de/de.php) related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003014

Wood stable carbon isotope composition (delta13C)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003015
alternative label	wood_delta13C
description	Carbon stable isotope ratio [ECSO:00001142], d13C [ECSO:00000669], of wood [NCIT:C93062]. Carbon stable isotope ratio, d13C, of wood.
comments	delta13C is calculated as the ratio of, 13C/12C in a sample to 13C/12C in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A more negative value for delta13C indicates the sample has a lower ratio of 13C/12C relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-50
maxAllowedValue	0
has broader	stable isotope ratio determination
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	carbon stable isotope ratio d13C wood carbon-13 atom isotope
scope note
has exact match	Wood Carbon content
has close match
has related match
examples	exact match: Wood carbon (C) isotope signature (delta 13C) [TRY:1213] (https://www.try-db.org/de/de.php) related match: Integrated water-use efficiency [BROT:16] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003015

Bark stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003031
alternative label	bark_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of bark [PO:0004518]. Nitrogen stable isotope ratio, d15N, of bark.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N nitrogen-15 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Bark nitrogen (N) isotope signature (delta 15N) [TRY:832] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003031

Leaf stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003032
alternative label	leaf_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of leaves [PO:0025034]. Nitrogen stable isotope ratio, d15N, of leaves.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N nitrogen-15 atom isotope
scope note
has exact match	Delta 15N
has close match
has related match
examples	exact match: Leaf nitrogen (N) isotope signature (delta 15N) [TRY:78] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003032

Stem stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003033
alternative label	stem_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of stems [PO:0009047]. Nitrogen stable isotope ratio, d15N, of stems.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N nitrogen-15 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Root nitrogen (N) isotope signature (delta 15N) [TRY:81] (https://www.try-db.org/de/de.php)
references	no linked references
date created	29/03/2023
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003033

Root stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003034
alternative label	root_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of roots [PO:0009005]. Nitrogen stable isotope ratio, d15N, of roots.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N nitrogen-15 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Root nitrogen (N) isotope signature (delta 15N) [TRY:81] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003034

Wood stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003035
alternative label	wood_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of wood [NCIT:C93062]. Nitrogen stable isotope ratio, d15N, of wood.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure plant structure	stem wood, secondary xylem
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N wood nitrogen-15 atom isotope
scope note
has exact match
has close match
has related match
examples	exact match: Wood nitrogen (N) isotope signature (delta 15N) [TRY:1230] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003035

Leaf xylem stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003052
alternative label	leaf_xylem_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of xylem sap [PO:0025539] from leaf [PO:0025034] xylem vessels [PO:0025417]. Nitrogen stable isotope ratio, d15N, of xylem sap from leaf xylem vessels.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N xylem vessel nitrogen-15 atom isotope xylem sap
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003052

Root xylem stable nitrogen isotope composition (delta15N)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003053
alternative label	root_xylem_delta15N
description	Nitrogen stable isotope ratio [ECSO:00001737], d15N [ECSO:00001738], of xylem sap [PO:0025539] from root [PO:0009005] xylem vessels [PO:0025417]. Nitrogen stable isotope ratio, d15N, of xylem sap from root xylem vessels.
comments	delta15N is calculated as the ratio of, 15N/14N in a sample to 15N/14N in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta15N indicates the sample has a lower ratio of 15N/14N relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-25
maxAllowedValue	75
has broader	stable isotope ratio determination
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	stable nitrogen isotope ratio d15N xylem vessel nitrogen-15 atom isotope xylem sap
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003053

Leaf stable oxygen isotope composition (delta18O)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003072
alternative label	leaf_delta18O
description	Oxygen stable isotope ratio [ECSO:00001740], d18O [ECSO:00001677], of leaves [PO:0025034]. Oxygen stable isotope ratio, d18O, of leaves.
comments	delta18O is calculated as the ratio of, 18O/16O in a sample to 18O/16O in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta18O indicates the sample has a lower ratio of 18O/16O relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-10
maxAllowedValue	50
has broader	stable isotope ratio determination
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	stable oxygen isotope ratio d18O oxygen-18 atom
scope note
has exact match
has close match
has related match
examples	exact match: Leaf oxygen (O) isotope signature (delta 18O) [TRY:3358] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003072

Stem water stable oxygen isotope composition (delta18O)

Property	Value
URI	https://w3id.org/APD/traits/trait_0003092
alternative label	stem_water_delta18O
description	Oxygen stable isotope ratio [ECSO:00001740], d18O [ECSO:00001677], of water [CHEBI:15377] in stems [PO:0009047]. Oxygen stable isotope ratio, d18O, of water in stems.
comments	delta18O is calculated as the ratio of, 18O/16O in a sample to 18O/16O in a standard, minus 1. The result is expressed in parts per thousand, commonly referred to as per mille. A lower value for delta18O indicates the sample has a lower ratio of 18O/16O relative to the standard.
value type	continuous variable
unit	{delta}[ppth] parts per thousand
minAllowedValue	-10
maxAllowedValue	50
has broader	stable isotope ratio determination
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	stable oxygen isotope ratio d18O water oxygen-18 atom d18O in water
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0003092

Plant canopy width

Property	Value
URI	https://w3id.org/APD/traits/trait_0010021
alternative label	plant_width
description	A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] width [PATO:0000600] dimension of a plant canopy [ENVO:01001242] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341]. This is a plant’s larger [NCIT:C49508] width dimension and is orthogonal to its height and breadth. Maximum width dimension of a plant’s vegetative canopy for a taxon in a given location; this is the larger width dimension and is orthogonal to its height and breadth.
comments	This trait is not meant to document the width of all individuals sampled, but instead the maximum width for a typical mature individual within a population or across a taxon.
value type	continuous variable
unit	m metre
minAllowedValue	0.001
maxAllowedValue	20
has broader	whole plant morphology trait
plant structure	whole plant
measured characteristic	width size
keyword	canopy vegetation
scope note
has exact match	plant width
has close match	Crown size
has related match
examples	exact match: Crown (canopy) width [TRY:325] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010021

Plant canopy breadth

Property	Value
URI	https://w3id.org/APD/traits/trait_0010022
alternative label	plant_breadth
description	A whole plant morphology trait [TO:0000398] which is the smaller [NCIT:C25376] width [PATO:0000600] dimension of a plant canopy [ENVO:01001242]. Shortest width dimension of a plant’s vegetative canopy; orthogonal to its height and width.
comments	This trait is not meant to document the breadth of all individuals sampled, but instead the maximum breadth for a typical mature individual within a population or across a taxon.
value type	continuous variable
unit	m metre
minAllowedValue	0.001
maxAllowedValue	20
has broader	whole plant morphology trait
plant structure	whole plant
measured characteristic	width size
keyword	canopy vegetation
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010022

Plant vegetative height

Property	Value
URI	https://w3id.org/APD/traits/trait_0010023
alternative label	plant_height
description	A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the vegetative shoot system [PO:0025607] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341]. Maximum vertical height of the vegetative shoot system for a taxon in a given location.
comments	This trait is not meant to document the height of all individuals sampled, but instead the maximum height for a typical mature individual within a population or across a taxon. The trait should only be scored for self-supporting plants, not climbers whose height is dependent on the surrounding vegetation. The documented height should be the maximum height achieved by vegetative shoots, not the height reached by reproductive shoots, as it represents the plant’s access to light.
value type	continuous variable
unit	m metre
minAllowedValue	0.001
maxAllowedValue	130
has broader	whole plant morphology trait
plant structure plant structure	whole plant stem
measured characteristic	height size
keyword	canopy vegetation vegetative shoot system
scope note	self-supporting plants
has exact match	Tree height Tree height plant height
has close match	plant height
has related match
examples	exact match: Plant height vegetative [TOP69] (https://top-thesaurus.org/index) exact match: Plant height vegetative [TRY:3106] (https://www.try-db.org/de/de.php) exact match: Plant_height_max [GIFT:1.6.2] (https://gift.uni-goettingen.de) exact match: maximum whole plant height [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: whole plant height [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) related match: minimum whole plant height [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Average height [BROT:10] (http://doi.org/10.1038/sdata.2018.135) exact match: MaxStemHeight_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	Pérez-Harguindeguy 2013 Falster 2003a
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010023

Stem diameter at breast height

Property	Value
URI	https://w3id.org/APD/traits/trait_0010024
alternative label	plant_diameter_breast_height
description	A stem morphology trait [TO:0000361] which is the maximum [NCIT:C25564] diameter at breast height [ECSO:00000549] for a taxon [NCIT:C40098] in a given population [PCO:0000001] or location [NCIT:C25341]. Maximum diameter at breast height (DBH) for a taxon in a given location. It is commonly referred to as ‘DBH’.
comments	This trait is always meant to be measured at ‘breast height’, 1.4 m above the ground surface. However, for short plants this will not always be the case, so it is advised to read the methods to confirm at what height plant diameters have been recorded. This trait is not meant to document the DBH of all individuals sampled, but instead the maximum DBH for a typical mature individual within a population or across a taxon.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	5000
has broader	stem morphology trait
plant structure plant structure	stem whole plant
measured characteristic	diameter size
keyword	tree diameter at breast height vegetation
scope note
has exact match	Stem diameter tree breast height diameter
has close match	Stem diameter stem diameter
has related match
examples	exact match: Stem diameter [TRY:21] (https://www.try-db.org/de/de.php) exact match: DBH_max [GIFT:1.8.2] (https://gift.uni-goettingen.de) exact match: diameter at breast height (1.3 m) [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: MaxStemDia_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	basal_diameter
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010024

Stem count

Property	Value
URI	https://w3id.org/APD/traits/trait_0010025
alternative label	stem_count
description	A shoot system morphology trait [TO:0000077] which is the count [NCIT:C25463] of stems [PO:0009047] on multi-stemmed plants [PO:0000003]. Count of stems on multi-stemmed plants.
comments
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	5
has broader	stem morphology trait
plant structure	stem
measured characteristic	has number of count
keyword	branchiness
scope note
has exact match	stem number
has close match
has related match
examples	close match: Plant biomass and allometry: Shoots per plant [TRY:1034] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stem_count_categorical
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010025

Height of a climbing plant

Property	Value
URI	https://w3id.org/APD/traits/trait_0010026
alternative label	plant_height_climbing_plant
description	A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the vegetative shoot system [PO:0025607] of a climbing plant that is not self-supporting and therefore has a height that depends on the surrounding vegetation. Maximum vertical height of the vegetative shoot system of a non-self-supporting taxon (i.e. a vine, liana, or other climbing plant) in a given location.
comments	While this trait is intended to indicate the height of climbing plants, the height of an epiphyte relative to the soil surface could also be documented here. All heights reported for this trait will be dependent on the height of the surrounding vegetation.
value type	continuous variable
unit	m metre
minAllowedValue	0.1
maxAllowedValue	100
has broader	whole plant morphology trait
plant structure	whole plant
measured characteristic	height size
keyword	canopy vegetation vegetative shoot system
scope note	applicable to plants that are not self-supporting
has exact match
has close match
has related match	plant height
examples	related match: Plant height [TRY:18] (https://www.try-db.org/de/de.php) related match: Average height [BROT:10] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010026

Plant reproductive height

Property	Value
URI	https://w3id.org/APD/traits/trait_0010027
alternative label	plant_height_reproductive
description	A whole plant morphology trait [TO:0000398] which is the maximum [NCIT:C25564] vertical [PATO:0001854] height [PATO:0000119] of the reproductive shoot system [PO_0025082]. Maximum vertical height of the reproductive shoot system for a taxon in a given location.
comments	This trait can be scored for any species, but is intended as an alternative to plant_height (maximum vegetative height) for species that do not have vegetative stems, but instead only leaves and reproductive stems that extend past the soil surface.
value type	continuous variable
unit	m metre
minAllowedValue	0.1
maxAllowedValue	100
has broader	whole plant morphology trait
plant structure	whole plant
measured characteristic	height size
keyword	vegetation reproductive shoot system
scope note
has exact match
has close match
has related match
examples	related match: Plant height [TRY:18] (https://www.try-db.org/de/de.php)
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010027

Stem length

Property	Value
URI	https://w3id.org/APD/traits/trait_0010028
alternative label	stem_length
description	A shoot system morphology trait [TO:0000077] which is the maximum [NCIT:C25564] length [PATO_0000122] of a plant’s stems [PO_0009047]. Maximum length or extent of a plant’s stems.
comments	This trait can be scored for any species, but is intended as an alternative to plant_height (maximum vegetative height) for species that are prostrate, crawling, creeping or similar.
value type	continuous variable
unit	m metre
minAllowedValue	0.01
maxAllowedValue	10
has broader	stem morphology trait
plant structure plant structure	stem whole plant
measured characteristic	length size
keyword	vegetation vegetative shoot system
scope note
has exact match
has close match
has related match
examples	exact match: Stem length [TRY:326] (https://www.try-db.org/de/de.php)
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010028

Plant spinescence

Property	Value
URI	https://w3id.org/APD/traits/trait_0010070
alternative label	plant_spinescence
description	A plant structure morphology trait [TO:0000839] which summarises the extent of physical structures a plant uses for defence [NCIT:C61556], including the size [PATO:0000117], strength [PATO:0001230], and density [PATO:0001019] of spine leaves [PO:0025173], spine stipules [PO:0025174], thorns [PO:0025172], and/or prickles [PO:0025169] and the pain [HP:0012531] inflicted by these structures. Degree to which a plant is defended by spines, thorns and/or prickles, as defined by P<U+00C3><U+00A9>rez-Harguindeguy et al. 2013.
comments	The values for this trait are exactly those defined by P<U+00C3><U+00A9>rez-Harguindeguy et al. 2013 (doi:10.1071/BT12225). See the trait plant_physical_defence for individual plant structures a plant uses for physical defence.
value type	categorical variable
has narrower	plant_spinescence_no_spines No spines. (level 0) plant_spinescence_low_density_soft_spines Low or very local density of soft spines <5mm long; plant may sting or prickle when hit carelessly but not impart strong pain. (level 1) plant_spinescence_high_density_soft_spines High density of soft spines, intermediate density of spines of intermediate hardness, or low density of hard, sharp spines >5mm long; plant causes actual pain when hit carelessly. (level 2) plant_spinescence_high_density_hard_spines_to_5mm Intermediate or high density of hard, sharp spines >5mm long; plant causes strong pain when hit carelessly. (level 3) plant_spinescence_high_density_hard_spines_to_20mm Intermediate or high density of hard, sharp spines >20mm long; plant may cause significant wounds when hit carelessly. (level 4) plant_spinescence_high_density_hard_spines_to_100mm Intermediate or high density of hard, sharp spines >100mm long; plant is dangerous to careless large mammals, including humans. (level 5)
has broader	leaf shape trait whole plant phenotype trait plant structure morphology trait
plant structure plant structure plant structure	leaf stem whole plant
measured characteristic	structure surface feature shape
keyword	spine leaf spine stipule thorn prickle defense pain spiny
scope note	none
has exact match
has close match
has related match
examples	close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php) close match: Spinescence [BROT:21] (http://doi.org/10.1038/sdata.2018.135) close match: Armature [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	spinescence
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010070

Embryo colour

Property	Value
URI	https://w3id.org/APD/traits/trait_0010110
alternative label	embryo_colour
description	A plant embryo cotyledon morphology trait [TO:0000775] which is the colour [PATO:0000014] of the plant embryo cotyledon [PO:0025470]. Binary variable distinguishing between embryos that are green versus colourless.
comments
value type	categorical variable
has narrower	embryo_colour_colourless Colourless embryo. embryo_colour_green Green embryo.
has broader	plant embryo morphology trait
plant structure	whole plant
measured characteristic	colour / color
keyword	plant embryo cotyledon plant embryo
scope note
has exact match	plant embryo cotyledon color
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010110

Cotyledon function

Property	Value
URI	https://w3id.org/APD/traits/trait_0010111
alternative label	cotyledon_function
description	A plant embryo morphology trait [TO:0000064] which indicates if the cotyledons [PO:0020030] have a photosynthesis [GO:0015979] or energy [PATO:0001021] storage [OBI:0302893] function [NCIT:C25518]. Binary variable distinguishing between cotyledons that function as leaves and can photosynthesise versus ones that function as energy reserve organs.
comments
value type	categorical variable
has narrower	cotyledon_function_foliaceous Cotyledons are photosynthetic and function as leaves. cotyledon_function_reserve Cotyledons function as energy reserve organs.
has broader	plant embryo morphology trait leaf morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	organismal quality function
keyword	cotyledon photosynthesis energy storage plant embryo seedling plant embryo cotyledon storage organ
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010111

Cotyledon position at germination

Property	Value
URI	https://w3id.org/APD/traits/trait_0010112
alternative label	cotyledon_position
description	A plant embryo morphology trait [TO:0000064] which indicates if the cotyledons [PO:0020030] emerge from the seed coat [PO:0009088] at germination [GO:0009845]. Binary variable distinguishing between seedlings where the cotyledon remains within the seed coat versus emerges from the seed coat at germination.
comments
value type	categorical variable
has narrower	cotyledon_position_cryptocotylar A type of seed germination in which the cotyledons remain within the seed coat at germination. cotyledon_position_phanerocotylar A type of seed germination in which the cotyledons emerge from the seed coat.
has broader	plant embryo morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	organismal quality structure position
keyword	cotyledon seed coat seed germination plant embryo seedling plant embryo cotyledon
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010112

Cotyledon hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0010113
alternative label	cotyledon_hairs
description	A plant embryo cotyledon morphology trait [TO:0000775] which is the pilosity [PATO:0000066] of the cotyledons [PO:0020030]. Binary variable distinguishing between glabrous versus hairy cotyledons.
comments
value type	categorical variable
has narrower	cotyledon_hairs_glabrous Cotyledons lack hairs. cotyledon_hairs_hairy Cotyledons have hairs.
has broader	plant embryo morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	texture pilosity
keyword	cotyledon plant embryo hairy seedling plant embryo cotyledon trichome
scope note
has exact match	cotyledon pilosity
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010113

Hypocotyl hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0010114
alternative label	seedling_hypocotyl_hairs
description	A seedling hypocotyl morphology trait [TO:0000769] which is the pilosity [PATO:0000066] of the hypocotyl [PO:0020100]. Binary variable distinguishing between glabrous versus hairy hypocotyls, where the hypocotyl is the embryonic axis to which the cotyledons are attached.
comments
value type	categorical variable
has narrower	seedling_hypocotyl_hairs_glabrous Hypocotyl lacks hairs. seedling_hypocotyl_hairs_hairy Hypocotyl has hairs.
has broader	plant embryo morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	texture pilosity
keyword	seedling hypocotyl morphology trait hypocotyl plant embryo hairy seedling trichome
scope note
has exact match	hypocotyl pilosity
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	hypocotyl_type
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010114

Seedling first true leaf type

Property	Value
URI	https://w3id.org/APD/traits/trait_0010160
alternative label	seedling_first_node_leaf_type
description	A plant embryo true leaf formation stage [PO:0001095] morphology trait which is whether the leaf [PO:0025034] at the first stem node [PO:0020141] above the epicotyl [PO:0020035] in a true leaf or a scale leaf. Binary variable distinguishing between seedlings where the first leaf is scale-like (cataphyll) versus leaf-like.
comments
value type	categorical variable
has narrower	seedling_first_node_leaf_type_scale-like The leaf at the first stem node above the epicotyl is scale-like, also referred to as a cataphyll. seedling_first_node_leaf_type_leaf-like The leaf at the first stem node above the epicotyl is leaf-like (broad).
has broader	plant embryo morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	organismal quality structure
keyword	plant embryo true leaf formation stage stem node epicotyl seedling
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seedling_first_leaf
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010160

Seedling first node leaf count

Property	Value
URI	https://w3id.org/APD/traits/trait_0010161
alternative label	seedling_first_node_leaf_count
description	A plant embryo true leaf formation stage [PO:0001095] morphology trait which is the leaf count [NCIT:C25463] at the first stem node [PO:0020141] above the epicotyl [PO:0020035]. Binary variable distinguishing between seedlings where the leaves at the first node are single versus paired.
comments
value type	categorical variable
has narrower	seedling_first_node_leaf_count_single There is a single leaf at the first stem node above the epicotyl. seedling_first_node_leaf_count_paired There is a pair of leaves at the first stem node above the epicotyl.
has broader	plant embryo morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	has number of count
keyword	plant embryo true leaf formation stage stem node epicotyl seedling
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	seedling_first_node
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010161

Seedling germination location

Property	Value
URI	https://w3id.org/APD/traits/trait_0010162
alternative label	seedling_germination_location
description	A plant embryo cotyledon morphology trait [TO:0000775] which is the position [PATO:0000140] of seed germination [GO:0009845] relative to the soil surface. Binary variable distinguishing between seedlings that germinate above versus below ground.
comments
value type	categorical variable
has narrower	seedling_germination_location_epigeal Germinant with one or more cotyledons emerging aboveground. seedling_germination_location_hypogeal Germinant with all cotyledons remaining belowground.
has broader	plant embryo morphology trait
plant structure plant structure	whole plant leaf
measured characteristic	organismal quality position
keyword	seed germination plant embryo seedling
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0010162

Leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011211
alternative label	leaf_area
description	A leaf area trait [TO:0000540] which is the 2-D [PATO:0001709] area [PATO:0001323] of the leaf lamina [PO:0020039] and petiole [PO:0020038]. For some species the area of a leaf analogue is reported, possibly a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335]. The 2-dimensional projected (scanned) area of the surface of a leaf or a leaf analogue (leaflet, cladode, phyllode).
comments	This trait includes measurements of leaves and leaf analogues, including leaflets, cladodes, and phyllodes. When a measurement is made on a leaf analogue, the tissue measured should be documented as part of the methods or noted as a context or under measurement_remarks. Leaflet area can alternatively be mapped to the trait leaflet_area. It is most common for leaf area measurements to include the petiole; the measurement methods should indicate if the petiole is included in or excluded from the leaf area measurements. For terete leaves, leaf area should still be recorded as the 2-dimensional projected area.
value type	continuous variable
unit	mm2 square millimetre
minAllowedValue	0.1
maxAllowedValue	1e+07
has broader	leaf size trait
plant structure	leaf
measured characteristic	area 2-D extent size
keyword	leaf lamina petiole leaflet cladode phyllode
scope note	Plants with leaves.
has exact match	leaf area trait Leaf area
has close match
has related match	leaf size
examples	exact match: Leaf area [TOP25] (https://top-thesaurus.org/index) exact match: Leaf area (in case of compound leaves: leaf, undefined if petiole in- or excluded) [TRY:3112] (https://www.try-db.org/de/de.php) close match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards) exact match: Leaf size [GIFT:4.4] (https://gift.uni-goettingen.de) exact match: leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Average leaf area [BROT:6] (http://doi.org/10.1038/sdata.2018.135)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011211

Leaflet area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011212
alternative label	leaflet_area
description	A leaf area trait [TO:0000540] which is the 2-D [PATO:0001709] area [PATO:0001323] of a single leaflet [PO:0020049] lamina [PO:0025060]. This measurement will not include the petiole [PO:0020038]. The 2-dimensional projected (scanned) area of the surface of the lamina of a single leaflet.
comments
value type	continuous variable
unit	mm2 square millimetre
minAllowedValue	0.1
maxAllowedValue	1e+06
has broader	leaf size trait
plant structure	leaf
measured characteristic	area 2-D extent size
keyword	leaflet leaf lamina
scope note
has exact match	leaflet area
has close match
has related match	leaf size
examples	close match: leaf area [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) close match: Average leaf area [BROT:6] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011212

Leaf length

Property	Value
URI	https://w3id.org/APD/traits/trait_0011213
alternative label	leaf_length
description	Length [PATO:0000122] of the leaf [PO:0025034] (including the petiole [PO:0020038]), from the leaf apex [PO:0020137] to the leaf axil [PO:0009023] (leaf base [PO:0020040]). Length of a leaf or a leaf analogue, from the leaf apex to the leaf axil (base), including the petiole.
comments	If a leaf analogue (leaflet, cladode, or phyllode) is measured instead of a true leaf, this should be noted as a context or under measurement_remarks.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.1
maxAllowedValue	1e+05
has broader	leaf size trait
plant structure	leaf
measured characteristic	length size
keyword	petiole leaf apex leaf base leaf lamina
scope note	Plants with leaves.
has exact match	leaf length
has close match
has related match	leaf size
examples	exact match: Leaf length [TOP13] (https://top-thesaurus.org/index) exact match: Leaf length [TRY:144] (https://www.try-db.org/de/de.php) exact match: Leaf length [GIFT:4.6] (https://gift.uni-goettingen.de) close match: maximum leaf length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) close match: minimum leaf length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) close match: Max_Blade_Length_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) close match: Max_Rachis_Length_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) close match: Max_Petiole_length_m [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011213

Leaf width

Property	Value
URI	https://w3id.org/APD/traits/trait_0011214
alternative label	leaf_width
description	The maximum [NCIT:C25564] width [PATO:0000921] axis of a leaf [PO:0025034] measured perpendicular to [PATO:0002434] the leaf length [PATO:0000122]. Maximum width axis of a leaf or a leaf analogue (a leaflet, cladode, or phyllode), measured perpendicular to the leaf length.
comments	This trait refers to the single line marking the maximum width of the leaf lamina. For a compound leaf the correct measure is the maximum width of the total leaf blade. It is the width measure that will be reported in taxonomic treatments. It is not the effective_leaf_width, a trait that records the diameter of the largest circle that can be drawn within the leaf lamina. If a leaf analogue (leaflet, cladode, or phyllode) is measured instead of a true leaf, this should be noted as a context or under measurement_remarks.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	1e+05
has broader	leaf size trait
plant structure	leaf
measured characteristic	width size
keyword	leaf lamina
scope note	Plants with leaves.
has exact match	leaf width
has close match
has related match	leaf size
examples	exact match: Leaf width [TOP17] (https://top-thesaurus.org/index) exact match: Leaf width [TRY:145] (https://www.try-db.org/de/de.php) exact match: Leaf width [GIFT:4.7] (https://gift.uni-goettingen.de) close match: maximum leaf width [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) close match: minimum leaf width [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011214

Leaf thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011215
alternative label	leaf_thickness
description	The thickness [PATO:0000915] of the leaf lamina [PO:0020039], measured away from the midrib [PO:0020139]. Thickness of the leaf lamina, generally measured away from the midrib.
comments	Leaf thickness measurements submitted from the ecological literature will generally avoid the midrib, but not avoid lower order leaf veins. In contrast, leaf thickness measurements from the taxonomic literature will usually include any veins or midribs, because in this literature leaf thickness is generally measured on succulent plants. The methods field should document whether the midrib was explicitly included, excluded, or ignored.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	51
has broader	leaf size trait
plant structure	leaf
measured characteristic	thickness size
keyword	leaf lamina
scope note	Plants with leaves.
has exact match	leaf thickness
has close match
has related match
examples	exact match: Leaf thickness [TOP20] (https://top-thesaurus.org/index) exact match: Leaf thickness [TRY:46] (https://www.try-db.org/de/de.php) exact match: Leaf thickness [GIFT:4.8] (https://gift.uni-goettingen.de) exact match: leaf thickness [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011215

Leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011216
alternative label	leaf_dry_mass
description	A leaf mass trait [TO:0000505] which is the dry [PATO:0001824] mass [PATO:0000125] of a leaf [PO:0025034] or leaf analogue (a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335]). The oven dry mass of a leaf or leaf analogue (leaflet, cladode, phyllode).
comments	This trait includes measurements of leaves and leaf analogues, including leaflets, cladodes, and phyllodes. When a measurement is made on a leaf analogue, the tissue measured should be documented as part of the methods or noted as a context or under measurement_remarks. Leaflet dry mass can alternatively be mapped to the trait leaflet_dry_mass. It is most common for leaf dry mass measurements to include the petiole; the measurement methods should indicate if the petiole is included in or excluded from the leaf dry mass measurements.
value type	continuous variable
unit	mg milligram
minAllowedValue	0.01
maxAllowedValue	15000
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass
keyword	leaf weight, leaf mass leaflet cladode phyllode
scope note
has exact match	leaf dry weight, leaf dry mass Leaf dry mass
has close match
has related match	leaf size
examples	exact match: Leaf dry mass [TOP40] (https://top-thesaurus.org/index) exact match: Leaf dry mass (single leaf) [TRY:55] (https://www.try-db.org/de/de.php) close match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards) exact match: leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011216

Leaflet dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011217
alternative label	leaflet_dry_mass
description	A leaf dry mass trait [TO:0001014] which is the dry mass of a single leaflet [PO:0020049]. This measurement will not include the petiole [PO:0020038]. The oven dry mass of a single leaflet.
comments
value type	continuous variable
unit	mg milligram
minAllowedValue	0.005
maxAllowedValue	15000
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass
keyword	leaflet leaf weight, leaf mass
scope note
has exact match
has close match	leaflet weight
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011217

Leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011218
alternative label	leaf_fresh_mass
description	A leaf mass trait [TO:0000505] which is the mass [PATO:0000125] of a fresh [EnvThes:21976] leaf [PO:0025034], indicating its mass at the time it is picked. For some species the mass of a leaf analogue is reported, possibly a leaflet [PO:0020049], cladode [PO:0025354], or phyllode [PO:0025335]. The fresh mass of a leaf or leaf analogue (leaflet, cladode, phyllode).
comments	This trait includes measurements of leaves and leaf analogues, including leaflets, cladodes, and phyllodes. When a measurement is made on a leaf analogue, the tissue measured should be documented as part of the methods or noted as a context or under measurement_remarks. It is most common for leaf fresh mass measurements to include the petiole; the measurement methods should indicate if the petiole is included in or excluded from the leaf fresh mass measurements. A source of variation for this trait will be how leaves are stored and if they are partially rehydrated before weighing; these details should be documented in the methods.
value type	continuous variable
unit	mg milligram
minAllowedValue	0.1
maxAllowedValue	30000
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass
keyword	leaf weight, leaf mass leaflet cladode phyllode
scope note	Plants with leaves
has exact match	leaf wet weight
has close match
has related match
examples	exact match: Leaf fresh mass [TOP35] (https://top-thesaurus.org/index) exact match: Leaf fresh mass [TRY:163] (https://www.try-db.org/de/de.php) exact match: leaf fresh mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011218

Petiole length

Property	Value
URI	https://w3id.org/APD/traits/trait_0011219
alternative label	petiole_length
description	Length [PATO:0000122] of the petiole [PO:0020038]), from the leaf base [PO:0020040] to the leaf axil [PO:0009023]. Length of the petiole from the leaf base to the leaf axil.
comments
value type	continuous variable
unit	cm
minAllowedValue	0
maxAllowedValue	1000
has broader	leaf size trait
plant structure	leaf
measured characteristic	length size
keyword	petiole
scope note
has exact match	petiole length Petiole length
has close match
has related match
examples
references	no linked references
date created	31/05/2023
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011219

Petiole width

Property	Value
URI	https://w3id.org/APD/traits/trait_0011220
alternative label	petiole_width
description	Width [PATO:0000921] of the petiole [PO:0020038]. Width of the petiole.
comments
value type	continuous variable
unit	cm
minAllowedValue	0
maxAllowedValue	200
has broader	leaf size trait
plant structure	leaf
measured characteristic	width size
keyword	petiole
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	31/05/2023
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011220

Number of leaflets per leaf

Property	Value
URI	https://w3id.org/APD/traits/trait_0011221
alternative label	leaflet_count
description	A leaf morphology trait [TO:0000748] which is the count [NCIT:C25463] of leaflets [PO:0020049] in a compound leaf [PO:0020043]. The count of leaflets that comprise a single leaf .
comments
value type	continuous variable
unit	{count}
minAllowedValue	1
maxAllowedValue	1000
has broader	leaf morphology trait
plant structure plant structure	leaf leaflet
measured characteristic	has number of count
keyword	leaflet compound leaf morphology trait compound leaf
scope note
has exact match	leaflet number
has close match
has related match
examples	exact match: Leaflet number per leaf [TRY:759] (https://www.try-db.org/de/de.php)
references	no linked references
date created	31/03/2025
date modified	31/03/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011221

Leaflet length

Property	Value
URI	https://w3id.org/APD/traits/trait_0011222
alternative label	leaflet_length
description	A leaf morphology trait [TO:0000748] which is length [PATO:0000122] of a leaflet [PO:0020049], from the leaflet apex [PO:0020137] to where the leaflet attaches to the leaf [PO:0025034] rachis [PO:020055]. The length of a single leaflet comprising a botanical leaf.
comments	This trait is intended to only apply to compound leaves, those with multiple leaflets per botanical leaf.
value type	continuous variable
unit	cm
minAllowedValue	0.1
maxAllowedValue	10
has broader	leaf size trait
plant structure plant structure	leaf leaflet
measured characteristic	length size
keyword	leaflet compound leaf morphology trait compound leaf
scope note	applicable to plants with compound leaves.
has exact match	leaflet length
has close match
has related match
examples	exact match: Leaflet_length [TOP14] (https://top-thesaurus.org/index) exact match: Leaflet length [TRY:985] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011222

Leaflet width

Property	Value
URI	https://w3id.org/APD/traits/trait_0011223
alternative label	leaflet_width
description	A leaf morphology trait [TO:0000748] which is the maximum [NCIT:C25564] width [PATO:0000921] axis of a single leaflet [PO:0020049] measured perpendicular to [PATO:0002434] the leaflet length [PATO:0000122]. Maximum width axis of a single leaflet, measured perpendicular to the leaflet length.
comments	This trait is intended to only apply to compound leaves, those with multiple leaflets per botanical leaf.
value type	continuous variable
unit	cm
minAllowedValue	0.1
maxAllowedValue	10
has broader	leaf size trait
plant structure plant structure	leaf leaflet
measured characteristic	width size
keyword	leaflet compound leaf morphology trait compound leaf
scope note	applicable to plants with compound leaves.
has exact match	leaflet width
has close match
has related match
examples	exact match: Leaflet_width [TOP18] (https://top-thesaurus.org/index) exact match: Leaflet width [TRY:986] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011223

Leaf mass per area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011230
alternative label	leaf_mass_per_area
description	A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of the leaf dry [PATO:0001824] mass [PATO:0000125] to the leaf [PO:0025034] area [PATO:0001323], measured on an entire leaf, including the entire leaf lamina [PO:0020039] and the petiole [PO:0020038]. The ratio of leaf dry mass to leaf area, measured on an entire leaf including the petiole; LMA. (The inverse of specific leaf area, SLA.)
comments	Leaf mass per area includes measurements submitted as specific_leaf_area (SLA); they are simply inverted. Leaf mass per area is explicitly measured on an entire leaf, including the petiole. Measurements made on a piece of the leaf lamina, excluding petiole and midrib, are a separate trait leaf_lamina_mass_per_area. If measurements are made on the entire leaf lamina, but the petiole is excluded, this should be noted in the methods or under measurement_remarks.
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	1
maxAllowedValue	3000
has broader	leaf size trait leaf mass trait
plant structure	leaf
measured characteristic	area ratio; proportion mass per unit area area density
keyword	leaf weight, leaf mass biomass allocation leaf lamina petiole
scope note
has exact match	specific leaf area Leaf mass per leaf area
has close match
has related match
examples	exact match: Leaf mass per area [TOP53] (https://top-thesaurus.org/index) close match: Specific leaf area [TOP50] (https://top-thesaurus.org/index) exact match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): undefined if petiole is in- or exclu [TRY:3117] (https://www.try-db.org/de/de.php) exact match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole included [TRY:3116] (https://www.try-db.org/de/de.php) close match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards) exact match: Specific Leaf Area (SLA) [GIFT:4.1] (https://gift.uni-goettingen.de) exact match: leaf area per leaf dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Average specific leaf area [BROT:8] (http://doi.org/10.1038/sdata.2018.135)
references	Pérez-Harguindeguy 2013 Wright 2004
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	specific_leaf_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011230

Leaf lamina mass per area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011231
alternative label	leaf_lamina_mass_per_area
description	A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of the leaf dry [PATO:0001824] mass [PATO:0000125] to leaf [PO:0025034] area [PATO:0001323], measured on a piece of the leaf lamina [PO:0020039] located away from major leaf lamina veins [PO:0020138]. The ratio of leaf dry mass to leaf area, measured on a piece of the leaf lamina that does not include the petiole or major veins.
comments	This trait only includes data made on subsamples of a leaf, usually a leaf punch sample associated with leaf strength measurements or from the herbivory literature. For whole-leaf LMA/SLA data, see the trait leaf_mass_per_area
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	0.1
maxAllowedValue	500
has broader	leaf size trait leaf mass trait
plant structure	leaf
measured characteristic	area ratio; proportion mass per unit area area density
keyword	leaf weight, leaf mass biomass allocation leaf lamina
scope note	Plants with leaves.
has exact match
has close match
has related match	specific leaf area
examples	related match: Leaf mass per area [TOP53] (https://top-thesaurus.org/index) related match: Specific leaf area [TOP50] (https://top-thesaurus.org/index) exact match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole excluded [TRY:3115] (https://www.try-db.org/de/de.php) close match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA) petiole, rhachis and midrib excluded [TRY:3086] (https://www.try-db.org/de/de.php) related match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): undefined if petiole is in- or exclu [TRY:3117] (https://www.try-db.org/de/de.php) related match: Leaf area per leaf dry mass (specific leaf area, SLA or 1/LMA): petiole included [TRY:3116] (https://www.try-db.org/de/de.php) related match: Leaf size SLA, leaf mass & LDMC [LEDA:2.1] (https://uol.de/en/landeco/research/leda/standards)
references	Pérez-Harguindeguy 2013
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011231

Leaf tissue density

Property	Value
URI	https://w3id.org/APD/traits/trait_0011232
alternative label	leaf_density
description	The ratio [PATO:0001470] of leaf [PO:0025034] dry [PATO:0001824] mass [PATO:0000125] to leaf fresh [EnvThes:21976] volume [PATO:0000918]. The ratio of leaf dry mass to leaf fresh volume.
comments
value type	continuous variable
unit	mg/mm3 milligram per cubic millimetre
minAllowedValue	0.05
maxAllowedValue	1.1
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass density, density mass ratio; proportion
keyword	leaf weight, leaf mass biomass allocation
scope note
has exact match	leaf mass density
has close match
has related match
examples	exact match: Leaf density [TOP56] (https://top-thesaurus.org/index) exact match: Leaf density (leaf tissue density, leaf dry mass per leaf volume) [TRY:48] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011232

Leaf area ratio (LAR)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011260
alternative label	leaf_area_ratio
description	A leaf area trait [TO:0000540] which is the ratio [PATO:0001470] of a plant’s [PO:0000003] total [NCIT:C25304] leaf [PO:0025034] area [PATO:0001323] to total plant dry mass [TO_0000352]. The ratio of total plant leaf area to total plant dry mass; LAR.
comments	Since the total plant dry mass is required for leaf area ratio measurements, this is standardly measured only on young plants or plants grown for experiments where the entire plant can be harvested. It is best practice for leaf area measurements to include the petiole.
value type	continuous variable
unit	mm2/mg square millimetre per milligram
minAllowedValue	0.05
maxAllowedValue	100
has broader	leaf size trait
plant structure plant structure	leaf whole plant
measured characteristic	area size ratio; proportion
keyword	whole plant dry mass biomass allocation
scope note	Plants with leaves.
has exact match	leaf area ratio
has close match	Leaf area
has related match
examples	exact match: Leaf area ratio [TOP88] (https://top-thesaurus.org/index) exact match: Leaf area per plant dry mass (leaf area ratio, LAR) [TRY:109] (https://www.try-db.org/de/de.php)
references	Poorter & Remkes 1990
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011260

Leaf mass fraction

Property	Value
URI	https://w3id.org/APD/traits/trait_0011261
alternative label	leaf_mass_fraction
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of a plant’s total leaf dry mass [APD:glossary/40080] to total plant dry mass [TO:0000352]. The ratio of leaf dry mass to total plant dry mass.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	1
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	leaf weight, leaf mass biomass allocation whole plant dry mass whole plant stem dry mass
scope note
has exact match
has close match
has related match
examples	exact match: Leaf mass ratio [TOP85] (https://top-thesaurus.org/index) exact match: Plant biomass and allometry: Leaf dry mass per plant dry mass (leaf weight ratio, LWR) [TRY:110] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011261

Leaf dry matter content (LDMC)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011262
alternative label	leaf_dry_matter_content
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of leaf dry mass [TO:0001014] to leaf [PO:0025034] fresh mass [APD:glossary/40044]. The ratio of leaf dry mass to leaf fresh mass; LDMC.
comments	Two similar traits are leaf_water_content_per_dry_mass, the ratio of leaf water content to leaf dry mass and leaf_water_content_per_fresh_mass, the ratio of leaf water content to leaf fresh mass.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.001
maxAllowedValue	2
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	leaf weight, leaf mass biomass allocation fresh mass
scope note
has exact match
has close match
has related match
examples	exact match: Leaf dry matter content [TOP45] (https://top-thesaurus.org/index) exact match: Leaf dry mass per leaf fresh mass (leaf dry matter content, LDMC) [TRY:47] (https://www.try-db.org/de/de.php) exact match: Leaf dry matter content (LDMC) [GIFT:4.9] (https://gift.uni-goettingen.de) exact match: leaf dry mass per leaf fresh mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Leaf dry matter content [BROT:14] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011262

Leaf fresh mass per leaf area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011263
alternative label	leaf_fresh_mass_per_area
description	A leaf area trait [TO:0000540] which is the ratio [PATO:0001470] of the leaf [PO:0025034] fresh mass [APD:glossary/40044]. to the leaf area [PATO:0001323], measured on an entire leaf, including the entire leaf lamina [PO:0020039] and the petiole [PO:0020038]. The ratio of leaf fresh mass to leaf area, measured on an entire leaf including the petiole.
comments	Leaf fresh mass per area is explicitly measured on an entire leaf, including the petiole.
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	200
maxAllowedValue	2000
has broader	leaf size trait leaf mass trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	leaf weight, leaf mass biomass allocation fresh mass leaf lamina petiole
scope note	Plants with leaves.
has exact match
has close match
has related match
examples	exact match: Leaf area per leaf fresh mass (specific leaf area (SLA or 1/LMA) based on leaf fresh mass) [TRY:125] (https://www.try-db.org/de/de.php)
references	Pérez-Harguindeguy 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011263

Leaf water content per unit leaf area (leaf succulence)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011264
alternative label	leaf_water_content_per_area
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf area [PATO:0001323]. The ratio of the mass of water in a fresh leaf (leaf water content) to leaf surface area; leaf succulence.
comments
value type	continuous variable
unit	g/m2 gram per square metre
minAllowedValue	10
maxAllowedValue	5000
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass per unit area ratio; proportion
keyword	leaf weight, leaf mass water leaf water content
scope note
has exact match
has close match	tissue succulence
has related match	leaf moisture content
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011264

Leaf water content per unit leaf dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011265
alternative label	leaf_water_content_per_dry_mass
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf dry mass [TO:0001014]. The ratio of the mass of water in a fresh leaf (leaf water content) to leaf dry mass.
comments	A similar trait that is recorded by a greater number of studies is leaf_dry_matter_content, the ratio of a leaf dry mass to leaf fresh mass.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	20
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	leaf weight, leaf mass water leaf water content
scope note
has exact match
has close match
has related match	leaf moisture content
examples	exact match: Leaf water content per leaf dry mass (not saturated) [TRY:3120] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011265

Leaf water content per unit leaf fresh mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011266
alternative label	leaf_water_content_per_fresh_mass
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a fresh [EnvThes:21976] leaf [PO:0025034] to leaf fresh mass [APD:glossary/40044]. The ratio of the mass of water in a fresh leaf (leaf water content) to leaf fresh mass
comments	A similar trait that is recorded by a greater number of studies is leaf_dry_matter_content, the ratio of a leaf dry mass to leaf fresh mass.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	leaf weight, leaf mass water leaf water content fresh mass
scope note
has exact match
has close match
has related match	leaf moisture content
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011266

Leaf water content per unit saturated leaf mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011267
alternative label	leaf_water_content_per_saturated_mass
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a saturated leaf [PO:0025034] to the mass of a saturated leaf. A saturated leaf is one that has been hydrated to its maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620]. The ratio of the mass of water in a leaf (leaf water content) to the mass of a saturated leaf. A saturated leaf is one that has been hydrated to its maximal water holding capacity and is at full turgor.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.1
maxAllowedValue	10
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	leaf weight, leaf mass water leaf water content fresh mass water holding capacity turgor
scope note
has exact match
has close match
has related match	leaf moisture content
examples	close match: Leaf water content per leaf dry mass (at saturation) [TRY:3122] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011267

Leaf cell wall fraction

Property	Value
URI	https://w3id.org/APD/traits/trait_0011268
alternative label	leaf_cell_wall_fraction
description	A leaf mass trait [TO:0000505] which is the ratio [PATO:0001470] of cell wall [GO:0009505] dry mass to leaf dry mass [TO:0001014]. The ratio of leaf cell wall dry mass to leaf dry mass. This is the fraction of cell wall material that can be recovered from total leaf biomass.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.1
maxAllowedValue	1
has broader	leaf mass trait
plant structure	leaf
measured characteristic	mass mass fraction ratio; proportion
keyword	leaf weight, leaf mass plant-type cell wall biomass allocation cell
scope note
has exact match
has close match
has related match
examples	exact match: Leaf cell wall mass per leaf dry mass [TRY:3375] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011268

Leaf type

Property	Value
URI	https://w3id.org/APD/traits/trait_0011310
alternative label	leaf_type
description	A leaf shape trait [TO:0000492] which is inclusive [NCIT:C166400] categories describing the shape [PATO:0000052] of a leaf [PO:0025034] as broad, needle-like, or scale-like. Variable distinguishing between general leaf form (shape) categories.
comments	Terms identifying the plant organ responsible for photosynthesis are mapped to the trait plant_photosynthetic_organ, while leaf_form_general only includes terms pertaining to the general form of the leaf. The trait leaf_shape provides detailed vocabulary to describe a leaf’s shape.
value type	categorical variable
has narrower	leaf_type_broadleaf Flat leaf lamina, regardless of whether the leaves are or are not toothed, lobed, or compound. leaf_type_needle Needle or awl-shaped leaf that is generally 3-dimensional. leaf_type_scale Leaf reduced to a small scale. leaf_type_leafless An adult stage plant that lacks leaves throughout the year.
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape structure
keyword	leaf shape
scope note	none
has exact match
has close match
has related match
examples	related match: Leaf form [TOP225] (https://top-thesaurus.org/index) related match: Leaf type [TRY:43] (https://www.try-db.org/de/de.php) exact match: Leaf_form_1 [GIFT:4.11.1] (https://gift.uni-goettingen.de) close match: Basic leaf shape [BROT:5] (http://doi.org/10.1038/sdata.2018.135)
references	Pérez-Harguindeguy 2013 Burrows 2001
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011310

Leaf shape

Property	Value
URI	https://w3id.org/APD/traits/trait_0011311
alternative label	leaf_shape
description	A leaf shape trait [TO:0000492] which is the shape [PATO:0000052] and form of a leaf [PO:0025034], considering the ratio [PATO:0001470] of leaf length [PATO:0000122] to leaf width [PATO:0000921], the position [PATO:0000140] of the maximum [NCIT:C25564] width dimension relative to the length axis [NCIT:C25154], and the curvature [PATO:0001591] of the leaf margin [PO:0020128]. Shapes of a leaf or leaflet.
comments	This trait includes terms to describe the shape of leaves, regardless of if they are broad leaves, scale-like leaves, or needle-like leaves. For compound leaves, it is generally the shape of the leaflet that is recorded. See the trait leaf_type for broad terms referencing leaf form (broadleaf, needle, scale) and plant_photosynthetic_organ for terms indicating which plant organ or tissue functions as the plant’s primary photosynthetic organ (e.g. phyllode, cladode, article).
value type	categorical variable
has narrower	leaf_shape_linear A 2-dimensional shape that is very long and narrow, has parallel sides and a length:width ratio between 12:1 and 6:1. leaf_shape_narrowly_oblong A 2-dimensional shape that is long and narrow, has parallel sides and a length:width ratio between 6:1 and 3:1. (Synonyms, lorate) leaf_shape_oblong A 2-dimensional shape that is somewhat elongated with parallel sides and a length:width ratio between 2:1 and 3:2. leaf_shape_linear_elliptical A 2-dimensional shape that is a very narrow oval, with curved margins, lacking a distinct point, and a length:width ratio between 12:1 and 6:1. leaf_shape_narrowly_elliptical A 2-dimensional shape that is a narrow oval, with curved margins, lacking a distinct point, and a length:width ratio between 6:1 and 3:1. leaf_shape_elliptical A 2-dimensional shape that is an oval, with curved margins, lacking a distinct point, and a length:width ratio between 2:1 and 3:2. (Synonyms, oval) leaf_shape_widely_elliptical A 2-dimensional shape that is a broad oval, with curved margins, lacking a distinct point, and a length:width ratio of approximately 6:5. leaf_shape_orbicular A 2-dimensional shape that is approximately circular. (Synonyms, circular) leaf_shape_oblate A 2-dimensional shape that is a broad oval, with curved margins, lacking a distinct point, and a length:width ratio of approximately 5:6. The petiole attaches along the broader side. leaf_shape_narrowly_lanceolate A 2-dimensional shape that is very narrowly egg-shaped or teardrop-shaped, with a length:width ratio between 12:1 and 6:1. The petiole attaches near the wide, curved portion of the leaf lamina. leaf_shape_lanceolate A 2-dimensional shape that is narrowly egg-shaped or teardrop-shaped, with a length:width ratio between 6:1 and 3:1. The petiole attaches near the wide, curved portion of the leaf lamina. leaf_shape_narrowly_ovate A 2-dimensional shape that is narrowly egg-shaped, with a length:width ratio between 3:1 and 2:1. The petiole attaches near the wide, curved portion of the leaf lamina. (Synonyms, lance_ovate, widely_lanceolate) leaf_shape_ovate A 2-dimensional shape that is egg-shaped, with a length:width ratio between 2:1 and 3:2. The petiole attaches near the wide, curved portion of the leaf lamina. leaf_shape_widely_ovate A 2-dimensional shape that is broadly egg-shaped with a length:width ratio of approximately 6:5. The petiole attaches near the wide, curved portion of the leaf lamina. leaf_shape_narrowly_oblanceolate A 2-dimensional shape that is very narrowly teardrop-shaped, with a length:width ratio between 12:1 and 6:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. (Synonyms, linear_oblanceolate) leaf_shape_oblanceolate A 2-dimensional shape that is narrowly teardrop-shaped, with a length:width ratio between 6:1 and 3:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. leaf_shape_narrowly_obovate A 2-dimensional shape that is narrowly egg-shaped or teardrop-shaped with a length:width ratio between 3:1 and 2:1. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. (Synonyms, oblance-obovate) leaf_shape_obovate A 2-dimensional shape that is egg-shaped with a length:width ratio between 2:1 and 3:2. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. leaf_shape_widely_obovate A 2-dimensional shape that is broadly egg-shaped with a length:width ratio of approximately 6:5. The leaf lamina has curved outer margins and narrows nearly to a point where it attaches to the petiole. leaf_shape_narrowly_triangular A 2-dimensional shape that is narrowly triangular to wedge-shaped, with a length:width ratio 6:1 to 3:1 and the petiole attaching near the widest portion of the leaf. (Synonyms, triangular) leaf_shape_triangular A 2-dimensional shape that is triangular to wedge-shaped, with a length:width ratio 2:1 to 3:2 and the petiole attaching near the widest portion of the leaf. leaf_shape_deltate A 2-dimensional shape that is triangular to wedge-shaped, with a length:width ratio of 1:1 and the petiole attaching along one of the broad sides. leaf_shape_obtriangular A 2-dimensional shape that is triangular to fan-shaped or wedge-shaped, with a length:width ratio 2:1 to 3:2 and the petiole attaching near the narrowest portion of the leaf. (Synonyms, flabelliform, fan-shaped, wedge-shaped, cuneate) leaf_shape_narrowly_rhomboidal A 2-dimensional shape that is like a narrow diamond, with approximately straight edges, widest near the middle, and a length:width ratio 6:1 to 3:1. The petiole attaches near one of the points. leaf_shape_rhomboidal A 2-dimensional shape that is approximately diamond or parallelogram-shaped, with approximately straight edges, widest near the middle, and a length:width ratio 2:1 to 3:2. The petiole attaches near one of the points. (Synonyms, rhombic, diamond) leaf_shape_trullate A 2-dimensional shape that is approximately parallelogram-shaped, widest near where the petiole attaches (base), and with a length:width ratio 2:1 to 3:2. leaf_shape_narrowly_obtrullate A 2-dimensional shape that is narrowly parallelogram-shaped, widest near the tip, and with a length:width ratio 3:1 to 6:1. leaf_shape_obtrullate A 2-dimensional shape that is approximately parallelogram-shaped, widest near the tip, and with a length:width ratio 2:1 to 3:2. leaf_shape_widely_obtrullate A 2-dimensional shape that is broadly parallelogram-shaped, widest near the tip, and with a length:width ratio of approximately 6:5. leaf_shape_acicular A 2-dimensional or 3-dimensional shape with a needlelike tip, often round in cross-section, with margins straight and parallel, and a length:width ratio greater than 12:1. (Synonyms, needle-shaped) leaf_shape_terete A 3-dimensional shape that is a long, narrow cylinder approximately circular in cross-section. Terete leaves often taper toward the tip. (Synonym, cylindrical) leaf_shape_cordate A 2-dimensional heart shape, approximately ovate with a cordate base, with the petiole attaching at the notch between the lobes. (Synonyms, cordiform) leaf_shape_obcordate A 2-dimensional heart shape, approximately ovate with a cordate apex, and the petiole attaching along the broad, curve side. leaf_shape_falcate A 2-dimensional sickle-shape, approximately lanceolate to linear and curved on one side. (Synonyms, scimitar-shaped, sickle-shaped, falciform, crescent-shaped) leaf_shape_reniform A 2-dimensional shape that is roughly kidney-shaped, wider than long with a rounded apex and two rounded, basal lobes that are smoothly concave at their intersection. (Synonyms, kidney-shaped) leaf_shape_spathulate A 2-dimensional shape that can be approximately oblong, obovate, or oblanceolate in outline with a long attenuate base, like a spoon. (Synonyms, spoon-shaped, clavate) leaf_shape_strap-shaped A 2-dimensional shape that is strap-shaped with a rounded tip and a length:width ratio greater than 12:1. leaf_shape_ensiform A 2-dimensional shape that is sword-shaped with a pointed tip and a length:width ratio greater than 12:1. (Synonyms, sword-shaped) leaf_shape_subulate A 2-dimensional shape that is awl-shaped, approximately narrowly oblong to narrowly triangular with a broad base. (Synonyms, awl-shaped) leaf_shape_peltate A 2-dimensional orbicular shape, distinguished by the petiole attaching near the center of the leaf lamina. leaf_shape_filiform A 2-dimensional shape that is long, thin, and typically flexuous, threadlike, filamentous.
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape length curvature structure
keyword	leaf shape leaf margin
scope note	Plants with leaves.
has exact match	leaf shape
has close match
has related match	leaflet shape
examples	close match: Leaf form [TOP225] (https://top-thesaurus.org/index) close match: Leaf shape [TRY:154] (https://www.try-db.org/de/de.php)
references	Simpson 2019 Ellis 2009 Systematics Association Committee for Descriptive Terminology 1962
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011311

Leaf base shape

Property	Value
URI	https://w3id.org/APD/traits/trait_0011312
alternative label	leaf_base_shape
description	A leaf shape trait [TO:0000492] which is the variation in shapes and forms of a leaf base [PO:0020040], especially considering the curvature [PATO:0001591] of the leaf margin [PO:0020128] near where it attaches to the petiole [PO:0020038]. The shape of the basal margin of a leaf.
comments
value type	categorical variable
has narrower	leaf_base_shape_acute Two sides of the leaf meet at an angle that is less than 90<U+00B0>. leaf_base_shape_attenuate Basal leaf margin abruptly incurved (concave), the two sides intersecting at an angle of less than 45<U+00B0>. leaf_base_shape_auriculate Basal leaf margin with two rounded lobes. The leaf margin above the lobes is concave. The petiole attaches in the incised region between the lobes. leaf_base_shape_cordate Basal leaf margin with two rounded lobes and the petiole attaches in the incised region between the lobes. The leaf margin above the lobes is rounded. leaf_base_shape_cuneate Basal leaf margin approximately straight, the two sides intersecting at an angle between 45<U+00B0> to 90<U+00B0>. leaf_base_shape_hastate Basal leaf margin with two lobes oriented outward, such that they are at approximately 90<U+00B0> relative to the central axis and the leaf margin is fairly flat at the base. leaf_base_shape_oblique The leaf base is asymmetrical, with the midrib not lying in the middle of the leaf where it attaches to the petiole. leaf_base_shape_obtuse Two sides of the leaf meet at an angle that is greater than 90<U+00B0>. leaf_base_shape_rounded Basal leaf margin is rounded. leaf_base_shape_sagittate Basal leaf margin with two lobes oriented downward, away from apex. The petiole attaches in the concave region between the lobes. leaf_base_shape_sheathing The leaf base is broad and envelops the stem. leaf_base_shape_truncate The basal leaf margin is cut straight across with the two sides intersecting at approximately 180<U+00B0>.
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape curvature
keyword	leaf shape leaf base leaf margin petiole
scope note	Plants with leaves.
has exact match	Lamina shape
has close match
has related match
examples
references	Simpson 2019 Ellis 2009
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011312

Leaf margin

Property	Value
URI	https://w3id.org/APD/traits/trait_0011313
alternative label	leaf_margin
description	A leaf shape trait [TO:0000492] which is the edge shape [PATO:0002267] of the leaf margin [PO:0020128], in particular the presence or absence of leaf lamina teeth [PO:0025518] and the shape [PATO:0000052] of the leaf lamina teeth (serrate, dentate, or crenate). Shape of the leaf margin, referencing incisions or projections in the outermost 1/8 of the leaf lamina.
comments	Terms describing leaf lamina projections and incisions that are greater than 1/8 of the distance to the midrib are captured under the traits leaf_lobation and leaf_division_detailed.
value type	categorical variable
has narrower	leaf_margin_entire Leaf margin lacks projections or indentations that extend less than 1/8 of the distance to the midrib. leaf_margin_toothed Leaf margin has small projections or lobes along the edge of the leaf lamina. This is a broad term that includes serrate, dentate, and crenulate. leaf_margin_toothed_serrate Type of toothed leaf margin with sharp teeth that are angled toward the tip of the leaf. (Synonyms, saw-toothed) leaf_margin_toothed_dentate Type of toothed leaf margin with sharp teeth that point outwards at right angles to the leaf margin. leaf_margin_toothed_crenate Type of toothed leaf margin with teeth that are rounded or obtuse, rather than having a sharp apex. [PO:0025519]
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape edge shape curvature
keyword	leaf shape leaf margin leaf lamina tooth
scope note	Plants with broad leaves.
has exact match	leaf margin morphology trait
has close match	leaf margin serrated
has related match
examples	exact match: Leaf margin type [TRY:963] (https://www.try-db.org/de/de.php) exact match: Leaf_margin_1 [GIFT:4.12.1] (https://gift.uni-goettingen.de)
references	Simpson 2019 Ellis 2009 Hickey 1973
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011313

Leaf margin posture

Property	Value
URI	https://w3id.org/APD/traits/trait_0011314
alternative label	leaf_margin_posture
description	A leaf shape trait [TO:0000492] which is the degree of curvature [PATO:0001591] or curling [PATO:0000405] of the leaf margin [PO:0020128] along the side of the leaf, at the leaf apex [PO:0020137], and at the leaf base [PO:0020040]. The posture (curvature or curling) of the leaf margin.
comments	This trait captures the posture of the leaf margin, where posture refers to the placement of different parts of the leaf’s margin relative to a flat plane. This trait’s definition and allowable levels will be further refined once more data are available to map onto possible trait values.
value type	categorical variable
has narrower	leaf_margin_posture_involute Leaf margins along the side of the leaf are rolled upward, such that the margin of the adaxial (upper) leaf surface is rolled inward. (Synonym, inrolled, incurved) leaf_margin_posture_revolute Leaf margins along the side of the leaf are rolled under or downward, such that the margin of the abaxial (lower) leaf surface is rolled inward. [PATO:0001963] (Synonym, recurved) leaf_margin_posture_flat Leaf lamina, including margins, is a 2-dimensional plane. [PATO:0000407] leaf_margin_posture_undulate Leaf margins are wavy in their 2-dimensional plane. (Synonyms, wavy, rippled, sinuate)
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape curvature
keyword	leaf shape curled leaf margin leaf apex leaf base
scope note	Plants with leaves
has exact match
has close match
has related match
examples
references	Simpson 2019
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011314

Leaf lobation

Property	Value
URI	https://w3id.org/APD/traits/trait_0011315
alternative label	leaf_lobation
description	A leaf shape trait [TO:0000492] which is the presence or absence of leaf lamina lobes [PO:0025517] along the leaf margin [PO:0020128] and how far the lobes extend toward the midrib [PO:0020139]. A trait that indicates whether a leaf lamina or leaflets display any divisions or dissections on a 2-dimensional plane that extend greater than 1/8 of the distance to the midrib.
comments	For terms pertaining to 2-dimensional leaf lamina divisions and incisions that are less than 1/8 of the distance to the midrib, see the trait leaf_margin. See also the trait leaf_division for more detailed leaf lobation (and leaf compoundness) trait values. Note that there might be some species that grade from deeply lobed simple leaves to compound leaves.
value type	categorical variable
has narrower	leaf_lobation_unlobed Leaf margin lacks projections or indentations that extend more than 1/8 of the distance to the midrib. leaf_lobation_lobed Leaf margin has incisions, either rounded or sharp-tipped, that extend more than 1/8 of the distance to the midrib. leaf_lobation_lobed_deep Leaf margin with lobes that extend more than halfway to the midrib. (Synonyms, parted, divided) leaf_lobation_lobed_shallow Leaf margin with lobes that extend less than halfway to the midrib. (Synonyms, cleft, sinuate)
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape edge shape curvature
keyword	leaf shape leaf lamina lobes leaf margin leaf midvein, midrib
scope note	Plants with well-developed leaves.
has exact match	leaf lamina lobes
has close match
has related match
examples
references	Simpson 2019 Ellis 2009 Hickey 1973 Wikipedia 2022 Glossary of leaf morphology
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011315

Leaf compoundness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011316
alternative label	leaf_compoundness
description	A leaf shape trait [TO:0000492] which is whether a leaf [PO:0025034] is a simple leaf [PO:0020042] or is divided into leaflets [PO:0020049] making it a compound leaf [PO:0020043]. A binary trait that indicates whether a leaf lamina is simple or divided into discontinuous leaflets (compound).
comments	See also the trait ‘leaf_division’ for more detailed leaf compoundness trait values. Note that there might be some species that grade from deeply lobed simple leaves (see trait leaf_lobation) to compound leaves.
value type	categorical variable
has narrower	leaf_compoundness_simple A leaf with a single undivided blade. [PO:0020042] leaf_compoundness_compound A leaf that is divided into multiple leaflets. [PO:0020043]
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape spatial pattern, distribution
keyword	leaf shape simple leaf leaflet compound leaf leaf midvein, midrib
scope note	Plants with well-developed leaves.
has exact match
has close match
has related match
examples	exact match: Leaf compoundness [TOP222] (https://top-thesaurus.org/index) exact match: Leaf compoundness [TRY:17] (https://www.try-db.org/de/de.php) exact match: leaf compoundness [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Leaf division degree [BROT:2] (http://doi.org/10.1038/sdata.2018.135)
references	Ellis 2009
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011316

Leaf divisions

Property	Value
URI	https://w3id.org/APD/traits/trait_0011317
alternative label	leaf_lamina_division
description	A leaf shape trait [TO:0000492] which indicates types [NCIT:C25284] of lobed leaves [PO:0025517] and compound leaves [PO:0020043], including the arrangement [NCIT:C61532] of the lobes or leaflets [PO:0020049] with respect to the midrib [PO:0020139] or other primary leaf veins [PO:0025413] and the number of orders of axes [NCIT:C25154] across which there are incisions or divisions. A trait that captures the types of lobed leaves and compound leaves, including the arrangement of the lobes or leaflets relative to the midrib or other primary veins and the number of orders of axes across which there are incisions or divisions.
comments	This trait provides detailed description of leaf divisions, beyond what is captured in leaf_compoundness (simple versus compound) and leaf_lobation (entire versus lobed). The term ‘divided’ is meant to indicate lobing, such as ‘trichotomously divided’, but as ‘divided’ is also used in some context to indicate the divisions of leaves into leaflets, it is avoided as a trait value for this trait.
value type	categorical variable
has narrower	leaf_lamina_division_pinnately_lobed Leaf with divisions that are symmetrical across a central axis, usually the midvein. leaf_lamina_division_palmately_lobed Leaf with divisions that radiate from a single point, usually the basal junction of major veins. leaf_lamina_division_pinnatifid Leaf with lobes that extend less than half-way to the midrib. leaf_lamina_division_bipinnatifid Leaf that is twice pinnatifid, with two series of lobes that extend less than half-way to the midrib. leaf_lamina_division_pinnatipartite Leaf with lobes that extend more than half-way to the midrib. leaf_lamina_division_pinnatisect Leaf with lobes that extend nearly to the midrib but the divisions are not separated into distinct leaflets. leaf_lamina_division_bipinnatipartite Leaf that is twice pinnatipartite, with two series of lobes that extend more than half-way to the midrib. leaf_lamina_division_bipinnatisect Leaf that is twice pinnatisect, with two series of lobes that extend nearly to the midrib but the divisions are not separated into distinct leaflets. leaf_lamina_division_trichotomously_lobed Leaf divided repeatedly into three lobes. leaf_lamina_division_dichotomously_lobed Leaf divided repeatedly into two lobes. leaf_lamina_division_bipartite Leaf divided into two lobes. leaf_lamina_division_tripartite Leaf divided into three lobes. leaf_lamina_division_pinnately_compound Leaf divided into leaflets that attach along a central axis. [PATO_0000410] leaf_lamina_division_palmately_compound Leaf divided into leaflets that attach to a central point. (Synonyms, digitate) leaf_lamina_division_bipinnate Leaf that is twice pinnate, having pinnate leaflets that are themselves pinnately divided. leaf_lamina_division_tripinnate Leaf that is three times pinnate, with pinnate leaflets that are twice more pinnately divided. leaf_lamina_division_trifoliate Leaf divided into three leaflets. (Synonyms, ternate, trifoliolate)
has broader	leaf shape trait
plant structure	leaf
measured characteristic	spatial pattern, distribution shape arrangement
keyword	leaf shape leaf lamina lobes compound leaf leaflet leaf midvein, midrib leaf margin pinnate
scope note	Plants with broad leaves.
has exact match
has close match
has related match
examples	related match: Leaf compoundness [TOP222] (https://top-thesaurus.org/index) related match: Leaf compoundness [TRY:17] (https://www.try-db.org/de/de.php)
references	Simpson 2019 Ellis 2009 Hickey 1973
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	leaf_division
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011317

Leaf lamina posture (leaf 3-dimensionality)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011318
alternative label	leaf_posture_numeric
description	A leaf shape trait [TO:0000492] which is the maximum [NCIT:C25564] perpendicular [PATO:0002434] height [PATO:0000119] of a leaf [PO:0025034] placed on a flat surface, indicating the 3-D extent [PATO:0001710] of the leaf. This generally corresponds with whether the leaf is curled [FLOPO:0010468]. The maximum perpendicular height of a leaf when positioned on a flat surface.
comments	A measure of how planar versus 3-dimensional a leaf is. The term posture refers to the placement of different parts of the leaf’s lamina relative to a flat plane.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0
maxAllowedValue	100
has broader	leaf shape trait
plant structure	leaf
measured characteristic	height shape 3-D extent
keyword	leaf shape leaf lamina
scope note	Plants with leaves
has exact match
has close match
has related match
examples
references	Simpson 2019 Grootemaat 2017
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	leaf_curliness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011318

Leaf lamina posture (leaf 3-dimensional shape)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011319
alternative label	leaf_lamina_posture
description	A leaf shape trait [TO:0000492] which is the 3-D shape [PATO:0002266] of the surface [NCIT:C25245] of the leaf lamina [PO:0020039], including its degree of curvature [PATO:0001591] and whether it is bent [PATO:0000617]. The 3-dimension shape of the surface of the leaf lamina, including its degree of curvature and bending.
comments
value type	categorical variable
has narrower	leaf_lamina_posture_concave A longitudinal posture term indicating the leaf lamina is curved upward along the central axis, such that the upper leaf lamina surface is concave. leaf_lamina_posture_conduplicate A longitudinal posture term indicating the leaf lamina is folded upward along the central axis. (Synonym, folded) leaf_lamina_posture_convex A longitudinal posture term indicating the leaf lamina is curved downward along the central axis, such that the lower leaf lamina surface is concave. (Synonym, concavoconvex, planoconvex) leaf_lamina_posture_convolute Leaf lamina is contorted. (Synonym, twisted) leaf_lamina_posture_flat The leaf lamina is a 2-dimensional plane. leaf_lamina_posture_incurved Leaf tip is curved upward. leaf_lamina_posture_involute A longitudinal posture term indicating the outer portions of the leaf lamina are rolled inward or upward. leaf_lamina_posture_plicate Leaf margin is pleated, with a series of longitudinal folds. leaf_lamina_posture_recurved Leaf tip is curved outward or downward. [PATO:0002211] leaf_lamina_posture_revolute A longitudinal posture term indicating the outer portions of the leaf lamina are rolled outward or downward. leaf_lamina_posture_undulate The leaf lamina surface is rippled or wavy in its 2-dimensional plane. [PATO:0000967] (Synonyms, wavy, rippled, sinuate)
has broader	leaf shape trait
plant structure	leaf
measured characteristic	shape 3-D shape 3-D extent
keyword	leaf shape leaf lamina
scope note	Plants with leaves
has exact match
has close match
has related match
examples
references	no linked references
date created	07/06/2023
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011319

Leaf glaucousness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011360
alternative label	leaf_glaucousness
description	A leaf morphology trait [TO:0000748] which is the presence or absence of epicuticular wax [PO:0025388] on the leaf lamina [PO:0020039] of a fully expanded adult phase leaf [PO:0006340], giving the leaf lamina surface a white-coloured [PATO:0000014] or blue-coloured waxy [PATO:0002384] appearance, termed glaucous. Variable describing whether mature adult phase leaves have a glaucous appearance, the colour due to being covered with a smooth, usually whitish to bluish epicuticular wax that can easily be rubbed off.
comments	The accompanying methods should specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type	categorical variable
has narrower	leaf_glaucousness_glaucous Mature (fully expanded) leaves on adult plants have a waxy coating, giving them a glaucous appearance. (Synonyms, waxy bloom, pruinose) leaf_glaucousness_not_glaucous Mature (fully expanded) leaves on adult plants do not have a glaucous appearance.
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	colour / color texture
keyword	epicuticular wax leaf lamina waxy
scope note	Plants with leaves.
has exact match	leaf gloss
has close match
has related match
examples
references	Simpson 2019
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	glaucous
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011360

Mature leaf hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011361
alternative label	leaf_hairs_adult_leaves
description	A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of a fully expanded adult phase leaf [PO:0006340]. Variable describing whether mature (fully expanded) adult phase leaves on adult plants have hairs (trichomes).
comments	This trait captures the presence or absence of hairs on mature (fully expanded), adult phase leaves on adult plants. Best practice is to sample leaves that are recently fully expanded, since older leaves have often shed many of their hairs. The accompanying methods should specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type	categorical variable
has narrower	leaf_hairs_adult_leaves_glabrous Mature (fully expanded) leaves on adult plants do not have hairs (trichomes). leaf_hairs_adult_leaves_hairy Mature (fully expanded) leaves on adult plants have hairs (trichomes).
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	texture pilosity
keyword	leaf pubescence leaf trichome leaf lamina vestiture hairy adult vascular leaf pilosity trichome
scope note	Plants with leaves.
has exact match	leaf lamina pubescence
has close match
has related match	leaf trichome density
examples	close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
references	Hanley 2007 Simpson 2019
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	leaf_hairs_adult
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011361

Juvenile phase leaf hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011362
alternative label	leaf_hairs_juvenile_leaves
description	A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of a juvenile vascular leaf [PO:0006339] (on adult or juvenile plants). Variable describing whether fully expanded juvenile phase leaves (on adult or juvenile plants) have hairs (trichomes).
comments	This trait captures the presence or absence of hairs on fully-expanded juvenile phase leaves. It is best practice for the methods to specify whether these leaves were sampled from adult or juvenile plants. The accompanying methods should also specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type	categorical variable
has narrower	leaf_hairs_juvenile_leaves_glabrous Juvenile phase leaves do not have hairs (trichomes). leaf_hairs_juvenile_leaves_hairy Juvenile phase leaves have hairs (trichomes).
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	texture pilosity
keyword	leaf pubescence leaf trichome leaf lamina vestiture hairy juvenile vascular leaf pilosity trichome
scope note	Plants with leaves.
has exact match
has close match	leaf lamina pubescence
has related match	leaf trichome density
examples	close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
references	Hanley 2007 Simpson 2019
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	leaf_hairs_juvenile
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011362

Immature leaf hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011363
alternative label	leaf_hairs_immature_leaves
description	A leaf pubescence trait [TO:0002722] which is the presence or absence of trichomes [PO:0006504] (hairs) on the leaf lamina [PO:0020039] of an adult phase leaf [PO:0006340] still in the vascular leaf expansion stage [PO:0001052]. Variable describing whether immature (still expanding) adult phase leaves on adult plants have hairs (trichomes).
comments	This trait captures the presence or absence of hairs on immature (still expanding) leaves on adult plants. The accompanying methods should specify if the trait was scored on the upper (adaxial) or lower (abaxial) surface of the leaf; in the absence of any notes, it is assumed the trait was scored on the upper (adaxial) leaf surface.
value type	categorical variable
has narrower	leaf_hairs_immature_leaves_glabrous Immature (still expanding) leaves do not have hairs (trichomes). leaf_hairs_immature_leaves_hairy Immature (still expanding) leaves have hairs (trichomes).
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	texture pilosity
keyword	leaf pubescence leaf trichome leaf lamina vestiture hairy adult vascular leaf vascular leaf expansion stage trichome
scope note	Plants with leaves.
has exact match
has close match	leaf lamina pubescence
has related match	leaf trichome density
examples	close match: Leaf emergences (pubescence, pruinescence, hairs, spines, thorns) [TRY:677] (https://www.try-db.org/de/de.php)
references	Hanley 2007 Simpson 2019
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011363

Leaf surface colour

Property	Value
URI	https://w3id.org/APD/traits/trait_0011364
alternative label	leaf_surface_colour
description	A leaf morphology trait [TO:0000748] which is which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the leaf lamina [PO:0020039]. Human visible dominant colour of the surface of a leaf.
comments
value type	categorical variable
has narrower	leaf_surface_colour_green Leaf lamina surface is green. leaf_surface_colour_grey_green Leaf lamina surface is grey-green. leaf_surface_colour_dark_green Leaf lamina surface is dark green. leaf_surface_colour_pale_green Leaf lamina surface is pale green. leaf_surface_colour_pale_grey_green Leaf lamina surface is pale grey-green. leaf_surface_colour_dark_grey_green Leaf lamina surface is dark grey-green. leaf_surface_colour_red Leaf lamina surface is red. leaf_surface_colour_purple Leaf lamina surface is purple.
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	colour / color
keyword	leaf lamina adult vascular leaf
scope note
has exact match	leaf lamina color
has close match
has related match
examples	exact match: Leaf color [TRY:582] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011364

Leaf surface reflectivity

Property	Value
URI	https://w3id.org/APD/traits/trait_0011365
alternative label	leaf_surface_reflectivity
description	A leaf morphology trait [TO:0000748] which is the reflectivity [PATO:0001297] of the surface [NCIT:C25245] of a mature [PATO:0001701] leaf lamina [PO:0020039]. Surface reflectivity of a leaf, indicating whether the leaf lamina is shiny or dull.
comments	There will be overlap between this trait and the trait leaf_glaucousness. This trait can be used to document surface reflectivity more broadly, not only distinguishing between leaf lamina that are glaucous versus not glaucous.
value type	categorical variable
has narrower	leaf_surface_reflectivity_dull Leaf surface appears dull, indicating it has a low reflectivity. leaf_surface_reflectivity_shiny Leaf surface appears shiny or glossy, indicating it has a high reflectivity. (Synonyms, glossy, lustrous, polished)
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	texture morphology reflectivity
keyword	leaf lamina adult vascular leaf
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011365

Leaf phyllotaxis

Property	Value
URI	https://w3id.org/APD/traits/trait_0011410
alternative label	leaf_phyllotaxis
description	A leaf [PO:0025034] spatial pattern [PATO:0000060] trait which is the phyllotaxy [TO:0006014] of leaves, indicating a leaf’s position [TO:0000541] relative to other leaves at a stem node [PO:0020141]. Leaf arrangement at a stem node, indicating the relative number and position of leaves at a single stem note.
comments	This trait only includes true phyllotaxis, the arrangement of leaves at a single node, while the trait leaf arrangement indicates how the leaves are presented along the entire shoot system. See the trait stem_growth_habit for whole-plant leaf pattern terms, such as rosette.
value type	categorical variable
has narrower	leaf_phyllotaxis_alternate A single leaf is attached at each stem node. leaf_phyllotaxis_opposite A pair of leaves are attached opposite to one another at each stem node. leaf_phyllotaxis_whorled Three or more leaves are attached at each stem node, arranged in a symmetrical pattern around the stem. (Synonyms, verticillate)
has broader	leaf position trait
plant structure	leaf
measured characteristic	spatial pattern, distribution arrangement
keyword	phyllotaxy stem node
scope note	Plants with leaves.
has exact match	phyllotaxy
has close match
has related match
examples	close match: Leaf distribution along the shoot axis (arrangement type) [TRY:16] (https://www.try-db.org/de/de.php) close match: Leaf_arrangement [GIFT:4.15.1] (https://gift.uni-goettingen.de)
references	Simpson 2019 Ellis 2009 Wikipedia 2022 Leaf arrangement on the stem
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011410

Leaf arrangement

Property	Value
URI	https://w3id.org/APD/traits/trait_0011411
alternative label	leaf_arrangement
description	A leaf [PO:0025034] spatial pattern [PATO:0000060] trait which is the position [TO:0000541] and arrangement [NCIT:C61532] of leaves between stem nodes [PO:0020141], indicating how leaves align and cluster along the vegetative shoot system [PO:0025607]. Leaf arrangement across stem nodes, indicating the 3-dimensional arrangement of leaves along the length of the stem.
comments	Leaf arrangement, as defined for this trait, indicates patterns of leaf arrangement across nodes, including both formal terms (decussate, distichous, spiral) as well as indications of leaf clustering due to variation in internode length. See the trait leaf phyllotaxis for terms describing leaf arrangement at a single node. See the trait stem_growth_habit for whole-plant leaf pattern terms, such as rosette.
value type	categorical variable
has narrower	leaf_arrangement_decussate When sequential pairs of opposite leaves attach at right angles to the pair above or below. leaf_arrangement_distichous When alternate or opposite leaves are arranged in two opposite rows. leaf_arrangement_scattered When leaves are attached along the entire length of stem. leaf_arrangement_spiral When alternate leaves are attached in a progressive spiral pattern along the stem. leaf_arrangement_fasciculate A tight bundle or cluster of leaves attaches to the stem in a single location, generally due to the shortening of internodes. (Synonym, tufted) leaf_arrangement_clustered When leaves are in any type of cluster, where the type of cluster is not further specified. The leaves may be clustered toward the base of the plant (a rosette), toward the top of the plant, toward the end of branches, or in distinct tuft (fascicles), but more information is not provided. leaf_arrangement_clustered_basal When leaves are clustered or crowded to the base of the stem, arranged in a rosette or other basal clustering. (Synonym, rosette) leaf_arrangement_crowded When internode length varies considerably along a stem, such that leaves appear crowded or tufted. (Synonym, congested) leaf_arrangement_rosette When leaves radiate from a central point at or near ground level. leaf_arrangement_imbricate When leaves lay tightly against the stem and are overlapping.
has broader	leaf position trait
plant structure	leaf
measured characteristic	spatial pattern, distribution arrangement
keyword	stem node vegetative shoot system phyllotaxy
scope note	Plants with leaves.
has exact match
has close match
has related match
examples	close match: Leaf distribution along the shoot axis (arrangement type) [TRY:16] (https://www.try-db.org/de/de.php) close match: Leaf_arrangement [GIFT:4.15.1] (https://gift.uni-goettingen.de)
references	Simpson 2019 Ellis 2009 Wikipedia 2022 Leaf arrangement on the stem
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011411

Leaf axil angle

Property	Value
URI	https://w3id.org/APD/traits/trait_0011412
alternative label	leaf_axil_angle
description	A leaf position trait [TO:0000541] which is the angle [PATO:0002326] of a leaf axil [PO:0009023], the space between a leaf [PO:0025034] and the stem [PO:0009047] to which it is attached. The angle between a shoot and a leaf that connects to the shoot axis.
comments	This trait records the angle between the upper surface of a leaf and the stem to which it is attached. The slope of the leaf lamina relative to the solar zenith is a separate trait, leaf_inclination_angle.
value type	continuous variable
unit	deg plane angle degree
minAllowedValue	0
maxAllowedValue	180
has broader	leaf position trait
plant structure	leaf
measured characteristic	angle orientation spatial pattern, distribution
keyword	leaf axil leaf attitude leaf lamina leaf angle
scope note	Plants with leaves
has exact match	leaf angle
has close match
has related match
examples	related match: Leaf angle (inclination, orientation) [TRY:3] (https://www.try-db.org/de/de.php)
references	no linked references
date created	31/10/2022
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011412

Leaf inclination angle

Property	Value
URI	https://w3id.org/APD/traits/trait_0011413
alternative label	leaf_inclination_angle
description	A leaf position trait [TO:0000541] which is the inclination angle [PATO:0002326] of the adaxial [PATO:0002047] leaf lamina [PO:0020039] surface relative to a horizontal [PATO:0001855] plane, such that an adaxial leaf surface facing the solar zenith [ECSO:00002723] has an angle of 0. The inclination angle of the upper (adaxial) leaf lamina surface relative to a horizontal plane, such that a leaf surface facing the solar zenith has an angle of 0.
comments	This trait records the slope of the leaf lamina relative to the ground surface. The angle between a stem and the leaf lamina is a separate trait, leaf_axil_angle. Complanate leaves can be scored as having a leaf_inclination_angle of 0.
value type	continuous variable
unit	deg plane angle degree
minAllowedValue	-90
maxAllowedValue	90
has broader	leaf position trait
plant structure	leaf
measured characteristic	angle orientation spatial pattern, distribution
keyword	leaf lamina solar zenith angle leaf attitude
scope note	Plants with leaves
has exact match
has close match	leaf attitude
has related match
examples	exact match: Leaf angle (inclination, orientation) [TRY:3] (https://www.try-db.org/de/de.php)
references	Niinemets 2010 Falster 2003b
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	leaf_angle
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011413

Leaf pendulousness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011414
alternative label	leaf_pendulousness
description	A leaf position trait [TO:0000541] which indicates whether leaves [PO:0025034] are pendulous [APD:glossary/40066], hanging loosely downward and possessing the ability to flutter. Binary variable describing whether leaves are pendulous, hanging loosely downward and possessing the ability to flutter.
comments	This is a binary categorical trait, scoring whether leaves are pendulous and therefore hanging loosely downward and able to flutter. Two additional traits, leaf_inclination_angle and leaf_axil_angle document the actual angle of the leaf lamina relative to, respectively, the solar zenith and the stem.
value type	categorical variable
has narrower	leaf_pendulousness_pendulous Leaves are pendulous. leaf_pendulousness_not_pendulous Leaves are not pendulous.
has broader	leaf position trait
plant structure	leaf
measured characteristic	spatial pattern, distribution orientation angle
keyword	leaf angle pendulous
scope note	Plants with leaves
has exact match
has close match	leaf attitude
has related match
examples	close match: Leaf angle (inclination, orientation) [TRY:3] (https://www.try-db.org/de/de.php)
references	Curtis 2012
date created	14/07/2021
date modified	05/02/2024
date reviewed	31/10/2022
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Ian J. Wright Peter Vesk Gregory J. Jordan Andy Leigh Patricia Lu-Irving Lily Dun David Coleman
change note	pendulous_leaves
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011414

Cuticle thickness on the lower leaf surface

Property	Value
URI	https://w3id.org/APD/traits/trait_0011510
alternative label	leaf_cuticle_thickness_abaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of cuticle [PO:0000022] on the abaxial [PATO:0002046] (lower) leaf lamina [PO:0020039]. Thickness of the cuticle on the lower leaf surface.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	0.1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	plant cuticle abaxial leaf lamina
scope note
has exact match
has close match	plant cuticle thickness
has related match
examples	close match: Leaf cuticula thickness [TRY:716] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	lower_cuticle_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011510

Cuticle thickness on the upper leaf surface

Property	Value
URI	https://w3id.org/APD/traits/trait_0011511
alternative label	leaf_cuticle_thickness_adaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of cuticle [PO:0000022] on the adaxial [PATO:0002047] (upper) leaf lamina [PO:0020039]. Thickness of the cuticle on the upper leaf surface.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	0.1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	plant cuticle adaxial leaf lamina
scope note
has exact match
has close match	plant cuticle thickness
has related match
examples	close match: Leaf cuticula thickness [TRY:716] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	upper_cuticle_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011511

Leaf epidermis thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011512
alternative label	leaf_epidermis_thickness
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] , in samples where the leaf lamina [PO:0020039] side (abaxial [PATO:0002046] versus adaxial [PATO:0002047]) is not specified. Thickness of the epidermis, where the leaf lamina side is not specified.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue epidermis leaf lamina abaxial adaxial
scope note
has exact match
has close match	epidermis thickness
has related match
examples	exact match: Leaf epidermis thickness [TRY:101] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	epidermis_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011512

Lower leaf side epidermis thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011513
alternative label	leaf_epidermis_thickness_abaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039]. Thickness of the epidermis on the lower side of the leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue epidermis abaxial leaf lamina
scope note
has exact match
has close match	epidermis thickness
has related match
examples	close match: Leaf epidermis thickness [TRY:101] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	epidermis_thickness_lower_leaf_surface
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011513

Upper leaf side epidermis thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011514
alternative label	leaf_epidermis_thickness_adaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of epidermis [PO:0005679] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039]. Thickness of the epidermis on the upper side of the leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue epidermis adaxial leaf lamina
scope note
has exact match
has close match	epidermis thickness
has related match
examples	close match: Leaf epidermis thickness [TRY:101] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	epidermis_thickness_upper_leaf_surface
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011514

Average leaf epidermal cell density

Property	Value
URI	https://w3id.org/APD/traits/trait_0011515
alternative label	leaf_epidermal_cell_density_both_sides
description	A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] to leaf lamina [PO:0020039] area [PATO_0001323], averaged across the abaxial [PATO:0002046] (lower) and adaxial [PATO:0002047] (upper) sides of the leaf lamina. Epidermal cell density averaged across the upper and lower sides of the leaf.
comments
value type	continuous variable
unit	{count}/mm2 reciprocal square millimetre
minAllowedValue	100
maxAllowedValue	10000
has broader	plant cell morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	has number of count density ratio; proportion count per unit area
keyword	cell epidermal cell abaxial adaxial leaf lamina
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	epidermal_cell_density_both_sides
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011515

Lower leaf side epidermal cell density

Property	Value
URI	https://w3id.org/APD/traits/trait_0011516
alternative label	leaf_epidermal_cell_density_abaxial
description	A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039] to leaf lamina area [PATO_0001323]. Epidermal cell density on the lower side of the leaf.
comments
value type	continuous variable
unit	{count}/mm2 reciprocal square millimetre
minAllowedValue	100
maxAllowedValue	10000
has broader	plant cell morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	has number of count density ratio; proportion count per unit area
keyword	cell epidermal cell abaxial leaf lamina
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	epidermal_cell_density_abaxial
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011516

Upper leaf side epidermal cell density

Property	Value
URI	https://w3id.org/APD/traits/trait_0011517
alternative label	leaf_epidermal_cell_density_adaxial
description	A plant cell morphology trait [TO:0002683] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of epidermal cells [PO:0004013] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039] to leaf lamina area [PATO_0001323]. Epidermal cell density on the upper side of the leaf.
comments
value type	continuous variable
unit	{count}/mm2 reciprocal square millimetre
minAllowedValue	100
maxAllowedValue	10000
has broader	plant cell morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	has number of count density ratio; proportion count per unit area
keyword	cell epidermal cell adaxial leaf lamina
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	epidermal_cell_density_adaxial
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011517

Lower leaf side hypodermis thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011518
alternative label	leaf_hypodermis_thickness_abaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of hypodermis [PO:0005051] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039]. Thickness of the hypodermis on the lower side of the leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue hypodermis abaxial leaf lamina
scope note
has exact match
has close match
has related match
examples	close match: Leaf hypodermis thickness [TRY:806] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	lower_hypodermis_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011518

Upper leaf side hypodermis thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011519
alternative label	leaf_hypodermis_thickness_adaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of hypodermis [PO:0005051] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039]. Thickness of the hypodermis on the upper side of the leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	1
maxAllowedValue	100
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue hypodermis adaxial leaf lamina
scope note
has exact match
has close match
has related match
examples	close match: Leaf hypodermis thickness [TRY:806] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	upper_hypodermis_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011519

Lower palisade mesophyll thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011520
alternative label	leaf_palisade_tissue_thickness_abaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of palisade mesophyll [PO:0005648] tissue [CARO:0000043] on the abaxial [PATO:0002046] (lower) side of the leaf lamina [PO:0020039]. Thickness of palisade mesophyll tissue on the lower side of the leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	10
maxAllowedValue	1000
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue palisade mesophyll abaxial leaf lamina
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	lower_palisade_cell_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011520

Upper palisade mesophyll thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011521
alternative label	leaf_palisade_tissue_thickness_adaxial
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of palisade mesophyll [PO:0005648] tissue [CARO:0000043] on the adaxial [PATO:0002047] (upper) side of the leaf lamina [PO:0020039]. Thickness of palisade mesophyll tissue on the upper side of the leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	10
maxAllowedValue	1000
has broader	portion of plant tissue morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue palisade mesophyll adaxial leaf lamina
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	upper_palisade_cell_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011521

Palisade cell length

Property	Value
URI	https://w3id.org/APD/traits/trait_0011522
alternative label	leaf_palisade_cell_length
description	A plant cell morphology trait [TO:0002683] which is the length [PATO:0000122] of individual palisade mesophyll cells [PO:0006206]. Length of individual leaf palisade cells.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	10
maxAllowedValue	1000
has broader	plant cell morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	length
keyword	cell palisade mesophyll cell leaf lamina
scope note
has exact match
has close match
has related match	plant cell length
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	palisade_cell_length
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011522

Palisade cell width

Property	Value
URI	https://w3id.org/APD/traits/trait_0011523
alternative label	leaf_palisade_cell_width
description	A plant cell morphology trait [TO:0002683] which is the width [PATO:0000921] of individual palisade mesophyll cells [PO:0006206]. Width of individual leaf palisade cells.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	1
maxAllowedValue	50
has broader	plant cell morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	width
keyword	cell palisade mesophyll cell leaf lamina
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	palisade_cell_width
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011523

Number of layers of palisade cells

Property	Value
URI	https://w3id.org/APD/traits/trait_0011524
alternative label	leaf_palisade_layer_number
description	A portion of plant tissue morphology trait [TO:0000843] which is the count [NCIT:C25463] of cell layers [UBERON:0000119] of palisade mesophyll cells [PO:0006206]. The number of layers of palisade cells in a leaf.
comments
value type	continuous variable
unit	{count}
minAllowedValue	1
maxAllowedValue	10
has broader	portion of plant tissue morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	has number of count
keyword	cell palisade mesophyll cell palisade mesophyll leaf lamina tissue
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	palisade_layer_number
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011524

Spongy mesophyll cell thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011525
alternative label	leaf_spongy_mesophyll_thickness
description	A portion of plant tissue morphology trait [TO:0000843] which is the thickness [PATO:0000915] of spongy mesophyll [PO:0005647] tissue [CARO:0000043] within the leaf lamina [PO:0020039]. Thickness of spongy mesophyll tissue in a leaf.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	10
maxAllowedValue	1000
has broader	portion of plant tissue morphology trait leaf morphology trait
plant structure	leaf
measured characteristic	thickness
keyword	tissue spongy mesophyll leaf lamina
scope note
has exact match
has close match
has related match
examples	exact match: Leaf mesophyll spongiophyll thickness [TRY:805] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	spongy_mesophyll_thickness
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011525

Cell cross-sectional area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011526
alternative label	cell_cross-sectional_area
description	A plant cell size [TO:0002684] trait which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a cell [CL:0000000]. Cell cross-sectional area.
comments	This trait can be used to map the area of cell types that are not already explicitly listed as their own trait. The cell type and tissue/organ being measured needs to be specified under measurement_remarks or as a method_context.
value type	continuous variable
unit	um2 square micrometre
minAllowedValue	100
maxAllowedValue	10000
has broader	plant cell morphology trait
plant structure	whole plant
measured characteristic	area 2-D extent size
keyword	cell plant cell size cross section
scope note
has exact match
has close match
has related match
examples	close match: Leaf epidermis cell area [TRY:338] (https://www.try-db.org/de/de.php) close match: Leaf mesophyll cell area [TRY:573] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011526

Stomatal density on the lower leaf surface

Property	Value
URI	https://w3id.org/APD/traits/trait_0011610
alternative label	leaf_stomatal_density_abaxial
description	A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] on the abaxial [PATO:0002046] (lower) leaf lamina [PO:0020039] to the leaf lamina area [PATO:0001323]. Density of stomata on the lower leaf surface.
comments
value type	continuous variable
unit	{count}/mm2 reciprocal square millimetre
minAllowedValue	1
maxAllowedValue	1000
has broader	leaf stomatal complex morphology trait
plant structure	leaf
measured characteristic	has number of count density ratio; proportion count per unit area
keyword	leaf stomatal complex frequency stomata; stomatal pore abaxial leaf lamina
scope note
has exact match	abaxial stomatal frequency
has close match	Stomatal density
has related match
examples	exact match: Stomata density abaxial [TOP371] (https://top-thesaurus.org/index) close match: Stomata density [TRY:63] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stomatal_density_abaxial
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011610

Stomatal density on the upper leaf surface

Property	Value
URI	https://w3id.org/APD/traits/trait_0011611
alternative label	leaf_stomatal_density_adaxial
description	A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] on the adaxial [PATO:0002047] (upper) leaf lamina [PO:0020039] to the leaf lamina area [PATO:0001323]. Density of stomata on the upper leaf surface.
comments
value type	continuous variable
unit	{count}/mm2 reciprocal square millimetre
minAllowedValue	0
maxAllowedValue	1000
has broader	leaf stomatal complex morphology trait
plant structure	leaf
measured characteristic	has number of count density ratio; proportion count per unit area
keyword	leaf stomatal complex frequency stomata; stomatal pore adaxial leaf lamina
scope note
has exact match	adaxial stomatal frequency
has close match	Stomatal density
has related match
examples	exact match: Stomata density adaxial [TOP372] (https://top-thesaurus.org/index) close match: Stomata density [TRY:63] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stomatal_density_adaxial
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011611

Stomatal density averaged across both leaf surfaces

Property	Value
URI	https://w3id.org/APD/traits/trait_0011612
alternative label	leaf_stomatal_density_average
description	A leaf stomatal complex frequency trait [TO:0000566] which is the ratio [PATO:0001470] of the count [NCIT:C25463] of stomata [PO:0008032] to leaf lamina [PO:0020039] area [PATO:0001323], averaged the across the abaxial [PATO:0002046] and adaxial [PATO:0002047] sides of the leaf lamina. Density of stomata averaged across both leaf surfaces.
comments
value type	continuous variable
unit	{count}/mm2 reciprocal square millimetre
minAllowedValue	0
maxAllowedValue	1000
has broader	leaf stomatal complex morphology trait
plant structure	leaf
measured characteristic	has number of count density ratio; proportion count per unit area
keyword	leaf stomatal complex frequency stomata; stomatal pore abaxial adaxial leaf lamina
scope note
has exact match	Stomatal density
has close match
has related match
examples	exact match: Stomata density trait [TOP370] (https://top-thesaurus.org/index) exact match: Stomata density [TRY:63] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stomatal_density_average
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011612

Stomatal distribution

Property	Value
URI	https://w3id.org/APD/traits/trait_0011613
alternative label	leaf_stomatal_distribution
description	A leaf stomatal complex frequency trait [TO:0000566] which indicates whether stomata [PO:0008032] occur on the abaxial [PATO:0002046] and/or adaxial [PATO:0002047] leaf lamina [PO:0020039]. Distribution of stomatal across the two leaf surfaces.
comments
value type	categorical variable
has narrower	leaf_stomatal_distribution_amphistomatic Leaves with stomata on both leaf surfaces. leaf_stomatal_distribution_amphistomatic_uneven Leaves with stomata on both leaf surfaces but more stomata on the lower leaf surface. leaf_stomatal_distribution_epistomatic Leaves that only have stomata on the upper (adaxial) leaf surface. leaf_stomatal_distribution_hypostomatic Leaves that only have stomata on the lower (abaxial) leaf surface.
has broader	leaf stomatal complex morphology trait
plant structure	leaf
measured characteristic	spatial pattern, distribution
keyword	leaf stomatal complex frequency stomata; stomatal pore abaxial adaxial leaf lamina
scope note
has exact match
has close match
has related match
examples	exact match: Stomata distribution to ab- and adaxial leaf surfaces [TRY:781] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stomatal_distribution
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011613

Stomatal hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011614
alternative label	leaf_stomatal_hairs
description	A leaf stomatal complex frequency trait [TO:0000566] which is the pilosity [PATO:0000066] (hairiness) of the stomatal pores [PO:0008032]. Variable indicating if stomata are covered by hairs and, if hairy, the density of hairs present.
comments
value type	categorical variable
has narrower	leaf_stomatal_hairs_stomata_covered_with_dense_hairs Stomata densely covered by hairs. leaf_stomatal_hairs_stomata_not_covered_by_hairs Stomata not hair-covered.
has broader	leaf stomatal complex morphology trait
plant structure	leaf
measured characteristic	texture pilosity
keyword	leaf stomatal complex frequency stomata; stomatal pore hairy
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stomatal_hairs
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011614

Guard cell length

Property	Value
URI	https://w3id.org/APD/traits/trait_0011615
alternative label	leaf_guard_cell_length
description	A leaf stomatal complex frequency trait [TO:0000566] which indicates the length [PATO:0000122] of a guard cell [PO:0000293]. Length of guard cells.
comments
value type	continuous variable
unit	um micrometre
minAllowedValue	10
maxAllowedValue	200
has broader	leaf stomatal complex morphology trait
plant structure	leaf
measured characteristic	length
keyword	cell leaf stomatal complex frequency guard cell stomata; stomatal pore
scope note
has exact match
has close match
has related match	plant cell length
examples	exact match: Stomata guard cell size [TRY:733] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	guard_cell_length
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011615

Leaf visible light transmission

Property	Value
URI	https://w3id.org/APD/traits/trait_0011710
alternative label	leaf_transmission
description	A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is transmitted [SWEET:procWave/Transmission] through a leaf. Proportion of incoming visible light (between 400-700 nm) that is transmitted through a leaf.
comments
value type	continuous variable
unit	umol/umol micromole per micromole
minAllowedValue	0.001
maxAllowedValue	1
has broader	leaf optical properties trait
plant structure	leaf
measured characteristic	dimensionless ratio ratio; proportion electromagnetic property radiation
keyword	optics visible light transmission photosynthetically active radiation
scope note
has exact match
has close match
has related match
examples	exact match: Leaf light transmittance [TRY:535] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011710

Leaf visible light absorption

Property	Value
URI	https://w3id.org/APD/traits/trait_0011711
alternative label	leaf_absorption
description	A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is absorbed [SWEET:procWave/Absorption] by a leaf. Proportion of incoming visible light (between 400-700 nm) that is absorbed by a leaf.
comments
value type	continuous variable
unit	umol/umol micromole per micromole
minAllowedValue	0.01
maxAllowedValue	1
has broader	leaf optical properties trait
plant structure	leaf
measured characteristic	dimensionless ratio ratio; proportion electromagnetic property radiation
keyword	optics visible light absorption photosynthetically active radiation
scope note
has exact match
has close match
has related match
examples	exact match: Leaf light absorption (extinction) [TRY:10] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011711

Leaf visible light reflection

Property	Value
URI	https://w3id.org/APD/traits/trait_0011712
alternative label	leaf_reflectance
description	A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming visible light [NCIT:C17732] that is reflected [SWEET:procWave/Reflection] by a leaf. Proportion of incoming visible light (between 400-700 nm) that is reflected by a leaf.
comments
value type	continuous variable
unit	umol/umol micromole per micromole
minAllowedValue	0.01
maxAllowedValue	1
has broader	leaf optical properties trait
plant structure	leaf
measured characteristic	dimensionless ratio ratio; proportion electromagnetic property radiation
keyword	optics visible light reflection photosynthetically active radiation
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011712

Leaf infra-red light reflection

Property	Value
URI	https://w3id.org/APD/traits/trait_0011713
alternative label	leaf_reflectance_near_infrared
description	A leaf [PO:0025034] optical [NCIT:C16940] properties trait which is the proportion [PATO:0001470] of incoming infrared radiation [ENVO:21001214] that is reflected [SWEET:procWave/Reflection] by a leaf. Proportion of incoming near infra-red light (between 750-10500 nm) that is reflected by a leaf.
comments
value type	continuous variable
unit	umol/umol micromole per micromole
minAllowedValue	0.1
maxAllowedValue	1
has broader	leaf optical properties trait
plant structure	leaf
measured characteristic	dimensionless ratio ratio; proportion electromagnetic property radiation
keyword	infrared radiation optics reflection radiation
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011713

Stem cross-sectional area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011811
alternative label	stem_cross_sectional_area
description	A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a stem [PO:0009047] segment which includes the bark [PO:0004518]. Cross-sectional area of a stem, including both wood and bark.
comments	This is not a trait itself, since it is dependent on the location sampled. However, values might be reported that are effectively contexts for another trait, such as bark thickness or Huber value.
value type	continuous variable
unit	mm2 square millimetre
minAllowedValue	1
maxAllowedValue	1000
has broader	stem morphology trait
plant structure	stem
measured characteristic	area size
keyword	cross section
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stem_cross_section_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011811

Wood cross-sectional area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011812
alternative label	sapwood_cross_sectional_area
description	A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of a stem [PO:0009047] segment from which the bark [PO:0004518] has been removed; this includes both sapwood [PO:0004513] and, if present heartwood [PO:0004512]. Cross-sectional area of a stem without bark.
comments	This is not a trait itself, since it is dependent on the location sampled. However, values might be reported that are effectively contexts for another trait, such as bark thickness or Huber value. Although it is defined as the diameter of the entire stem without bark, it is generally measured on narrow branches that have not yet developed heartwood and therefore is the cross-section of sapwood.
value type	continuous variable
unit	mm2 square millimetre
minAllowedValue	1
maxAllowedValue	1000
has broader	stem morphology trait
plant structure	stem
measured characteristic	area size
keyword	sapwood cross section wood, secondary xylem
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	stem_cross_sectional_area_no_bark
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011812

Terminal twig cross-sectional area

Property	Value
URI	https://w3id.org/APD/traits/trait_0011813
alternative label	branch_terminal_twig_cross_sectional_area
description	A stem morphology trait [TO:0000361] which is the cross-sectional [NCIT:C63795] area [PATO:0001323] of the most distal [PATO:0001234] branch [PO:0025073]. Cross-sectional area of the terminal twig, measured just beyond the final branching node.
comments
value type	continuous variable
unit	mm2 square millimetre
minAllowedValue	0.05
maxAllowedValue	50
has broader	stem morphology trait
plant structure	stem
measured characteristic	area size
keyword	branch cross section
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	twig_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011813

Terminal twig length

Property	Value
URI	https://w3id.org/APD/traits/trait_0011814
alternative label	branch_terminal_twig_length
description	A stem morphology trait [TO:0000361] which is the length [PATO:0000122] of the most distal [PATO:0001234] branch [PO:0025073]. Length of the terminal twig, the length of stem beyond the final branching node.
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	1
maxAllowedValue	1000
has broader	stem morphology trait
plant structure	stem
measured characteristic	length
keyword	branch
scope note
has exact match
has close match
has related match
examples	exact match: Twig length [TRY:1202] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	twig_length
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011814

Wood density

Property	Value
URI	https://w3id.org/APD/traits/trait_0011815
alternative label	wood_density
description	A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of a unit of wood [NCIT:C93062] from a stem [PO:0009047] assessed after drying to its volume [PATO:0000918] assessed without drying. The ratio of wood dry mass to wood fresh volume, measured on a stem segment without bark.
comments
value type	continuous variable
unit	mg/mm3 milligram per cubic millimetre
minAllowedValue	0.05
maxAllowedValue	1.4
has broader	stem morphology trait
plant structure	stem
measured characteristic	mass volume density ratio; proportion mass density, density
keyword	wood wood, secondary xylem biomass allocation
scope note
has exact match	secondary xylem volumetric density, wood volumetric density Wood density
has close match
has related match
examples	exact match: Wood density [TOP284] (https://top-thesaurus.org/index) exact match: Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density [TRY:4] (https://www.try-db.org/de/de.php) exact match: Stem specific density (SSD) [GIFT:4.3] (https://gift.uni-goettingen.de) exact match: stem wood density [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Stem specific density [BROT:11] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011815

Herbaceous stem density

Property	Value
URI	https://w3id.org/APD/traits/trait_0011816
alternative label	stem_density
description	A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of a unit of herbaceous [PATO:0002352] stem [PO:0009047] assessed after drying to its volume [PATO:0000918] assessed without drying. The ratio of stem dry mass to stem fresh volume, measured on an herbaceous stem segment.
comments	This trait is specifically for non-woody or partially woody stems that otherwise are outliers for wood density
value type	continuous variable
unit	mg/mm3 milligram per cubic millimetre
minAllowedValue	0.1
maxAllowedValue	1.4
has broader	stem morphology trait
plant structure	stem
measured characteristic	mass volume density ratio; proportion mass density, density
keyword	herbaceous biomass allocation
scope note
has exact match
has close match
has related match
examples	exact match: Stem specific density (SSD, stem dry mass per stem fresh volume) or wood density [TRY:4] (https://www.try-db.org/de/de.php) exact match: Stem specific density (SSD) [GIFT:4.3] (https://gift.uni-goettingen.de) exact match: stem wood density [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Stem specific density [BROT:11] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011816

Stem hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0011817
alternative label	stem_hairs
description	A stem morphology trait [TO:0000361] which is the presence or absence of trichomes [PO:0006504] (hairs) on a stem [PO_0009047]. Variable describing whether plant stems have hairs (trichomes).
comments	As stems often lose hairs as they age, the context field should indicate the stem age.
value type	categorical variable
has narrower	stem_hairs_glabrous Adult plant stems do not have hairs (trichomes). stem_hairs_hairy Adult plant stems have hairs (trichomes).
has broader	stem morphology trait
plant structure	stem
measured characteristic	texture pilosity
keyword	trichome
scope note
has exact match	stem hairiness
has close match
has related match
examples	close match: Stem emergences (pubescence, hairs, spines, thorns) [TRY:1256] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011817

Xylem to leaf area ratio (XLA)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011910
alternative label	xylem_to_leaf_area_ratio
description	A leaf morphology trait [TO:0000748] which is the ratio [PATO:0001470] of xylem vessel [PO:0025417] area [PATO:0001323] in a petiole [PO:0020038] to the attached leaf lamina [PO:0020039] area. For a single leaf, the ratio of xyem cross-sectional area at the base of the petiole to the leaf lamina area.
comments	This trait is similar to Huber value, but at the level of a single leaf, not the entire branch.
value type	continuous variable
unit	cm2{xylem}/mm2{leaf}
minAllowedValue	0.01
maxAllowedValue	100
has broader	leaf morphology trait
plant structure	leaf
measured characteristic	area size dimensionless ratio ratio; proportion
keyword	biomass allocation xylem water transport leaf lamina
scope note
has exact match
has close match
has related match
examples
references	Blackman 2023
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011910

Huber value

Property	Value
URI	https://w3id.org/APD/traits/trait_0011911
alternative label	huber_value
description	A stem morphology trait [TO:0000361] which is the ratio [PATO:0001470] of sapwood [PO:0004513] area [PATO:0001323] to the total distal [PATO:0001234] leaf [PO:0025034] area. For a single branch, the ratio of sapwood cross-sectional area at the base of the branch to the branch’s total leaf area.
comments
value type	continuous variable
unit	mm2{sapwood}/mm2{leaf} square millimetre per square millimetre
minAllowedValue	1e-06
maxAllowedValue	0.2
has broader	stem morphology trait
plant structure plant structure	leaf stem
measured characteristic	area size dimensionless ratio ratio; proportion
keyword	sapwood biomass allocation
scope note
has exact match
has close match
has related match
examples	exact match: Stem, branch or twig sapwood cross-sectional area per supported leaf area (Huber value) [TRY:171] (https://www.try-db.org/de/de.php)
references	Tyree & Ewers 1991
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011911

Leaf dry mass to stem dry mass ratio

Property	Value
URI	https://w3id.org/APD/traits/trait_0011912
alternative label	leaf_mass_to_stem_mass_ratio
description	A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of a plant’s total leaf [PO:0025034] dry [PATO:0001824] mass [PATO:0000125] to its total stem [PO:0009047] and branch [PO:0025073] dry mass. The ratio of whole plant leaf dry mass to whole plant stem dry mass.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	10
has broader	leaf mass trait stem mass trait
plant structure plant structure plant structure	leaf stem whole plant
measured characteristic	mass mass fraction ratio; proportion
keyword	biomass allocation whole plant stem dry mass whole plant leaf dry mass
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	leaf_mass_to_stem_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011912

Stem dry mass to vegetative shoot dry mass ratio (support fraction)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011913
alternative label	stem_mass_to_shoot_mass_ratio
description	A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of stem [PO:0009047] and branch [PO:0025073] dry [PATO:0001824] mass [PATO:0000125] to vegetative shoot system [PO:0025607] dry mass. The ratio of whole plant stem dry mass to whole plant vegetative shoot dry mass. That is, the proportion of aboveground vegetative biomass that is comprised of stems and branches, rather than leaves; support fraction.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0
maxAllowedValue	1
has broader	leaf mass trait stem mass trait
plant structure plant structure plant structure	leaf stem whole plant
measured characteristic	mass mass fraction ratio; proportion
keyword	branch vegetative shoot system biomass allocation whole plant stem dry mass whole plant leaf dry mass
scope note
has exact match
has close match
has related match
examples	close match: Plant biomass and allometry: Supportive structure (stem, twig, leaf petiole, leaf sheeth) dry mass p [TRY:3059] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	support_fraction
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011913

Side branch dry mass to whole plant dry mass ratio

Property	Value
URI	https://w3id.org/APD/traits/trait_0011914
alternative label	branch_mass_fraction
description	A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of side branch [PO:0025073] dry [PATO:0001824] mass [PATO:0000125] to whole plant [PO:0000003] dry mass. The ratio of side branch dry mass to whole plant dry mass. That is, the proportion of total plant biomass that is comprised of all branches off the main shoot axis.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0
maxAllowedValue	1
has broader	stem mass trait
plant structure plant structure	stem whole plant
measured characteristic	mass mass fraction ratio; proportion
keyword	branch biomass allocation whole plant dry mass
scope note
has exact match
has close match
has related match
examples	exact match: Plant biomass and allometry: Branch dry mass per plant dry mass [TRY:137] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011914

Stem dry matter content (SDMC)

Property	Value
URI	https://w3id.org/APD/traits/trait_0011915
alternative label	stem_dry_matter_content
description	A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of stem [PO:0009047] (or branch [PO:0025073]) dry [PATO:0001824] mass [PATO:0000125] to stem (or branch) fresh mass [APD:glossary/40044]. The ratio of stem (or branch) dry mass to stem (or branch) fresh mass; SDMC.
comments	A related trait is stem_water_content_per_saturated_mass.
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	100
maxAllowedValue	1000
has broader	stem mass trait
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	branch fresh mass water
scope note
has exact match
has close match
has related match
examples	exact match: Stem dry mass per stem fresh mass (stem dry matter content, StDMC) [TRY:1181] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011915

Stem water content per unit saturated stem mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011916
alternative label	stem_water_content_per_saturated_mass
description	A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a stem [PO:0009047] to stem fresh mass [APD:glossary/40044] in a stem that is at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620]. The ratio of the mass of water in a stem (stem water content) to total stem mass, measured in a saturated stem. A saturated stem is one that has been hydrated to its maximal water holding capacity and is at full turgor.
comments	A related trait is stem_dry_matter_content.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.1
maxAllowedValue	10
has broader	stem mass trait
plant structure	stem
measured characteristic	mass mass fraction ratio; proportion
keyword	water water holding capacity turgor fresh mass
scope note
has exact match
has close match
has related match	stem moisture content
examples	close match: Wood (sapwood) water content [TRY:1097] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011916

Stem mass fraction

Property	Value
URI	https://w3id.org/APD/traits/trait_0011917
alternative label	stem_mass_fraction
description	A plant dry mass trait [TO:0000352] which is the ratio [PATO:0001470] of whole plant [PO:0000003] stem [PO:0009047] dry [PATO:0001824] mass [PATO:0000125] to whole plant dry mass [APD:glossary/40080]. The ratio of stem dry mass to whole plant dry mass. That is, the proportion of total plant biomass that is comprised of stems and branches.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	1
has broader	stem mass trait
plant structure plant structure	stem whole plant
measured characteristic	mass mass fraction ratio; proportion
keyword	biomass allocation whole plant stem dry mass whole plant dry mass
scope note
has exact match
has close match
has related match
examples	exact match: Plant biomass and allometry: Stem dry mass per plant dry mass [TRY:136] (https://www.try-db.org/de/de.php) related match: stem dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011917

Stem saturated water content per unit stem dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0011918
alternative label	stem_saturated_water_content_per_dry_mass
description	A stem mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in a saturated stem [PO:0009047] to stem dry [PATO:0001824] mass [PATO:0000125], where a saturated stem is one at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091]. The ratio of the mass of water in a saturated stem to stem dry mass.
comments	Similar traits include stem_dry_matter_content, the ratio of a stem dry mass to stem fresh mass and stem_water_content_per_saturated_mass the ratio of the mass of water in a stem to saturated stem mass.
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	20
has broader	stem mass trait
plant structure plant structure	stem whole plant
measured characteristic	mass mass fraction ratio; proportion
keyword	biomass allocation whole plant stem dry mass whole plant dry mass
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0011918

Bark morphology, Eucalyptus

Property	Value
URI	https://w3id.org/APD/traits/trait_0012010
alternative label	bark_morphology_eucalyptus
description	A bark morphology trait which is the texture [PATO:0000150] of the bark [PO:0004518] of Eucalyptus taxa. Broad categories that describe the texture of Eucalyptus bark.
comments	This trait just records the morphology of taxa in the genera Eucalyptus, Corymbia, and Angophora.
value type	categorical variable
has narrower	bark_morphology_eucalyptus_eucalypt_box Eucalyptus, Corymbia or Angophora taxa with ‘Box’ type bark. bark_morphology_eucalyptus_eucalypt_gum Eucalyptus, Corymbia or Angophora taxa with ‘Gum’ type bark. bark_morphology_eucalyptus_eucalypt_stocking Eucalyptus, Corymbia or Angophora taxa with ‘Gum’ type bark above a rough barked stocking. bark_morphology_eucalyptus_eucalypt_ironbark Eucalyptus, Corymbia or Angophora taxa with ‘Ironbark’ type bark. bark_morphology_eucalyptus_eucalypt_peppermint Eucalyptus, Corymbia or Angophora taxa with ‘Peppermint’ type bark. bark_morphology_eucalyptus_eucalypt_stringybark Eucalyptus, Corymbia or Angophora taxa with ‘Stringy’ type bark. bark_morphology_eucalyptus_eucalypt_ribbonbark Eucalyptus, Corymbia or Angophora taxa with ‘Gum’ type bark that sheds in ribbons.
has broader	bark morphology trait
plant structure	bark
measured characteristic	structure texture
scope note	taxa in the genera Eucalyptus, Corymbia, and Angophora
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012010

Bark thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0012011
alternative label	bark_thickness
description	A bark morphology trait which is the thickness [PATO:0000915] of the bark [PO:0004518]. The corresponding stem [PO:0009047] diameter [PATO:0001334] should also be recorded so that a scaled measure, the bark thickness to stem diameter ratio, can also be calculated. Thickness of the bark on the stem.
comments	The corresponding stem diameter should also be recorded so that a scaled measure, the bark thickness to stem diameter ratio, can also be calculated.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	200
has broader	bark morphology trait
plant structure	bark
measured characteristic	thickness
scope note
has exact match	Bark thickness
has close match
has related match
examples	exact match: Bark thickness [TOP283] (https://top-thesaurus.org/index) exact match: Bark thickness [TRY:24] (https://www.try-db.org/de/de.php) exact match: Bark thickness (inner bark) [TRY:3355] (https://www.try-db.org/de/de.php) exact match: Bark thickness (outer bark) [TRY:3356] (https://www.try-db.org/de/de.php) exact match: Average bark thickness [BROT:9] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012011

Scaled bark thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0012012
alternative label	bark_thickness_index
description	A bark morphology trait which is the ratio [PATO:0001470] of two times bark [PO:0004518] thickness [PATO:0000915] to the stem [PO:0009047] diameter [PATO:0001334]. The ratio of two times a single bark thickness measure to stem diameter, to adjust for increasing bark thickness with increasing stem diameter.
comments
value type	continuous variable
unit	mm/mm millimetre per millimetre
minAllowedValue	0.001
maxAllowedValue	1
has broader	bark morphology trait
plant structure plant structure	bark stem
measured characteristic	thickness index
keyword	biomass allocation
scope note
has exact match
has close match
has related match
examples	close match: Bark thickness [TOP283] (https://top-thesaurus.org/index) close match: Bark thickness [TRY:24] (https://www.try-db.org/de/de.php) close match: Bark thickness (inner bark) [TRY:3355] (https://www.try-db.org/de/de.php) close match: Bark thickness (outer bark) [TRY:3356] (https://www.try-db.org/de/de.php)
references	Lawes 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012012

Bark density

Property	Value
URI	https://w3id.org/APD/traits/trait_0012013
alternative label	bark_density
description	The ratio [PATO:0001470] of bark [PO:0004518] dry [PATO:0001824] mass [PATO:0000125] to the corresponding fresh [EnvThes:21976] bark volume [PATO:0000918]. The ratio of bark dry mass to bark fresh volume.
comments
value type	continuous variable
unit	mg/mm3 milligram per cubic millimetre
minAllowedValue	0.1
maxAllowedValue	1.1
has broader	bark morphology trait
plant structure	bark
measured characteristic	mass volume density ratio; proportion mass density, density
keyword	biomass allocation
scope note
has exact match
has close match
has related match
examples	exact match: Bark density [TOP282] (https://top-thesaurus.org/index) exact match: Bark density (bark dry mass per bark volume) [TRY:248] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012013

Bark dry mass per unit bark surface area

Property	Value
URI	https://w3id.org/APD/traits/trait_0012014
alternative label	bark_dry_mass_per_surface_area
description	The ratio [PATO:0001470] of bark [PO:0004518] dry [PATO:0001824] mass [PATO:0000125] to the corresponding bark surface area [EnvThes:21564]. The ratio of bark dry mass to bark surface area.
comments
value type	continuous variable
unit	g/cm2 gram per square centimetre
minAllowedValue	0.01
maxAllowedValue	1
has broader	bark morphology trait
plant structure	bark
measured characteristic	mass area density per unit area ratio; proportion mass density, density
keyword	biomass allocation
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	bark_mass_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012014

Bark water content per unit bark dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012015
alternative label	bark_water_content_per_dry_mass
description	The ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in bark [PO:0004518] (bark water content [APD:glossary/40014]) to bark dry [PATO:0001824] mass. The ratio of the mass of water in bark (bark water content) to bark dry mass.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.01
maxAllowedValue	1
has broader	bark morphology trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	water bark water content
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012015

Bark water content per unit saturated bark mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012016
alternative label	bark_water_content_per_saturated_mass
description	A bark mass trait [TO:0000952] which is the ratio [PATO:0001470] of the mass [PATO:0000125] of water [CHEBI:15377] in bark [PO:0004518] to the fresh mass [APD:glossary/40044] of bark that is at maximum [NCIT:C25564] water holding capacity [IOBC:200906064703668091], known as full turgor [PATO:0001620]. The ratio of the mass of water in bark (bark water content) to total bark mass, measured in saturated bark. Saturated bark is bark that has been hydrated to its maximal water holding capacity and is at full turgor.
comments
value type	continuous variable
unit	g/g gram per gram
minAllowedValue	0.1
maxAllowedValue	10
has broader	bark morphology trait
plant structure	bark
measured characteristic	mass mass fraction ratio; proportion
keyword	water water holding capacity turgor fresh mass
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012016

Root diameter

Property	Value
URI	https://w3id.org/APD/traits/trait_0012111
alternative label	root_diameter
description	A root morphology trait [TO:0000043] which is the maximum [NCIT:C25564] root [PO:0009005] diameter [PATO:0001334] of a plant. Maximum root diameter.
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	1000
has broader	root morphology trait
plant structure	root
measured characteristic	diameter size
keyword	root morphology trait
scope note
has exact match	root width
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012111

Root system morphology

Property	Value
URI	https://w3id.org/APD/traits/trait_0012112
alternative label	root_system_classification
description	A root system morphology trait [TO:0000985] which are categorical [STATO:0000252] trait values that capture the branchiness [PATO:0002009] and texture [PATO:0000150] of the root system [PO:0025025] and the root system depth [TO:0001058]. Categorical root system descriptions that indicate the branchiness and texture of the root system and the depth of the root system.
comments	The trait values listed are those specified in the root system classification scheme proposed by Cannon in 1949.
value type	categorical variable
has narrower	root_system_classification_type_i The primary root is fibrous or fleshy and is relatively long; mesophytic. root_system_classification_type_ii The primary root is relatively short and is fibrous. Roots are well branched in a dry habitat but branch little in one that is wet. root_system_classification_type_iii The primary root is usually long and fibrous. The first order lateral roots are well branched and are usually short; they are rather evenly distributed along the primary root; xerophytic. root_system_classification_type_iv The primary root is usually long and slender like that of Type II. The first order lateral roots are well branched. Those closest to the surface of the ground, or not far below, are relatively long; xerophytic and mesophytic. root_system_classification_type_v The primary root is divided-forked or branched-and fibrous, and with few first order lateral roots, or, in age, apparently with none; xerophytic. root_system_classification_type_vi The primary root is usually as in Type V. The first order lateral roots are mostly on the upper portion of the primary root and are thus relatively superficial; xerophytic or in well aerated sandy soil where the rainfall is not light. root_system_classification_type_vii Adventitious roots that are in one group and arise on a short horizontal, or vertical, axis of the shoot. They are fibrous or fleshly; mesophytic and hydrophytic. root_system_classification_type_viii Adventitious roots that are in one group and are fibrous and fleshy, or fibrous and thick. The fibrous adventitious roots are absorbing-anchoring roots and are usually with laterals to the second order; meophytic. root_system_classification_type_ix Adventitious roots that arise on the aerial stem and branches of some herbaceous taxa, shrubs, and trees; on creepers, vines, and runners; and on subterranean rhizomes and stolons.
has broader	root morphology trait
plant structure	root
measured characteristic	volume 3-D extent size texture spatial pattern, distribution
keyword	root system root system depth root branchiness root texture
scope note
has exact match
has close match
has related match
examples
references	Cannon 1949
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	root_morphology
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012112

Fine root volume to coarse root volume ratio

Property	Value
URI	https://w3id.org/APD/traits/trait_0012113
alternative label	root_fine_root_coarse_root_ratio
description	A root system morphology trait [TO:0000985] which is the ratio [PATO:0001470] of the volume [PATO:0000918] of fine roots [EnvThes:20581] (those <0.5mm in diameter) to the volume of coarse roots [APD:glossary/40026] (those >0.5 mm in diameter). The ratio of the volume of fine roots (<0.5mm diameter) to the volume of coarse roots (>0.5mm diameter).
comments
value type	continuous variable
unit	mm3/mm3 cubic millimetre per cubic millimetre
minAllowedValue	0.05
maxAllowedValue	50
has broader	root morphology trait
plant structure	root
measured characteristic	volume ratio; proportion
keyword	fine root root branchiness coarse root
scope note
has exact match
has close match
has related match
examples	close match: Fine root dry mass fraction that is taproot (taproot dry mass per fine root dry mass) [TRY:2011] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012113

Root biomass depth distribution coefficient

Property	Value
URI	https://w3id.org/APD/traits/trait_0012114
alternative label	root_distribution_coefficient
description	A root system morphology trait [TO:0000985] which is a model parameter [STATO:0000034] indicating the distribution [PATO:0000060] of a plant’s roots [PO:0009005] with depth [PATO:0001595]. To calculate the model parameter, the required input is a plot of the cumulative root distribution from the surface, where [Y] is the cumulative fraction of roots from the surface to a depth [d]. An asymptotic nonlinear model is used to describe the vertical root distribution: Y = 1 - b^d where Y is the cumulative root fraction from the surface to soil depth d in centimeters and b the estimated parameter. Root biomass depth distribution coefficient (‘B’ from Gale & Grigal 1987), where high values indicate that root biomass is allocated deeper in the soil. To calculate the model parameter, the required input is a plot of the cumulative root distribution from the surface, where [Y] is the cumulative fraction of roots from the surface to a depth [d]. An asymptotic nonlinear model is used to describe the vertical root distribution: Y = 1 - b^d where Y is the cumulative root fraction from the surface to soil depth d in centimeters and b the estimated parameter.
comments
value type	continuous variable
unit	{dimensionless}
minAllowedValue	0.5
maxAllowedValue	1
has broader	root morphology trait
plant structure	root
measured characteristic	mass model parameter spatial pattern, distribution
keyword	root branchiness root system depth root system
scope note
has exact match
has close match
has related match
examples	close match: Root evenness of dry mass vertical distribution [TRY:3033] (https://www.try-db.org/de/de.php)
references	Gale and Grigal 1987
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012114

Root system type (presence of taproot)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012115
alternative label	root_system_type
description	A root system morphology trait [TO:0000985] which indicates if the root system [PO:0025025] is one where the primary root [PO:0020127] remains the dominant root, developing into a taproot, a long [PATO:0000573], thick [PATO:0000591] root with few branches, or whether the lateral roots [PO:0020121] are of similar length to the primary root. Variable describing whether a plant has a tap root.
comments
value type	categorical variable
has narrower	root_system_type_tap_root_absent Tap root not present. root_system_type_tap_root_present Tap root present. root_system_type_tap_root_sometimes_present Tap root sometimes present.
has broader	root morphology trait
plant structure	root
measured characteristic	structure spatial pattern, distribution
keyword	root system primary root lateral root root branchiness
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	tap_root; root_tap_root_status
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012115

Specific root length (SRL)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012116
alternative label	root_specific_root_length
description	A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of root [PO:0009005] length [PATO:0000122] to root dry mass [TO:0000078]. The ratio of root length to root dry mass; SRL.
comments
value type	continuous variable
unit	m/g metre per gram
minAllowedValue	0.5
maxAllowedValue	5000
has broader	root morphology trait
plant structure	root
measured characteristic	length ratio; proportion mass
keyword	biomass allocation root system primary root lateral root
scope note
has exact match
has close match
has related match
examples	exact match: Specific root length trait [TOP935] (https://top-thesaurus.org/index) exact match: Root length per root dry mass (specific root length, SRL) [TRY:1080] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	specific_root_length
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012116

Specific tap root length (STRL)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012117
alternative label	root_specific_taproot_length
description	A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of primary root [PO:0020127] length [PATO:0000122] to primary root dry mass [TO:0000078] The ratio of taproot length (primary root length) to taproot dry mass; STRL.
comments	This trait is usually measured during the very early stage of growth, capturing how efficiently seedlings reach reliable water.
value type	continuous variable
unit	m/g metre per gram
minAllowedValue	0.5
maxAllowedValue	5000
has broader	root morphology trait
plant structure	root
measured characteristic	length ratio; proportion mass
keyword	biomass allocation root system primary root
scope note
has exact match
has close match
has related match
examples	close match: Coarse root length per coarse root dry mass (specific coarse root length, SRL) [TRY:1550] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	specific_taproot_length
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012117

Root surface area per unit root dry mass (specific root area)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012118
alternative label	root_specific_root_area
description	A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of root [PO:0009005] surface area [EnvThes:21564] to root dry mass [TO:0000078]. The ratio of root surface area to root dry mass; specific root area.
comments
value type	continuous variable
unit	mm2/mg square millimetre per milligram
minAllowedValue	0.001
maxAllowedValue	1
has broader	root morphology trait
plant structure	root
measured characteristic	area size ratio; proportion area density mass density, density
keyword	biomass allocation root system
scope note
has exact match
has close match
has related match
examples	exact match: Root surface area per root dry mass (specific root area) [TRY:1089] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note	specific_root_area
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012118

Root wood density

Property	Value
URI	https://w3id.org/APD/traits/trait_0012119
alternative label	root_wood_density
description	A root morphology trait [TO:0000043] which is the ratio [PATO:0001470] of the dry mass [PATO:0000125] of a unit of wood [PO_0005848] from a root [PO:0009005]to its fresh fresh [EnvThes:21976] volume [PATO:0000918]. The ratio of woody root dry mass to woody root fresh volume.
comments
value type	continuous variable
unit	mg/mm3 milligram per cubic millimetre
minAllowedValue	0.05
maxAllowedValue	10
has broader	root morphology trait
plant structure	root
measured characteristic	mass volume density ratio; proportion mass density, density
keyword	biomass allocation root system wood, secondary xylem
scope note
has exact match
has close match
has related match
examples	exact match: Root tissue density (root dry mass per root volume) [TRY:82] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012119

Root to shoot ratio

Property	Value
URI	https://w3id.org/APD/traits/trait_0012120
alternative label	root_shoot_ratio
description	A whole plant morphology trait [TO:0000398] which is the ratio [PATO:0001470] of a plant’s root dry mass [TO:0000078] to its shoot dry mass [TO:0000552]. The ratio of total root system dry mass to total shoot system dry mass.
comments	This trait is usually measured on experimentally grown seedlings where it is possible to collect the entire root system.
value type	continuous variable
unit	mg{root}/mg{shoot} milligram per milligram
minAllowedValue	0.01
maxAllowedValue	15
has broader	root morphology trait stem morphology trait
plant structure plant structure plant structure	root whole plant stem
measured characteristic	mass ratio; proportion ratio; proportion mass fraction mass density, density
keyword	biomass allocation root system
scope note
has exact match	root to shoot ratio
has close match
has related match
examples	exact match: Root/shoot ratio [TRY:9] (https://www.try-db.org/de/de.php) exact match: Shoot:root ratio [BROT:22] (http://doi.org/10.1038/sdata.2018.135)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012120

Root dry matter content (RDMC)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012121
alternative label	root_dry_matter_content
description	A root mass trait [TO:0000279] which is the ratio [PATO:0001470] of root dry mass [TO:0000078] to root [PO:0009005] fresh mass [APD:glossary/40044]. The ratio of root dry mass to root fresh mass; root dry matter content, RDMC.
comments
value type	continuous variable
unit	mg/g milligram per gram
minAllowedValue	10
maxAllowedValue	1000
has broader	root morphology trait
plant structure	root
measured characteristic	mass mass fraction ratio; proportion
keyword	biomass allocation root system fresh mass
scope note
has exact match
has close match
has related match
examples	exact match: Root dry mass per root fresh mass (root dry matter content, RDMC) [TRY:785] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012121

Root mass fraction

Property	Value
URI	https://w3id.org/APD/traits/trait_0012122
alternative label	root_mass_fraction
description	A root mass trait [TO:0000279] which is the ratio [PATO:0001470] of the total root dry mass [TO:0000078] to whole plant [PO:0000003] dry mass [APD:glossary/40080]. The ratio of total root system dry mass to whole plant dry mass.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.01
maxAllowedValue	1
has broader	root morphology trait
plant structure plant structure	root whole plant
measured characteristic	mass mass fraction ratio; proportion
keyword	biomass allocation whole plant dry mass root system
scope note
has exact match
has close match
has related match
examples	exact match: Plant biomass and allometry: Root mass fraction (fraction of root dry mass per whole plant dry mass) [TRY:470] (https://www.try-db.org/de/de.php) related match: root dry mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012122

Root vascular anatomy

Property	Value
URI	https://w3id.org/APD/traits/trait_0012123
alternative label	root_vascular_anatomy
description	A root system morphology trait [TO:0000985] which is the arrangement [NCIT:C61532] or type [NCIT:C25284] of the root [PO:0020127] vascular system [PO:0003011]. The arrangement or type of the root vascular system.
comments
value type	categorical variable
has narrower	root_vascular_anatomy_cord_roots A root anatomy found within three genera of Mirbelieae (Daviesia, Euchilopsis, Phyllota) wherein networks of interlinking vascular strands exist (Crisp 2003; Pate and Dixon 1996). root_vascular_anatomy_unistelar_roots The commonly observed root anatomy, wherein there is a single vascular strand which displays secondary thickening.
has broader	root morphology trait
plant structure	root
measured characteristic	volume 3-D extent size texture spatial pattern, distribution
keyword	root system root vascular cambium
scope note
has exact match
has close match
has related match
examples
references	Cannon 1949
date created	03/06/2025
date modified	03/06/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012123

Storage organ length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012151
alternative label	storage_organ_length
description	A plant structure morphology trait [TO:0000839] which is the length [PATO:0000122] of a plant’s storage organ [EFO:0000997], where a storage organ is a plant part (organ [PO:0009008], tissue [PO:0009007], or other plant structure [PO:0009011]) whose function [NCIT:C25518] is carbohydrate [CHEBI:16646] (energy [PATO:0001021]) storage. The length of an underground plant structure (tissues, organs, or other structures) where a plant stores excess carbohydrates (energy).
comments	This can be the length of any of the structures documented in the trait storage_organ and should be documented as an entity context.
value type	continuous variable
unit	mm millimetre
minAllowedValue	1
maxAllowedValue	1000
has broader	stem morphology trait root morphology trait
plant structure plant structure	root stem
measured characteristic	length size
keyword	storage organ energy
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012151

Storage organ diameter

Property	Value
URI	https://w3id.org/APD/traits/trait_0012152
alternative label	storage_organ_diameter
description	A plant structure morphology trait [TO:0000839] which is the diameter [PATO:0001334] of a plant’s storage organ [EFO:0000997], where a storage organ is a plant part (organ [PO:0009008], tissue [PO:0009007], or other plant structure [PO:0009011]) whose function [NCIT:C25518] is carbohydrate [CHEBI:16646] (energy [PATO:0001021]) storage. The width of an underground plant structure (tissues, organs, or other structures) where a plant stores excess carbohydrates (energy).
comments	This can be the length of any of the structures documented in the trait storage_organ and should be documented as an entity context.
value type	continuous variable
unit	mm millimetre
minAllowedValue	1
maxAllowedValue	1000
has broader	stem morphology trait root morphology trait
plant structure plant structure	root stem
measured characteristic	diameter size
keyword	storage organ energy
scope note
has exact match
has close match	tuberous root tuber diameter
has related match
examples	exact match: Belowground plant organ diameter [TRY:2550] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012152

Seed accessory cost fraction

Property	Value
URI	https://w3id.org/APD/traits/trait_0012221
alternative label	accessory_cost_fraction
description	A reproductive shoot system [PO:0025082] biomass allocation [EnvThes:21360] trait which is one minus the ratio [PATO:0001470] of total plant seed [PO:0009010] dry [PATO:0001824] mass [PATO:0000125] to total plant reproductive shoot system tissue dry mass where reproductive tissues include all flower buds [PO:0000056], flowers [PO:0009046], fruits [PO:0009001], dispersal [EnvThes:21038] tissues, and seeds produced during the developmental process involved in reproduction [GO:0003006]. The fraction of total reproductive investment required to mature a seed that is invested in non-seed tissues. It is calculated as one minus, the ratio of total biomass investment in seeds to total biomass investment in all reproductive tissues, including flower buds, flowers, fruits, dispersal tissues, aborted seeds, and successfully matured seeds.
comments
value type	continuous variable
unit	mg/mg milligram per milligram
minAllowedValue	0.01
maxAllowedValue	1
has broader	reproductive shoot system morphology trait
plant structure plant structure plant structure plant structure	reproductive shoot system flower fruit seed
measured characteristic	mass
keyword	biomass allocation reproduction accessory cost
scope note
has exact match
has close match
has related match
examples	close match: Plant biomass and allometry: Reproductive dry mass per plant dry mass (reproductive weight fraction) [TRY:1046] (https://www.try-db.org/de/de.php)
references	Lord & Westoby 2006
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012221

Seed accessory cost mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012222
alternative label	accessory_cost_mass
description	A reproductive shoot system [PO:0025082] biomass allocation [EnvThes:21360] trait which is the sum of the dry [PATO:0001824] mass [PATO:0000125] of all flower buds [PO:0000055], flowers [PO:0009046], fruits [PO:0009001], aborted seeds, and dispersal [EnvThes:21038] tissues produced during the developmental process involved in reproduction [GO:0003006], but excludes mature [PATO:0001701] seed [PO:0009010] dry mass. The mass of seed accessory costs, which is the total biomass investment in all reproductive tissues, excluding biomass invested in successfully matured seeds; it includes all biomass invested in flower buds, flowers, fruits, dispersal tissues and aborted seeds.
comments
value type	continuous variable
unit	mg milligram
minAllowedValue	0.01
maxAllowedValue	10000
has broader	reproductive shoot system morphology trait
plant structure plant structure plant structure plant structure	reproductive shoot system flower fruit seed
measured characteristic	mass
keyword	biomass allocation reproduction accessory cost
scope note
has exact match
has close match
has related match
examples
references	Lord & Westoby 2006
date created	14/07/2021
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012222

Bud length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012301
alternative label	bud_length
description	A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of the bud [PO:0000055] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the bud. Linear dimension from the base to the apex of a bud, even if this is not the longest dimension.
comments
value type	continuous variable
unit	cm
minAllowedValue	0.01
maxAllowedValue	200
has broader	floral organ morphology trait
plant structure plant structure	reproductive shoot system bud
measured characteristic	length size
scope note
has exact match	reproduction corolla perianth
has close match
has related match
examples
references	no linked references
date created	05/02/2024
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012301

Bud width

Property	Value
URI	https://w3id.org/APD/traits/trait_0012302
alternative label	bud_width
description	A floral organ morphology trait [TO:0000862] which is the width [PATO:0000921] of the bud [PO:0000055] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the bud. Width is the greatest linear dimension [PATO:0001708] perpendicular to [PATO:0002434] to the bud length [PATO:0000122] Greatest linear width dimension of a bud, sometimes referred to as ‘diameter’ in buds where the width and height are identical; orthogonal to the bud length, the base-to-apex dimension.
comments
value type	continuous variable
unit	cm
minAllowedValue	0.01
maxAllowedValue	200
has broader	floral organ morphology trait
plant structure plant structure	reproductive shoot system bud
measured characteristic	width size
scope note
has exact match	reproduction corolla perianth
has close match
has related match
examples
references	no linked references
date created	05/02/2024
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012302

Buds per inflorescence

Property	Value
URI	https://w3id.org/APD/traits/trait_0012351
alternative label	buds_per_inflorescence
description	A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of buds [PO:0000055] in an inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers or the entire reproductive shoot system [PO:0025082]. The count of buds in an inflorescence, where an inflorescence can be either a single cluster of flowers or the entire reproductive shoot system.
comments
value type	continuous variable
unit	{count}/{count}
minAllowedValue	1
maxAllowedValue	1000
has broader	floral organ morphology trait
plant structure plant structure plant structure	reproductive shoot system bud inflorescence
measured characteristic	has number of dimensionless ratio ratio; proportion count tolerance sensitivity
scope note
has exact match	reproduction reproductive shoot system inflorescence
has close match
has related match
examples
references	no linked references
date created	05/02/2024
date modified	05/02/2024
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012351

Inflorescence length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012352
alternative label	inflorescence_length
description	A floral organ morphology trait [TO:0000862] which is the vertical [PATO:0001854] length [PATO:0000122] (height [PATO:0000119]) of the inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers [PO:0009046] or the entire reproductive shoot system [PO:0025082]. The vertical length (height) of an inflorescence.
comments
value type	continuous variable
unit	cm
minAllowedValue	0.01
maxAllowedValue	1500
has broader	perianth morphology trait
plant structure plant structure plant structure	reproductive shoot system bud inflorescence
measured characteristic	length size
keyword	floral organ morphology trait flower reproductive shoot system inflorescence morphology trait
scope note
has exact match	inflorescence length
has close match
has related match
examples	exact match: Inflorescence length (height) [TRY:2817] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012352

Inflorescence diameter

Property	Value
URI	https://w3id.org/APD/traits/trait_0012353
alternative label	inflorescence_diameter
description	A floral organ morphology trait [TO:0000862] which is the diameter [PATO:0001334] of the inflorescence [PO:0009049], where an inflorescence can be either a single cluster of flowers [PO:0009046] or the entire reproductive shoot system [PO:0025082]. The diameter (of greatest width dimension) of an inflorescence.
comments
value type	continuous variable
unit	cm
minAllowedValue	0.01
maxAllowedValue	1500
has broader	perianth morphology trait
plant structure plant structure plant structure	reproductive shoot system bud inflorescence
measured characteristic	diameter size
keyword	floral organ morphology trait flower reproductive shoot system inflorescence morphology trait
scope note
has exact match	inflorescence diameter
has close match
has related match
examples	exact match: Inflorescence diameter [TRY:3587] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012353

Number of androecium parts in each whorl (Androecium structural merism)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012410
alternative label	flower_androecium_structural_merism
description	An androecium morphology trait [TO:0000994] which is the total count [NCIT:C25463] of stamens [PO:0009029], staminodes [PO:0009077] or stamen bundles (fascicles [NCIT:C79692]) in each floral whorl [PO:0025023]. The number of stamens or stamen bundles (fascicles) in one whorl, considering both fertile stamens and staminodes.
comments	This character is not applicable for spiral or irregular stamen arrangements, nor when there is a single structural stamen (i.e., one stamen, no staminodes; e.g., Chloranthus, Chloranthaceae). Definition derived from Schoenenberger et al. (2020) for their character ‘332. Androecium structural merism (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	100
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction androecium stamen staminode fascicle merism whorl, floral whorl
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012410

Androecium structural phyllotaxis

Property	Value
URI	https://w3id.org/APD/traits/trait_0012411
alternative label	flower_androecium_structural_phyllotaxis
description	An androecium morphology trait [TO:0000994] which is the spatial pattern or distribution [PATO:0000060] (phyllotaxy [TO:0006014]) of stamen [PO:0009029], staminodes [PO:0009077] or stamen bundles (fascicles [NCIT:C79692]) in each floral whorl [PO:0025023]. Structural phyllotaxy of the androecium, considering both fertile stamens and staminodes. In cases of stamen fascicles, it is the phyllotaxis of fascicles, not individual stamens that are recorded.
comments	This character is not applicable when there is a single structural stamen (i.e., one stamen, no staminodes; e.g., Chloranthus, Chloranthaceae). Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘330. Androecium structural phyllotaxy (D1)’
value type	categorical variable
has narrower	flower_androecium_structural_phyllotaxis_whorled Stamens and staminodes arranged in whorls. flower_androecium_structural_phyllotaxis_spiral Stamens and staminodes have a spiral arrangement. flower_androecium_structural_phyllotaxis_irregular Stamens and staminodes have an irregular arrangement.
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution
keyword	reproduction androecium phyllotaxy stamen staminode fascicle whorl, floral whorl
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012411

Number of androecium structural whorls

Property	Value
URI	https://w3id.org/APD/traits/trait_0012412
alternative label	flower_androecium_structural_whorls_count
description	An androecium morphology trait [TO:0000994] which is the total count [NCIT:C25463] of whorls [PO:0025023] comprising the androecium [PO:0009061], considering both stamen [PO:0009029] and staminodes [PO:0009077]. If stamens are grouped into fascicles [NCIT:C79692], the trait is the count of whorls of fascicles. The number of whorls, considering both fertile stamens and staminodes. In cases of stamen fascicles, the count of whorls of fascicles is recorded.
comments	This character is not applicable for spiral or irregular stamen arrangements, or when there is a single structural stamen (i.e., one stamen, no staminodes; e.g., Chloranthus, Chloranthaceae). Definition derived from Schoenenberger et al. (2020) for their character ‘331. Number of androecium structural whorls (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	100
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction androecium stamen staminode fascicle merism whorl, floral whorl
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012412

Anther attachment

Property	Value
URI	https://w3id.org/APD/traits/trait_0012413
alternative label	flower_anther_attachment
description	An androecium morphology trait [TO:0000994] which indicates how the filament [PO:0009067] is attached to [PATO:0001667] the tissue [CARO:0000043] connecting the two thecae [PO:0009069] (pollen [PO:0025281] sacs) of the anthers [PO:0009066], affecting the spatial pattern [PATO:0000060] of the stamens [PO:0009029]. Anther attachment refers to the area of insertion of the filament on the anther connective (i.e. the tissue connecting the two thecae of an anther).
comments	Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘312. Anther attachment (D1)’
value type	categorical variable
has narrower	flower_anther_attachment_basifixed Filament attached to the base of the connective. Laminar stamens are classified as basifixed. flower_anther_attachment_dorsifixed Filament attached to the dorsal side of the anther. flower_anther_attachment_dorsifixed_at_base Filament attached to the dorsal side of the anther, at the base. flower_anther_attachment_ventrifixed Filament attached to the ventral side of the anther.
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution
keyword	reproduction androecium stamen anther pollen connective anther theca filament
scope note
has exact match
has close match
has related match	anther attachment quality
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012413

Connective extension (apical)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012414
alternative label	flower_anther_connective_extension
description	An androecium morphology trait [TO:0000994] which is whether anthers [PO:0009066] have protrusions [NCIT:C50710] that extend above the two theca [PO:0009069] (pollen [PO:0025281] sacs), and, if present, the shape [PATO:0000052] and the relative length [PATO:0000122] of the protrusion relative to the length of the entire anther. Apical (distal) connective extensions (also called ‘distal connective protrusions’) are sterile anther structures that distally extend beyond the level of the thecae (i.e., the two lateral pairs of pollen sacs of a tetrasporangiate anther).
comments	This trait records both absence or presence of anther connective extensions and also the shape of these extensions in terms of length in relation to the length of the thecae. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘314. Connective extension (apical) (D1)’
value type	categorical variable
has narrower	flower_anther_connective_extension_absent Apical connective extensions are lacking. flower_anther_connective_extension_short_extension Connective extension less than a third of the length of anther. flower_anther_connective_extension_long_extension Connective extension more than a third but less than length of anther; e.g. Idiospermum australiense. flower_anther_connective_extension_very_long_extension Connective extension more than length of anther; e.g. Galbulimima belgraveana. flower_anther_connective_extension_present_general Apical connective extensions are present but no information on their length relative to the anthers.
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	structure shape length
keyword	reproduction androecium stamen anther pollen connective anther theca
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012414

Anther dehiscence

Property	Value
URI	https://w3id.org/APD/traits/trait_0012415
alternative label	flower_anther_dehiscence
description	An androecium morphology trait [TO:0000994] which is the length [PATO:0000122] and shape [PATO:0000052] of the stomia [PO:0020101] that form the anther dehiscence zone [PO:0005011]; stomia are the usually slit-like [PATO:0002482] openings in the anther [PO:0009066] through which pollen [PO:0025281] is released. Anther dehiscence refers to the type of opening of the anther when releasing its pollen through the stomia.
comments	Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘313. Anther dehiscence (D1)’
value type	categorical variable
has narrower	flower_anther_dehiscence_longitudinal_slit Anther with longitudinal slits that extend along the entire length of each theca. flower_anther_dehiscence_poricidal Anther where dehiscence of longitudinal slits is incomplete, with pollen exiting via a small pore at the tip. flower_anther_dehiscence_pores_many Anther with pollen exiting from many pores along its length. flower_anther_dehiscence_h_valvate Anther with two valves on longitudinal hinges, opening horizontally as salon doors; when the stomium bifurcates at its distal and/or proximal end and thus a valve is formed. flower_anther_dehiscence_flap_valvate Anther with one or more flap-like valves on horizontal hinges, opening vertically. flower_anther_dehiscence_transverse_slit Anther with horizontal slit. flower_anther_dehiscence_short_basal_slits Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the base of the theca. flower_anther_dehiscence_short_central_slits Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the center of the theca. flower_anther_dehiscence_short_apical_slits Anther where dehiscence of longitudinal slits is incomplete and only occurs over a short extent at the apical end of the theca. flower_anther_dehiscence_t_valvate Anther with T-shaped stomium; reduced transverse slit at base. flower_anther_dehiscence_common_stomium_confluent_thecae The two thecae of a single anther are fused at one end with a single pore through which pollen exits. flower_anther_dehiscence_circular Anther with a transverse slit that goes entirely around the anther.
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	structure shape length
keyword	reproduction androecium stamen anther pollen stomium anther dehiscence zone
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012415

Anther orientation

Property	Value
URI	https://w3id.org/APD/traits/trait_0012416
alternative label	flower_anther_orientation
description	An androecium morphology trait [TO:0000994] which is the position [PATO:0000140] and orientation [PATO:0000133] of the stomia [PO:0020101] that form the anther dehiscence zone [PO:0005011]; stomia are the slit-like [PATO:0002482] openings in the anther [PO:0009066] through which pollen [PO:0025281] is released. Orientation of anther at anthesis.
comments	Anthers of angiosperms are rather uniform in their basic structure. They normally have four microsporangia (pollen sacs) that are arranged pair-wise in two thecae. The two microsporangia of a theca usually release their pollen grains through a common opening (stomium). Often, it is difficult to establish anther orientation clearly in a flower as this is a gradual feature with many intermediate stages. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘311. Anther orientation (D1)’
value type	categorical variable
has narrower	flower_anther_orientation_apical When thecae are positioned in a transverse position at the tip of the connective and thus dehisce upward in the flower (e.g., Sinofranchetia, Endress and Hufford, 1989). flower_anther_orientation_extrorse When the stomia of the thecae face the floral periphery. flower_anther_orientation_introrse When the stomia of the thecae face the floral centre. flower_anther_orientation_latrorse When pollen is released toward the side (i.e., toward neighbouring anthers).
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution orientation position
keyword	reproduction androecium stamen anther pollen stomium anther dehiscence zone
scope note
has exact match
has close match	anther orientation
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012416

Flower colour

Property	Value
URI	https://w3id.org/APD/traits/trait_0012417
alternative label	flower_colour
description	A floral organ morphology trait [TO:0000862] which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the flower [PO:0009046. The petals [PO:0009032] (corolla [PO:0009059]), sepals [PO:0009031] (calyx [PO:0009060]), or the entire perianth [PO:0009058] may be responsible for the main colour observed. The dominant flower colour, where the entire flower, the petals, or the sepals may be responsible for the observed colour.
comments	Most data mapped to flower_colour has probably been scored based on perianth colour, but if a dataset does not differentiate between the two traits it is mapped to flower_colour.
value type	categorical variable
has narrower	flower_colour_blue_purple Flower colour is blue or purple. flower_colour_green Flower colour is green. flower_colour_pink Flower colour is pink. flower_colour_red_brown Flower colour is red or brown. flower_colour_white_cream Flower colour is white or cream. flower_colour_yellow_orange Flower colour is yellow or orange. flower_colour_grey Flower colour is grey. flower_colour_black Flower colour is black.
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	colour / color
keyword	reproduction petal corolla sepal calyx perianth
scope note
has exact match	flower color
has close match	corolla color
has related match
examples	exact match: Flower color [TOP242] (https://top-thesaurus.org/index) exact match: Flower color [TRY:207] (https://www.try-db.org/de/de.php) exact match: Flower_colour [GIFT:3.21.1] (https://gift.uni-goettingen.de) exact match: flower color [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012417

Perianth colour

Property	Value
URI	https://w3id.org/APD/traits/trait_0012418
alternative label	perianth_colour
description	A floral organ morphology trait [TO:0000862] which is the dominant (primary [NCIT:C25251]) colour [PATO:0000014] of the perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]). The dominant colour of the perianth parts (sepals, petals and tepals).
comments	This trait refers explicitly to the colour of the petals, sepals and tepals, while the trait ‘flower colour’ refers to the colour of the entire flower. See also data under flower_colour. Most data mapped to flower_colour has probably been scored based on perianth colour, but if a dataset does not differentiate between the two traits it is mapped to flower_colour.
value type	categorical variable
has narrower	perianth_colour_white Perianth colour is white. perianth_colour_yellow Perianth colour is yellow. perianth_colour_orange Perianth colour is orange. perianth_colour_red Perianth colour is red. perianth_colour_pink Perianth colour is pink. perianth_colour_purple Perianth colour is purple. perianth_colour_blue Perianth colour is blue. perianth_colour_green Perianth colour is green. perianth_colour_grey Perianth colour is grey. perianth_colour_brown Perianth colour is brown. perianth_colour_black Perianth colour is black.
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	colour / color
keyword	reproduction petal corolla sepal calyx perianth
scope note
has exact match
has close match	corolla color
has related match
examples	exact match: Flower color [TOP242] (https://top-thesaurus.org/index) exact match: Flower color [TRY:207] (https://www.try-db.org/de/de.php) exact match: Flower_colour [GIFT:3.21.1] (https://gift.uni-goettingen.de) exact match: flower color [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	no linked references
date created	19/05/2023
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012418

Flower length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012419
alternative label	flower_length
description	A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of the flower [PO:0009046] from the base of the receptable [PO:0009064] or the base of an inferior ovary [PO:0009072] to the tip of the flower, measured at anthesis [PO:0009068]. It excludes the length of the pedicel [PO:0030112]. The length of the flower.
comments
value type	continuous variable
unit	cm
minAllowedValue	0.01
maxAllowedValue	200
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	length
keyword	reproduction petal corolla sepal calyx perianth
scope note
has exact match	flower length
has close match
has related match
examples
references	no linked references
date created	19/05/2023
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012419

Flower diameter

Property	Value
URI	https://w3id.org/APD/traits/trait_0012420
alternative label	flower_diameter
description	A floral organ morphology trait [TO:0000862] which is the maximum diameter [PATO:0001334] of the flower [PO:0009046] at anthesis [PO:0009068], considering all floral organs, but most commonly the diameter from petal [PO:0009032] tip to petal tip. The maximum diameter of the flower at anthesis, considering all floral organs, but most commonly the diameter from petal tip to petal tip.
comments
value type	continuous variable
unit	cm
minAllowedValue	0.01
maxAllowedValue	200
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	diameter
keyword	reproduction petal corolla sepal calyx perianth
scope note
has exact match	flower diameter
has close match
has related match
examples
references	no linked references
date created	19/05/2023
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012420

Floral orientation

Property	Value
URI	https://w3id.org/APD/traits/trait_0012421
alternative label	flower_orientation
description	A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] and orientation [PATO:0000133] of the flower [PO:0009046]. The position and orientation of the flower.
comments
value type	categorical variable
has narrower	flower_orientation_up Flower orientation is vertical, with the floral entrance pointing up. flower_orientation_lateral Flower orientation is horizontal, with the floral entrance perpendicular to main vertical axis. flower_orientation_down Flower orientation is pendant, with the floral entrance pointing down. flower_orientation_mixed Flower orientation is mixed, with different flowers exhibiting different orientations.
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution orientation position
keyword	reproduction petal corolla sepal calyx perianth
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	19/05/2023
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012421

Maximum flower number

Property	Value
URI	https://w3id.org/APD/traits/trait_0012422
alternative label	flower_count_maximum
description	A floral organ morphology trait [TO:0000862] which is the maximum [NCIT:C25564] count [NCIT:C25463] of flowers [PO:0009046] on an individual [EFO:0000542]. Maximum flower number produced.
comments
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	1e+06
has broader	floral organ morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count maximum
keyword	reproduction perianth corolla
scope note
has exact match
has close match
has related match
examples	exact match: Flower number per plant [TRY:2999] (https://www.try-db.org/de/de.php)
references	no linked references
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012422

Number of fertile stamens

Property	Value
URI	https://w3id.org/APD/traits/trait_0012431
alternative label	flower_fertile_stamens_count
description	A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of fertile [PATO:0000955] stamens [PO:0009029] on a hermaphrodite [PATO:0001340] flower [PO:0009046] or male [PATO:0000384] flower. Count of fertile stamens in bisexual or male flowers; does not score female flowers.
comments	Definition derived from Schoenenberger et al. (2020) for their character ‘301. Number of fertile stamens (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	10000
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction androecium stamen
scope note	male or hermaphroditic flowers
has exact match	stamen number
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012431

Filament presence and shape

Property	Value
URI	https://w3id.org/APD/traits/trait_0012432
alternative label	flower_filament
description	A floral organ morphology trait [TO:0000862] which is whether a filament [PO:0009067] is present [PATO:0000467] or absent [PATO:0000462], and if present, the shape [PATO:0000052] and the relative length [PATO:0000122], width [PATO:0000921], and thickness [PATO:0000915] of the filament. Trait that considers the absence or presence of the filament, and in the latter case, the shape of the filament. Shape is considered in terms of length and width and is defined in relation to anther length/width.
comments	The width of the filament may thus either be broad as in laminar (e.g., Eupomatiaceae) or bulky stamens (e.g., Chloranthaceae), or narrow (filamentous) as found in many core eudicots groups (e.g., Rosaceae). This trait is considered inapplicable when filaments are entirely fused with each other or to the perianth. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘305. Filament (D1)’
value type	categorical variable
has narrower	flower_filament_absent No filaments present flower_filament_present_general Filaments present, with no specific defining characteristics. flower_filament_long_and_narrow Filaments have a long and narrow shape. (Synonyms, filiform, slender) flower_filament_short_and_narrow Filaments have a short and narrow shape; subsessile anthers. flower_filament_long_and_wide Filaments have a long and wide laminar shape. flower_filament_short_and_wide Filaments have a short and wide laminar shape. flower_filament_long_and_very_wide Filaments have a long and very wide shape; for instance, the petal-like (outer) stamens of Nymphaea. flower_filament_petaloid Filaments are long and wide and colorful, e.g. Canna. flower_filament_fused_into_synandrium Filament structure formed when stamens fused together. flower_filament_fused_into_fascicle_stalks Special case where the stamens are fused into the stalk of a fascicle of flowers. flower_filament_long_and_narrow_divided_in_two Filaments have a long and narrow shape that is divided in two; e.g. Adoxa. flower_filament_short_general Filament is short relative to the anther.
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	shape structure
keyword	reproduction androecium stamen filament anther
scope note	inapplicable when filaments are entirely fused with each other or to the perianth
has exact match
has close match	filament 2-D shape
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012432

Fusion of filaments

Property	Value
URI	https://w3id.org/APD/traits/trait_0012433
alternative label	flower_filament_fusion
description	A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of the stamen [PO:0009029] (or staminode [PO:0009077]) filaments [PO:0009067] that is fused [NCIT:C124537] together at anthesis [PO:0009068]. Fusion of stamen (and staminode) filaments among each other at anthesis (congenitally or postgenitally) is recorded on a continuous scale, from 0 (free filaments) to 1 (filaments fused along their entire length).
comments	Partial fusion is recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the length of filaments). Definition derived from Schoenenberger et al. (2020) for their character ‘306. Fusion of filaments (C1)’
value type	continuous variable
unit	{dimensionless}
minAllowedValue	0
maxAllowedValue	1
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	dimensionless ratio ratio; proportion
keyword	reproduction androecium stamen filament connective staminode
scope note
has exact match
has close match
has related match	filament attached to
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012433

Fusion of filaments to inner perianth series

Property	Value
URI	https://w3id.org/APD/traits/trait_0012434
alternative label	flower_filament_fusion_to_inner_perianth
description	A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of the stamen [PO:0009029] (or staminode [PO:0009077]) filaments [PO:0009067] that are fused [NCIT:C124537] to the innermost perianth [PO:0009058] organ [PO:0009008] at anthesis [PO:0009068]. The fusion of filaments with the innermost perianth organs at anthesis on a continuous scale, from 0 (filaments completely free from perianth) to 1 (filaments fused along their entire length with the perianth; the ‘entire length’ of the filament is defined as the distance between the floral base and the joint between filament and anther).
comments	Partial fusion is recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the length of perianth parts). If two (or more) stamen whorls (including staminodial whorls) differ in their extent of filament fusion with the perianth, this trait is recorded as a range of values. For example, if the filaments of an outermost stamen whorl are fused, up to 90% of their length, but an inner whorl only up to 50%, the trait is assigned a range of 0.5 to 0.9. The rationale for this is to provide a general trait that allows comparison of fusion among all angiosperms. Definition derived from Schoenenberger et al. (2020) for their character ‘308. Fusion of filaments to inner perianth series (C1)’
value type	continuous variable
unit	{dimensionless}
minAllowedValue	0
maxAllowedValue	1
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	dimensionless ratio ratio; proportion
keyword	reproduction androecium stamen filament connective staminode perianth
scope note
has exact match
has close match
has related match	filament attached to
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012434

Gynoecium phyllotaxis

Property	Value
URI	https://w3id.org/APD/traits/trait_0012441
alternative label	flower_gynoecium_phyllotaxis
description	A gynoecium [PO:0009062] morphology trait which is either the count [NCIT:C25463] of whorls [PO:0025023] of carpels [PO:0009030] comprising the gynoecium or other descriptors of carpel phyllotaxy [TO:0006014]. Structural phyllotaxy of the gynoecium.
comments	This trait is not applicable to unicarpellate flowers. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘400. Gynoecium phyllotaxy (D1)’
value type	categorical variable
has narrower	flower_gynoecium_phyllotaxis_one_whorl Flowers with a single whorl of carpels. flower_gynoecium_phyllotaxis_two_or_more_whorls Flowers with two or more whorls of carpels. flower_gynoecium_phyllotaxis_spiral Flowers with carpels in a spiral arrangement.
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution orientation structure has number of count
keyword	reproduction gynoecium carpel whorl, floral whorl phyllotaxy
scope note	Not applicable to unicarpellate flowers
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012441

Placentation

Property	Value
URI	https://w3id.org/APD/traits/trait_0012442
alternative label	flower_gynoecium_placentation
description	A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] of the ovules [PO:0020003] within the ovary [PO:0009072], known as placentation [APD:glossary/40074]. The different types of placentation in apocarpous/unicarpellate and in syncarpous gynoecia.
comments	In apocarpous and unicarpellate gynoecia, placentation is often described as marginal as the ovules are usually attached along the ventral slit (i.e., the zone where the carpel margins become postgenitally closed during carpel development). In syncarpous gynoecia there are three main types of placentation, 1) axile placentation refers to ovaries where the ovules are placed in the angle between carpel flanks in the center of the ovary; 2) in ovaries with parietal placentation the ovules attach to the ovary wall where two carpels meet; 3) in ovaries with free-central placentation the ovules are attached to a central column that emerges from the base of the ovary and protrudes into the non-septate ovary. It is not unusual that syncarpous gynoecia show a transition from proximally axile to a distally parietal placentation (e.g., Polemoniaceae). Basal and apical placentation may occur both in apocarpous/unicarpellate and in syncarpous gynoecia. Usually there is one longitudinal series of ovules attached to each carpel margin. However, both in apocarpous/unicarpellate and in syncarpous gynoecia there may be more than one series of ovules at the flanks of a carpel. These latter cases are referred to as laminar (or laminar-diffuse) placentation (e.g., in some Nymphaeaceae). A placenta that is protruding from its surroundings and has more than two series of ovules (either axile or parietal) is called protruding-diffuse. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘412. Placentation (D1)’
value type	categorical variable
has narrower	flower_gynoecium_placentation_axile Ovules are placed in the angle between carpel flanks in the center of the ovary; in syncarpous gynoecia. flower_gynoecium_placentation_apical Ovules attach at the top of the ovary; pendulous; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia. flower_gynoecium_placentation_basal Ovules attach at the base of the ovary; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia. flower_gynoecium_placentation_parietal Ovules attach to the ovary wall where two carpels meet; in syncarpous gynoecia. flower_gynoecium_placentation_marginal Ovules attach along the ventral slit; in apocarpous and unicarpellate gynoecia. flower_gynoecium_placentation_free_central Ovules are attached to a central column that emerges from the base of the ovary and protrudes into the non-septate ovary; in syncarpous gynoecia. flower_gynoecium_placentation_laminar When there is more than one series of ovules at the flanks of a carpel; in syncarpous gynoecia and apocarpous and unicarpellate gynoecia.
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	structure spatial pattern, distribution position
keyword	reproduction gynoecium carpel plant ovary placentation plant ovule
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012442

Fusion of ovaries

Property	Value
URI	https://w3id.org/APD/traits/trait_0012443
alternative label	flower_ovary_fusion
description	A floral organ morphology trait [TO:0000862] which, for flowers with multiple carpels, is the proportion [PATO:0001470] of the length [PATO:0000122] of the ovaries [PO:0009072] that are fused [NCIT:C124537] together. Degree of ovary fusion expressed as a fraction of the total length of the ovary (from the floral base to the apex of the ovary).
comments	Fusion of styles and stigmas is not taken into account for this trait. Not applicable when there is a single carpel. Definition derived from Schoenenberger et al. (2020) for their character ‘403. Fusion of ovaries (C1)’
value type	continuous variable
unit	{dimensionless}
minAllowedValue	0
maxAllowedValue	1
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	dimensionless ratio ratio; proportion structure
keyword	reproduction gynoecium carpel plant ovary
scope note	Not applicable when there is a single carpel.
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012443

Ovary position

Property	Value
URI	https://w3id.org/APD/traits/trait_0012444
alternative label	flower_ovary_position
description	A floral organ morphology trait [TO:0000862] which is the position [PATO:0000140] of the plant ovary [PO:0009072] relative to the other flower [PO:0009046] organs [PO:0009008], in particular whether the ovary is attached above or below the perianth [PO:0009058] organs (the petals [PO:0009032] and sepals [PO:0009031]). The position of the flower ovary relative to the insertion level of the remaining floral organs.
comments	The ovary is the part of the gynoecium where the ovules are produced. The ovary may be located on the receptacle and thus be positioned above the insertion level of the remaining floral organs (i.e., the ovary is superior and the flower is hypogynous). Alternatively, the ovary may be embedded in the receptacle and therefore be located below the insertion level of the remaining floral organs (i.e., the ovary is inferior and the flower is epigynous). Flowers with a hypanthium may either have a superior ovary (perigyny; e.g., many Rosaceae) or an inferior ovary (epiperigyny). It is also possible that the ovary is inferior to a certain degree only, such as half-inferior, if the receptacle is surrounding the ovary to its mid-level. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘102. Ovary position (D1)’
value type	categorical variable
has narrower	flower_ovary_position_inferior Flower with inferior ovary, where the ovary is embedded in the receptacle and therefore located below the insertion level of the remaining floral organs. flower_ovary_position_superior Flower with superior ovary, where the ovary is positioned above the insertion level of the remaining floral organs. flower_ovary_position_half_inferior Flower where the receptacle is surrounded by the ovary to its mid-level. flower_ovary_position_one_quarter_inferior Flower where the receptacle surrounds up to one quarter of the ovary. flower_ovary_position_three_quarters_inferior Flower where the receptacle surrounds more than three quarters of the ovary.
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution orientation position
keyword	reproduction gynoecium carpel plant ovary perianth
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012444

Number of ovules per functional carpel

Property	Value
URI	https://w3id.org/APD/traits/trait_0012445
alternative label	flower_ovules_per_functional_carpel_count
description	A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of ovules [PO:0020003] per fertile [PATO:0000955] carpel [PO:0009030]. Number of ovules per carpel recorded as a continuous character (with integer values of 1 and above).
comments	Reduced (sterile) carpels are not taken into account here. Definition derived from Schoenenberger et al. (2020) for their character ‘411. Number of ovules per functional carpel (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	1e+05
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction gynoecium carpel plant ovary plant ovule
scope note	Only applicable to fertile carpels.
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012445

Perianth differentiation

Property	Value
URI	https://w3id.org/APD/traits/trait_0012461
alternative label	flower_perianth_differentiation
description	A floral organ morphology trait [TO:0000862] which indicates whether petals [PO:0009032] and sepals [PO:0009031] are differentiated or identical and the various ways they can be identical or different. The ways in which perianth organs may look different from each other in a given flower.
comments	Typically, outer perianth parts are sepaloid and protect the other floral organs during floral development, while inner organs are often petaloid and play a role in pollinator attraction (Endress, 1994). However, it is also possible that all parts are either sepaloid or petaloid but remain differentiated in shape, size, and/or texture. In case of spiral perianths, differentiation may be continuous (i.e., gradual), whereby two successively initiated organs are very similar or only slightly different, while the outermost and innermost organs at both ends of the spiral are very different from each other (e.g., Chimonanthus). In the special case of perianths consisting of a single whorl, they are scored as undifferentiated. This perianth differentiation trait may be seen as both functional (the parting vs. sharing of functions among perianth parts) and developmental (the expression of a genetic program for different forms of perianth parts vs. a single program for a single type of perianth part morphology). Within-whorl differentiation, whereby organs of the same whorl take different forms, is common in zygomorphic flowers (e.g., Balsaminaceae, Fabaceae, Orchidaceae) but is not taken into account with this trait. This trait is not applicable when the perianth is absent. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘234. Perianth differentiation (D1)’
value type	categorical variable
has narrower	flower_perianth_differentiation_undifferentiated Flower were all tepals are alike, including flowers with a single, undifferentiated whorl. flower_perianth_differentiation_marked_differentiation_outer_sepaloid_inner_petaloid A typical flower with a distinct calyx and corolla. flower_perianth_differentiation_marked_differentiation_among_petaloid_tepals A flower with tepals that all resemble petals, but with marked differentiation between whorls of tepals. flower_perianth_differentiation_marked_differentiation_among_sepaloid_tepals A flower with tepals that all resemble sepals, but with marked differentiation between whorls of tepals. flower_perianth_differentiation_marked_differentiation A flower with marked differentiation between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals. flower_perianth_differentiation_continuous_differentiation_outer_sepaloid_inner_petaloid A flower with multiple whorls, where there is continuous differentiation between the outer sepaloid parts and the inner petaloid parts. flower_perianth_differentiation_continuous_differentiation_among_petaloid_tepals A flower with series of tepals that all resemble petals, which display continuous, gradual differentiation between whorls. flower_perianth_differentiation_continuous_differentiation_among_sepaloid_tepals A flower with series of tepals that all resemble sepals, which display continuous, gradual differentiation between whorls. flower_perianth_differentiation_weak_differentiation A flower with weak between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals. flower_perianth_differentiation_very_weak_differentiation A flower with very weak between whorls, but without an indication if the perianth whorls are petals, sepals, or tepals.
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	structure shape
keyword	reproduction perianth sepal petal corolla calyx
scope note	Not applicable when the perianth is absent
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012461

Fusion of perianth

Property	Value
URI	https://w3id.org/APD/traits/trait_0012462
alternative label	flower_perianth_fusion
description	A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]) that are fused [NCIT:C124537] together at anthesis [PO:0009068]. Fusion of perianth organs (congenital or postgenital) at anthesis, recorded on a continuous scale, from 0 (free parts) to 1 (parts fused along their entire length).
comments	Partial fusion was recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the length of perianth parts). In case of multiple whorls, this trait records within-whorl fusion. For example, if organs within each whorl are fused along their entire length, a trait value of 1 is recorded. In cases where organs of two whorls are fused into a common tubular structure, such as frequently observed in monocots (e.g., Polygonatum), the perianth is considered to be fused. If the two (or more) whorls differed in their extent of fusion, a range of trait values may be recorded. For example, if the calyx is only basally fused, up to 10% of its length, but the corolla is entirely fused, a range of 0.1 to 1 is recorded as the trait value for the species. Definition derived from Schoenenberger et al. (2020) for their character ‘204. Fusion of perianth (C1)’
value type	continuous variable
unit	{dimensionless}
minAllowedValue	0
maxAllowedValue	1
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	dimensionless ratio ratio; proportion structure
keyword	reproduction perianth sepal petal corolla calyx
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012462

Number of perianth parts in each whorl (Perianth merism)

Property	Value
URI	https://w3id.org/APD/traits/trait_0012463
alternative label	flower_perianth_merism
description	A perianth morphology trait [TO:0000275] which is the count [NCIT:C25463] of perianth [PO:0009058] parts (sepals [PO:0009031], petals [PO:0009032], and tepals [PO:0009033]) in each floral whorl [PO:0025023]. The number of perianth parts in each whorl, recorded as a continuous trait (with integer values of 1 and above).
comments	Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent. Definition derived from Schoenenberger et al. (2020) for their character ‘232. Perianth merism (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	50
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count orientation
keyword	reproduction perianth sepal petal corolla calyx merism whorl, floral whorl
scope note	Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent.
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012463

Number of perianth parts

Property	Value
URI	https://w3id.org/APD/traits/trait_0012464
alternative label	flower_perianth_parts_count
description	A perianth morphology trait [TO:0000275] which is the total count [NCIT:C25463] of perianth [PO:0009058] parts, including sepals [PO:0009031], petals [PO:0009032], and any form of tepal [PO:0009033]. The total number of perianth parts, including sepals, petals, or any form of tepal.
comments	A value of zero was scored when the perianth is absent. In flowers with perianth whorls fused along their complete length (e.g., Convolvulus), perianth parts are counted based on merism (e.g., if a calyx has 5 distinct sepals and the corolla is entirely fused, then the corolla is often interpreted to consist of five fused petals), anatomy (e.g., number of vascular traces), development (e.g., number of primordia), or comparison with closely related taxa. Definition derived from Schoenenberger et al. (2020) for their character ‘201. Number of perianth parts (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	1000
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction perianth sepal petal corolla calyx
scope note
has exact match	tepal number
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012464

Perianth phyllotaxis

Property	Value
URI	https://w3id.org/APD/traits/trait_0012465
alternative label	flower_perianth_phyllotaxis
description	A perianth morphology trait [TO:0000275] which is the phyllotaxy [TO:0006014] of the perianth [PO:0009058] parts (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]), at anthesis [PO:0009068] Perianth phyllotaxis at anthesis.
comments	Perianth parts may be organised in one or more whorls or along a continuous spiral, usually with wide divergence angles more or less equal to 137.5 degrees. Less frequently, perianth phyllotaxis may be irregular. Perianth phyllotaxis at anthesis may differ from phyllotaxis of perianth part primordia at their inception and it is not uncommon that spirally initiated perianths become whorled later through development. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘230. Perianth phyllotaxy (D1)’, with more information of differentiating between trait values in this manuscript.
value type	categorical variable
has narrower	flower_perianth_phyllotaxis_irregular Perianth parts lacking any symmetrical organisation. flower_perianth_phyllotaxis_spiral Perianth parts organised along a continuous spiral, usually with wide divergence angles more or less equal to 137.5 degrees, the Fibonacci ‘golden angle’. flower_perianth_phyllotaxis_whorled Perianth parts organised in one or more whorls.
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution orientation
keyword	reproduction perianth sepal petal corolla calyx whorl, floral whorl phyllotaxy
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012465

Symmetry of perianth

Property	Value
URI	https://w3id.org/APD/traits/trait_0012466
alternative label	flower_perianth_symmetry
description	A perianth morphology trait [TO:0000275] which is the symmetry [PATO:0000965] of the perianth [PO:0009058] (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]), whereby a perianth is only considered actinomorphic [PATO:0001328] if all perianth whorls [PO:0025023] are actinomorphic. Trait capturing the symmetry of the perianth.
comments	This trait is applied to the perianth as a whole. In case of flowers with two or more perianth whorls, species are considered actinomorphic if all whorls are actinomorphic and as zygomorphic if one or more whorls are zygomorphic. Trait is not applicable when the perianth is missing. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘207. Symmetry of perianth (D1)’
value type	categorical variable
has narrower	flower_perianth_symmetry_actinomorphic_general Perianth displaying radial symmetry. flower_perianth_symmetry_actinomorphic_rotational Perianth that can be equally divided into three or more identical sections that, when rotated around the center of the flower by some number of degrees, exactly match each other in orientation and shape. flower_perianth_symmetry_actinomorphic_strictly_polysymmetric Polysymmetry, with three or more planes of bilateral symmetry. flower_perianth_symmetry_actinomorphic_spiral Perianth lacking a distinct plane of symmetry because their perianth parts are spirally arranged. flower_perianth_symmetry_asymmetric Perianth lacking any planes of symmetry. flower_perianth_symmetry_disymmetric Perianth with two orthogonal planes of bilateral symmetry. flower_perianth_symmetry_zygomorphic Monosymmetric, with a single plane of bilateral symmetry.
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	spatial pattern, distribution symmetry orientation
keyword	reproduction perianth sepal petal corolla calyx whorl, floral whorl
scope note	Not applicable when the perianth is missing
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012466

Number of perianth whorls

Property	Value
URI	https://w3id.org/APD/traits/trait_0012467
alternative label	flower_perianth_whorls_count
description	A perianth morphology trait [TO:0000275] which is the count [NCIT:C25463] of floral whorls [PO:0025023] that comprise the perianth [PO:0009058] (petals [PO:0009032], sepals [PO:0009031], and tepals [PO:0009033]). The number of perianth whorls as a continuous trait (with integer values of 1 and above).
comments	Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent. Definition derived from Schoenenberger et al. (2020) for their character ‘231. Number of perianth whorls (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	15
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction perianth sepal petal corolla calyx whorl, floral whorl
scope note	Not applicable when perianth phyllotaxis is spiral or irregular or when the perianth is absent.
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012467

Pollen grain aperture shape

Property	Value
URI	https://w3id.org/APD/traits/trait_0012471
alternative label	flower_pollen_aperture_shape
description	A floral organ morphology trait [TO:0000862] which is the shape [PATO:0000052], structure [PATO:0000141], and position [PATO:0000140] of the pollen grain [PO:0025281] apertures [GO:0062074]. Pollen grains are often described according to the shape, structure, and position of their apertures. This trait describes the main shape of the many described aperture types.
comments	Aperture terminology is partly determined by the position of the aperture on the pollen grain (either at the pole or at the equator) and pollen grain polarity, in turn, is determined by the spatial orientation of the microspore in the meiotic tetrad. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘5002. Aperture shape (D1)’’
value type	categorical variable
has narrower	flower_pollen_aperture_shape_absent Pollen that lacks an aperture. flower_pollen_aperture_shape_colporus Pollen aperture shape that combines the groove of colpate and the pore of porate apertures. flower_pollen_aperture_shape_colpus Pollen aperture shape is elongate and grooved and may be positioned globally or equatorially. flower_pollen_aperture_shape_pore Pollen aperture shape is round and pore-like and may be positioned globally or equatorially. flower_pollen_aperture_shape_ring_like Aperture that completely encircles the pollen. flower_pollen_aperture_shape_spiraperturate Pollen grain with one or more spiral apertures. flower_pollen_aperture_shape_sulcus Pollen aperture shape is elongate and grooved and positioned at the pole. flower_pollen_aperture_shape_syncolpus Pollen grain with two or more simple (or compound) colpi, the ends of which join at the pole. flower_pollen_aperture_shape_ulcus Pollen aperture shape is round and pore-like and positioned at the pole.
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	shape structure position spatial pattern, distribution
keyword	reproduction androecium stamen anther pollen aperture pollen pollen tube anther theca
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012471

Number of pollen grain apertures

Property	Value
URI	https://w3id.org/APD/traits/trait_0012472
alternative label	flower_pollen_apertures_count
description	A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of pollen apertures [GO:0062074] per pollen [PO:0025281] grain. The number of apertures per pollen grain as a continuous character.
comments	A pollen aperture is a structurally delimited region of the pollen grain wall through which the pollen tube emerges during pollen germination and which plays a role in harmomegathy. A value of zero is scored when pollen is inaperturate (i.e. when there is no distinct aperture). Aperturate pollen is scored with integer values of 1 and above. Definition derived from Schoenenberger et al. (2020) for their character ‘5000. Number of apertures (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	100
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction androecium stamen anther pollen aperture pollen pollen tube anther theca
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012472

Pollen grain length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012473
alternative label	flower_pollen_length
description	A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of a pollen [PO:0025281] grain. The largest dimension of the pollen grain.
comments	Palynologists typically describe pollen dimensions (and shape) using two measures, the polar axis (P) and the total breadth or diameter of the equatorial axis. Here we use P or E or another measure, whichever is the longest.
value type	continuous variable
unit	um micrometre
minAllowedValue	0
maxAllowedValue	1000
has broader	androecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	length
keyword	reproduction pollen
scope note
has exact match
has close match
has related match
examples
references	no linked references
date created	19/05/2023
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012473

Number of structural carpels

Property	Value
URI	https://w3id.org/APD/traits/trait_0012481
alternative label	flower_structural_carpels_count
description	A floral organ morphology trait [TO:0000862] which is the count [NCIT:C25463] of both fertile [PATO:0000955] and female sterile [PATO:0000892] carpels [PO:0009030] in a hermaphrodite [PATO:0001340] flower [PO:0009046] or female [PATO:0000383] flower. Number of fertile or sterile carpels in bisexual or female flowers, recorded as a continuous character (with integer values of 1 and above).
comments	Includes the number of co-occurring carpellodes (sterile carpels) because this number is often more easily obtained from the literature than the actual number of fertile carpels. However, the number of carpellodes in male flowers is ignored for this character. In multicarpellate, unilocular gynoecia with complete carpel fusion up to the stigma (e.g., Primula), it may be difficult to assess the number of carpels unequivocally. In such cases, the number of carpels is scored only if it is well established based on atomical or developmental investigations. Similarly, in gynoecia where one or more carpels are reduced (e.g., in the pseudomonomerous gynoecia of some Arecaceae, Stauffer et al., 2002), the total number of structural carpels is only scored when unequivocally determined in the literature. Definition derived from Schoenenberger et al. (2020) for their character ‘401. Number of structural carpels (C1)’
value type	continuous variable
unit	{count}
minAllowedValue	0
maxAllowedValue	10000
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	has number of count
keyword	reproduction gynoecium carpel
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012481

Floral structural sex

Property	Value
URI	https://w3id.org/APD/traits/trait_0012482
alternative label	flower_structural_sex_type
description	A floral organ morphology trait [TO:0000862] which is the phenotypic sex [PATO:0001894] of a flower [PO:0009046], indicating whether individual flowers have both male [PATO:0000384] and female [PATO:0000383] reproductive structures [UBERON:0005156] (bisexual) or only one type of reproductive organ (unisexual). Trait that captures floral sex, distinguishing among three states, bisexual, incompletely unisexual, and unisexual.
comments	Defined by Schoenenberger et al. (2020) for their character ‘100. Floral structural sex (D1)’. See the trait ‘sex_type’ for plant sex type.
value type	categorical variable
has narrower	flower_structural_sex_type_bisexual Hermaphrodite; flowers with both male and female reproductive organs. flower_structural_sex_type_incompletely_unisexual Male flowers with pistillodes and/or female flowers with staminodes. flower_structural_sex_type_unisexual Flowers with either male or female reproductive organs.
has broader	floral organ morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	structure organismal quality
keyword	reproduction phenotypic sex perianth corolla hermaphrodite
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012482

Style differentiation

Property	Value
URI	https://w3id.org/APD/traits/trait_0012483
alternative label	flower_style_differentiation
description	A floral organ morphology trait [TO:0000862] which is whether a style [PO:0006469] is present [PATO:0000467] or absent [PATO:0000462], and if present, the shape [PATO:0000052] and the length [PATO:0000122], width [PATO:0000921], and thickness [PATO:0000915] of the style relative to the dimensions of the ovary [PO:0009072]. Presence or absence of a style, and, when present, the shape of the style in terms of length and width in relation to ovary length and width.
comments	This trait does not distinguish between fused or free styles. Definition and trait values derived from Schoenenberger et al. (2020) for their character ‘404. Style differentiation (D1)’
value type	categorical variable
has narrower	flower_style_differentiation_absent Stigma sessile. flower_style_differentiation_present_length_and_shape_unknown Style present but shape unknown. flower_style_differentiation_present_short_and_thick Style present, and short and thick. flower_style_differentiation_present_short_and_narrow Style present, and short and narrow, including short filiform styles. flower_style_differentiation_present_long_and_wide Style present, and long and wide. flower_style_differentiation_present_long_and_narrow Style present, and long and narrow, including long filiform styles. flower_style_differentiation_present_petaloid Style present and petaloid. flower_style_differentiation_continuous When the style is an apical extension of carpel with decurrent stigma.
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	shape amount (presence or absence in organism 2-D extent structure
keyword	reproduction gynoecium style plant ovary carpel
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012483

Fusion of styles

Property	Value
URI	https://w3id.org/APD/traits/trait_0012484
alternative label	flower_style_fusion
description	A floral organ morphology trait [TO:0000862] which is the proportion [PATO:0001470] of the length [PATO:0000122] of styles [PO:0006469] that are fused [NCIT:C124537] together. The degree of fusion of styles at anthesis, recorded on a continuous scale, from 0 (free styles) to 1 (styles fused along their entire length, but excluding the stigmatic region).
comments	Partial fusion is recorded using an approximate number between these two extremes (e.g., 0.1 corresponds to basal fusion, 0.5 to fusion along half of the total length of the styles). This character is not applicable in unicarpellate flowers. Definition derived from Schoenenberger et al. (2020) for their character ‘406. Fusion of styles (C1)’
value type	continuous variable
unit	{dimensionless}
minAllowedValue	0
maxAllowedValue	1
has broader	gynoecium morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	dimensionless ratio ratio; proportion structure
keyword	reproduction gynoecium style carpel
scope note
has exact match
has close match
has related match
examples
references	Schönenberger 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	24/05/2023
reviewed by	Hervé Sauquet
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012484

Petal length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012485
alternative label	flower_petal_length
description	A floral organ morphology trait [TO:0000862] which is the length [PATO:0000122] of a petal [PO_0009032]. The length of an individual flower petal.
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	1
maxAllowedValue	100
has broader	perianth morphology trait
plant structure plant structure	reproductive shoot system flower
measured characteristic	length size
keyword	reproduction petal corolla perianth petal morphology trait
scope note	applicable to flowering plants
has exact match	petal length
has close match
has related match
examples	exact match: Flower petal length [TRY:3576] (https://www.try-db.org/de/de.php)
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012485

Fruit dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012511
alternative label	fruit_dry_mass
description	A fruit morphology trait [TO:0002629] which is the dry [PATO:0001824] mass [PATO:0000125] of a fruit [PO:0009001], including seeds [PO:0009010]. Fruit dry mass, including seeds. Fruits will generally be oven-dried.
comments
value type	continuous variable
unit	mg milligram
minAllowedValue	1e-04
maxAllowedValue	1e+06
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	mass
keyword	reproduction
scope note
has exact match	fruit dry weight
has close match
has related match
examples	exact match: Fruit dry mass [TOP112] (https://top-thesaurus.org/index) exact match: Fruit dry mass [TRY:919] (https://www.try-db.org/de/de.php) exact match: Fruit dry mass [BROT:25] (http://doi.org/10.1038/sdata.2018.135)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012511

Fruit length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012512
alternative label	fruit_length
description	A fruit morphology trait [TO:0002629] which is the length [PATO:0000122] of a fresh [EnvThes:21976] fruit [PO:0009001] from the fruit proximal end [PO:0008002] (base) to the fruit distal end [PO:0008001] (apex). Linear dimension from the base to the apex of a fresh fruit, even if this is not the longest dimension.
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	2000
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	length size
keyword	reproduction
scope note
has exact match	fruit length
has close match
has related match
examples	exact match: Fruit length [TOP92] (https://top-thesaurus.org/index) exact match: Fruit length [TRY:918] (https://www.try-db.org/de/de.php) exact match: Fruit length [GIFT:3.13] (https://gift.uni-goettingen.de) close match: maximum fruit length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) close match: minimum fruit length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: AverageFruitLength_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) exact match: MinFruitLength_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) exact match: MaxFruitLength_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012512

Fruit width

Property	Value
URI	https://w3id.org/APD/traits/trait_0012513
alternative label	fruit_width
description	A fruit morphology trait [TO:0002629] which is the width [PATO:0000921] of a fresh [EnvThes:21976] fruit [PO:0009001], where width is the greatest linear dimension [PATO:0001708] perpendicular to [PATO:0002434] to the fruit length [PATO:0000122] and fruit height [PATO:0000119]. Greatest linear width dimension of a fresh fruit, sometimes referred to as ‘diameter’ in fruits where the width and height are identical; orthogonal to the fruit length, the base-to-apex dimension.
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	2000
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	width size
keyword	reproduction
scope note
has exact match	fruit width
has close match
has related match
examples	exact match: Fruit width [TOP96] (https://top-thesaurus.org/index) exact match: Fruit width (diameter) [TRY:917] (https://www.try-db.org/de/de.php) exact match: Fruit width [GIFT:3.14] (https://gift.uni-goettingen.de) exact match: AverageFruitWidth_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) exact match: MinFruitWidth_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) exact match: MaxFruitWidth_cm [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012513

Fruit breadth

Property	Value
URI	https://w3id.org/APD/traits/trait_0012514
alternative label	fruit_height
description	A fruit morphology trait [TO:0002629] which is the height [PATO:0000119] of a fresh [EnvThes:21976] fruit [PO:0009001], where height is the shorter linear width dimension [PATO:0001708] perpendicular to [PATO:0002434] both the fruit length [PATO:0000122] and fruit width [PATO:0000921]. Shorter linear width dimension of a fresh fruit and generally not reported in fruits where the width and height are identical; orthogonal to the length, the base-to-apex dimension.
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	2000
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	height size
keyword	reproduction
scope note
has exact match
has close match
has related match
examples	exact match: Fruit thickness [TOP100] (https://top-thesaurus.org/index) exact match: Fruit height [GIFT:3.15] (https://gift.uni-goettingen.de)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note	fruit_breadth
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012514

Fruit wall thickness

Property	Value
URI	https://w3id.org/APD/traits/trait_0012515
alternative label	fruit_wall_thickness
description	A fruit morphology trait [TO:0002629] which is the thickness [PATO:0000915] of the entire pericarp [PO:0009084] (the exocarp [PO:0009085], mesocarp [PO:0009087], and endocarp [PO:0009086] combined), although it often refers just to the fleshy [PATO:0002351] components of the pericarp (most frequently the exocarp and mesocarp). Fruit wall thickness of a mature fresh fruit. This term sometimes applies explicitly to the entire pericarp (exocarp + mesocarp + endocarp), but more frequently will have a functional meaning, referring to the outer fleshy layer which might be just some components of the peripcarp (most frequently the exocarp and mesocarp).
comments
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	1000
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	thickness
keyword	reproduction pericarp exocarp mesocarp endocarp
scope note
has exact match
has close match
has related match
examples
references	Simpson 2019
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note	fruit_wall_width
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012515

Fruit type

Property	Value
URI	https://w3id.org/APD/traits/trait_0012516
alternative label	fruit_type
description	A fruit morphology trait [TO:0002629] which is the botanical classification [NCIT:C25161] or type [NCIT:C25284] of a fruit [PO:0009001]. Botanical fruit type classification.
comments	Although this trait is explicitly for fruit types, it includes a single term strobilus to describe the ‘fruiting body’ of non-flowering plants.
value type	categorical variable
has narrower	fruit_type_achene A simple, dry, indehiscent fruit with a single seed that is attached to the pericarp at one point only. (Synonyms, cypsela) fruit_type_anthocarp A ripened ovary surrounded by attached accessory floral tissues; the floral tissues may be fleshy or dry. (Synonyms, accessory_fruit, pseudocarp, diclesium) fruit_type_berry A simple, indehiscent fruit with one to many seeds embedded in a fleshy pericarp. fruit_type_capsule A dry, dehiscent fruit composed of two or more united carpels; different types of capsules distinguished by type of dehiscence (e.g. valves, slits, pores). fruit_type_caryopsis A simple, dry, indehiscent fruit with a single seed which is fused to the pericarp at one point only; the fruit type of all Poaceae (Synonyms, grain) fruit_type_compound_fruit_other A fruit derived from more than one flower. (Synonyms, aggregate_fruit) fruit_type_drupe A simple indehiscent fruit with a hard, stony endocarp and a fleshy mesocarp. fruit_type_follicle A dry, dehiscent fruit derived from a single carpel that splits along one suture. fruit_type_legume A dry, dehiscent fruit derived from a single carpel that splits along two sutures; applies to Fabaceae. (Synonyms, pod) fruit_type_legume_indehiscent A dry, indehiscent fruit derived from a single carpel; applies to Fabaceae. fruit_type_mericarp A fruitlet (seed-containing portion of a fruit) that is a half-carpel; the individual fruitlets separate from one another at maturity; a specific type of schizocarp. (In some datasets, term likely refers to any fruitlet derived from a portion of a schizocarp that separates at maturity, not explicitly just a half-carpel.) fruit_type_multiple_fruit A fruit composed of several fruitlets developing from a single flower (e.g. raspberries, magnolias); fruits developing from more than one flower are included under compound (aggregate) fruits. (Synonym, apocarp) fruit_type_nut A simple, dry, indehiscent fruit with a single seed and a hard pericarp. fruit_type_nutlet A diminutive of nut, referring to a simple, dry, indehiscent structure with a single seed that might be an entire fruit (nut; monocarp) or a portion of a fruit, such as the individual components derived from an apocarp (fruit with multiple unfused ovaries) or schizocarp (fruit with multiple ovaries that separate at maturity). fruit_type_pepo A fleshy fruit derived from a single ovary that is divided into several carpels, has multiple seeds, and has a tough rind when mature. fruit_type_pome A fleshy fruit with the bulk of the fleshy tissue derived from flora tissue that surrounds the inferior ovary. fruit_type_pyrene A fleshy fruit in which the seed(s) are covered by a hard layer of endocarp. fruit_type_samara A winged, dry fruit. fruit_type_schizocarp Category that includes all fruits derived from a compound pistil that separate into individual carpellary constituents at maturity; broad terms that includes dry and fleshy fruits, some with accessory tissues, others without. The individual seed-containing units that a schizocarp breaks apart into include mericarps, nutlets, and fruitlets. fruit_type_silique A type of capsule (dry, dehiscent fruit) where the fruit is divided into two parts by a thin partition; more than three times as long as wide; applies to Brassicaceae (Synonyms, silicle, siliqua) fruit_type_strobilus A modified reproductive shoot system found in many nonflowering plants, especially gymnosperms, comprised of overlapping bracts (scales). (Synonyms, cone) fruit_type_syncarp A compound fruit produced by the adhesion of the fruits from several flowers. fruit_type_syconium A fleshy compound fruit, in which the fruitlets are enclosed in accessory material, such as the hollow-centered fruits of Moraceae. (Synonyms, synconium, fig) fruit_type_utricle A bladdery or inflated dry fruit with a single seed; essential an achene in which the pericarp is significantly larger than the mature seed.
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	structure composition shape
keyword	reproduction
scope note
has exact match	fruit type
has close match
has related match
examples	exact match: Fruit type [TRY:99] (https://www.try-db.org/de/de.php) exact match: Fruit_type_1 [GIFT:3.16.1] (https://gift.uni-goettingen.de) exact match: fruit type [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Fruit type [BROT:26] (http://doi.org/10.1038/sdata.2018.135)
references	Kew Seed Information Database 2022 Beentje 2010 Spjut 2003
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012516

Fruit fleshiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0012517
alternative label	fruit_fleshiness
description	A fruit morphology trait [TO:0002629] which indicates whether a mature [PATO:0001701] fresh [EnvThes:21976] fruit [PO:0009001] is fleshy [PATO:0002351]. Binary variable, dividing entire, true fruits into ‘dry’ versus ‘fleshy’ based on whether their botanical fruit type is fleshy when mature and fresh.
comments	This trait explicitly refers to fruit fleshiness, not diaspore or seed fleshiness; diaspore fleshiness is a separate trait.
value type	categorical variable
has narrower	fruit_fleshiness_dry Fruits are dry when mature and fresh. fruit_fleshiness_fleshy Fruits are fleshy when mature and fresh.
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	composition structure texture
keyword	reproduction fleshy
scope note
has exact match
has close match
has related match
examples	close match: Fruit type [TRY:99] (https://www.try-db.org/de/de.php) exact match: Fruit_dryness_1 [GIFT:3.18.1] (https://gift.uni-goettingen.de)
references	Kew Seed Information Database 2022 Beentje 2010 Spjut 2003
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012517

Fruit dehiscence

Property	Value
URI	https://w3id.org/APD/traits/trait_0012518
alternative label	fruit_dehiscence
description	A fruit morphology trait [TO:0002629] which indicates whether mature [PATO:0001701] fruits [PO:0009001] are dehiscent [GO:0009900]. Binary variable, dividing fruits into ‘dehiscent’ versus ‘indehiscent’ based on whether the individual fruit units are dehiscent when mature.
comments
value type	categorical variable
has narrower	fruit_dehiscence_dehiscent Fruits are dehiscent when mature. fruit_dehiscence_indehiscent Fruits are indehiscent when mature.
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	structure
keyword	reproduction dehiscence
scope note
has exact match
has close match
has related match
examples	close match: Fruit type [TRY:99] (https://www.try-db.org/de/de.php) exact match: Dehiscence_1 [GIFT:3.17.1] (https://gift.uni-goettingen.de)
references	Kew Seed Information Database 2022 Beentje 2010 Spjut 2003
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012518

Fruit colour

Property	Value
URI	https://w3id.org/APD/traits/trait_0012519
alternative label	fruit_colour
description	A fruit morphology trait [TO:0002629] which is the primary [NCIT:C25251] colour [PATO:0000014] of the exocarp [PO:0009085] of a mature [PATO:0001701] fruit [PO:0009001]. Human visible dominant colour of a fresh fruit’s exocarp (outermost layer) at maturity.
comments	If a fruit is multiple colours, they are listed as a space-delimited string.
value type	categorical variable
has narrower	fruit_colour_pink Fruit’s exocarp is pink. fruit_colour_red Fruit’s exocarp is red. fruit_colour_orange Fruit’s exocarp is orange. fruit_colour_yellow Fruit’s exocarp is yellow. fruit_colour_green Fruit’s exocarp is green. fruit_colour_blue Fruit’s exocarp is blue. fruit_colour_purple Fruit’s exocarp is purple. fruit_colour_white Fruit’s exocarp is white. fruit_colour_cream Fruit’s exocarp is cream. fruit_colour_brown Fruit’s exocarp is brown. fruit_colour_black Fruit’s exocarp is black. fruit_colour_grey Fruit’s exocarp is grey.
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	colour / color
keyword	reproduction exocarp pericarp
scope note
has exact match	fruit color exocarp color
has close match
has related match
examples	exact match: Fruit/seed color [TRY:585] (https://www.try-db.org/de/de.php) close match: Dispersal unit color [TRY:392] (https://www.try-db.org/de/de.php) exact match: Fruit_colour [GIFT:3.22.1] (https://gift.uni-goettingen.de) exact match: MainFruitColors [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0) close match: FruitColorDescription [PalmTraits] (https://doi.org/10.1038/s41597-019-0189-0)
references	Kissling 2019 Delmas 2020
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012519

Fruit surface hairiness

Property	Value
URI	https://w3id.org/APD/traits/trait_0012521
alternative label	fruit_surface_hairs
description	A fruit morphology trait [TO:0002629] which is the presence or absence of trichomes [PO:0006504] (hairs) on the surface (exocarp [PO:0009085]) of a mature [PATO:0001701] fruit [PO:0009001]. Surface hairiness of a mature fruit, including vestiture (trichome cover) and bristles.
comments
value type	categorical variable
has narrower	fruit_surface_hairs_glabrous Mature fruit do not have hairs (trichomes). fruit_surface_hairs_hairy Mature fruit have hairs (trichomes).
has broader	fruit morphology trait
plant structure plant structure	reproductive shoot system fruit
measured characteristic	texture pilosity
keyword	trichome fruit morphology trait
scope note
has exact match	fruit hairiness
has close match
has related match
examples
references	no linked references
date created	01/04/2025
date modified	01/04/2025
date reviewed
reviewed by	no reviewers
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012521

Seed dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012610
alternative label	seed_dry_mass
description	A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of a mature [PATO:0001701] seed [PO:0009010]. Dry mass of a mature seed, including both oven dried and air-dried samples.
comments	Standard methods people will have used to dry seeds include, ‘fresh’ (at dispersal, mature); ‘air dried’ (at local ambient conditions); ‘seed bank air dried’ (to 15% relative humidity); and ‘oven dried’ (>100 deg C for a set number of hours; e.g. seed bank standard is 103 deg C for 17 hours). It is expected that some observations in AusTraits mapped onto ‘seed_dry_mass’ will actually include both the seed and some dispersal tissue, if the two cannot easily be separated; these should be mapped to ‘diaspore_dry_mass’.
value type	continuous variable
unit	mg milligram
minAllowedValue	1e-05
maxAllowedValue	1e+06
has broader	seed morphology trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass
keyword	reproduction
scope note
has exact match	seed weight, seed mass seed mass
has close match
has related match
examples	exact match: Seed dry mass [TOP111] (https://top-thesaurus.org/index) exact match: Seed dry mass [TRY:26] (https://www.try-db.org/de/de.php) close match: seed weight & seed shape [LEDA:5.2] (https://uol.de/en/landeco/research/leda/standards) exact match: Seed mass [GIFT:3.2] (https://gift.uni-goettingen.de) exact match: seed mass [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata) exact match: Seed dry mass [BROT:27] (http://doi.org/10.1038/sdata.2018.135)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022 Saatkamp 2018
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note	seed_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012610

Diaspore dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012611
alternative label	diaspore_dry_mass
description	A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of a seed [PO:0009010] and the seed dispersal [EnvThes:21042] structures [PATO:0000141]. Dry mass of a plant’s dispersal unit, including the seed and any dispersal appendages; synonymous with the term dispersule and often synonymous with the term disseminule.
comments	Data mapped onto this trait have been very explicitly defined as a ‘diaspore’. There will be additional values mapped as either ‘seed_dry_mass’ or ‘fruit_dry_mass’ that are also diaspores, but have identified only as being a ‘seed’ or ‘fruit’. In addition, it is expected that some observations in AusTraits mapped onto ‘seed_dry_mass’ will actually include both the seed and some dispersal tissue, if the two cannot easily be separated.
value type	continuous variable
unit	mg milligram
minAllowedValue	1e-05
maxAllowedValue	1e+06
has broader	seed morphology trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass
keyword	dispersal reproduction seed dispersal
scope note
has exact match
has close match
has related match
examples	exact match: Dispersule dry mass trait [TOP110] (https://top-thesaurus.org/index) exact match: Dispersal unit dry mass [TRY:233] (https://www.try-db.org/de/de.php) close match: seed weight & seed shape [LEDA:5.2] (https://uol.de/en/landeco/research/leda/standards)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note	diaspore_mass
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012611

Seed embryo and endosperm dry mass

Property	Value
URI	https://w3id.org/APD/traits/trait_0012612
alternative label	seed_dry_mass_reserve
description	A seed morphology trait [TO:0000184] which is the dry [PATO:0001824] mass [PATO:0000125] of the components of a seed [PO:0009010] that provide energy [PATO:0001021] to a seedling [EnvThes:20698], generally the embryo [PO:0009009] and, if present, the endosperm [PO:0009089]. Dry mass of a seed’s embryo and endosperm (if present), the components of a seed that provide energy to a seedling. These tissues are distinct from a seed’s dispersal structures or protective tissues.
comments
value type	continuous variable
unit	mg milligram
minAllowedValue	1e-05
maxAllowedValue	1e+05
has broader	seed morphology trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	mass
keyword	energy plant embryo endosperm reproduction seedling storage organ
scope note
has exact match
has close match
has related match
examples
references	Westoby 1990 Lamont & Groom 2013
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note	seed_mass_reserve
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012612

Seed length

Property	Value
URI	https://w3id.org/APD/traits/trait_0012613
alternative label	seed_length
description	A seed morphology trait [TO:0000184] which is the greatest linear dimension [PATO:0001708] (axis) of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed width [PATO:0000921] and seed height [PATO:0000119] axes [NCIT:C25154]. Longest linear seed dimension.
comments	Observations of this trait in AusTraits are almost certainly an undocumented mix of dimensions of true seeds and diaspores. When all or part of the fruit is not easily separated from the true seed, many researchers will have recorded and submitted a diaspore length, but labeled the data ‘seed length’. The intent of this measurement is that it excludes appendages, although it is likely some submitted measurements include small appendages that are not easily separated from the seed.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	1000
has broader	seed morphology trait
plant structure plant structure	reproductive shoot system seed
measured characteristic	length size
keyword	reproduction
scope note
has exact match	seed length
has close match
has related match
examples	exact match: Seed length [TOP91] (https://top-thesaurus.org/index) exact match: Seed length [TRY:27] (https://www.try-db.org/de/de.php) close match: seed weight & seed shape [LEDA:5.2] (https://uol.de/en/landeco/research/leda/standards) exact match: Seed length [GIFT:3.10] (https://gift.uni-goettingen.de) exact match: seed length [BIEN] (https://bien.nceas.ucsb.edu/bien/biendata)
references	Pérez-Harguindeguy 2013 Kew Seed Information Database 2022 Saatkamp 2018
date created	14/07/2021
date modified	05/02/2024
date reviewed	30/11/2021
reviewed by	Elizabeth Wenk Hervé Sauquet Russell Barrett Carl Gosper Lydia Guja Gregory J. Jordan Mark Ooi Karen D. Sommerville Lily Dun
change note
is in scheme	https://w3id.org/APD/traits
identifier	trait_0012613

Seed width

Property	Value
URI	https://w3id.org/APD/traits/trait_0012614
alternative label	seed_width
description	A seed morphology trait [TO:0000184] which is the longest [NCIT:C25558] width [PATO:0000921] dimension of a seed [PO:0009010] and is perpendicular to [PATO:0002434] the seed length [PATO:0000122] and seed height [PATO:0000119] axes [NCIT:C25154]. Longest width dimension of a seed; orthogonal to the length and height axes.
comments	Observations of this trait in AusTraits are almost certainly an undocumented mix of dimensions of true seeds and diaspores. When all or part of the fruit is not easily separated from the true seed, many researchers will have recorded and submitted a diaspore width, but labeled the data ‘seed width’. The intent of this measurement is that it excludes appendages, although it is likely some submitted measurements include small appendages that are not easily separated from the seed.
value type	continuous variable
unit	mm millimetre
minAllowedValue	0.01
maxAllowedValue	1000